Review of genus-group names in the family Tenebrionidae (Insecta, Coleoptera)

Authors: Patrice Bouchard, Yves Bousquet, Rolf L. Aalbu, Miguel A. Alonso-Zarazaga, Ottó Merkl, Anthony E. Davies

Abstract

A review of genus-group names for darkling beetles in the family Tenebrionidae (Insecta: Coleoptera) is presented. A catalogue of 4122 nomenclaturally available genus-group names, representing 2307 valid genera (33 of which are extinct) and 761 valid subgenera, is given. For each name the author, date, page number, gender, type species, type fixation, current status, and first synonymy (when the name is a synonym) are provided. Genus-group names in this family are also recorded in a classification framework, along with data on the distribution of valid genera and subgenera within major biogeographical realms. A list of 535 unavailable genus-group names (e.g., incorrect subsequent spellings) is included. Notes on the date of publication of references cited herein are given, when known.

The following genera and subgenera are made available for the first Anemiadena Bouchard & Bousquet, subgen. nov. (in Cheirodes Gené, 1839), Armigena Bouchard & Bousquet, subgen. nov. (in Nesogena Mäklin, 1863), Debeauxiella Bouchard & Bousquet, subgen. nov. (in Hyperops Eschscholtz, 1831), Hyperopsis Bouchard & Bousquet, subgen. nov. (in Hyperops Eschscholtz, 1831), Linio Bouchard & Bousquet, subgen. nov. (in Nilio Latreille, 1802), Matthewsotys Bouchard & Bousquet, gen. nov., Neosolenopistoma Bouchard & Bousquet, subgen. nov. (in Eurynotus W. Kirby, 1819), Paragena Bouchard & Bousquet, subgen. nov. (in Nesogena Mäklin, 1863), Paulianaria Bouchard & Bousquet, gen. nov., Phyllechus Bouchard & Bousquet, gen. nov., Prorhytinota Bouchard & Bousquet, subgen. nov. (in Rhytinota Eschscholtz, 1831), Pseudorozonia Bouchard & Bousquet, subgen. nov. (in Rozonia Fairmaire, 1888), Pseudothinobatis Bouchard & Bousquet, gen. nov., Rhytinopsis Bouchard & Bousquet, subgen. nov. (in Thalpophilodes Strand, 1942), Rhytistena Bouchard & Bousquet, subgen. nov. (in Rhytinota Eschscholtz, 1831), Spinosdara Bouchard & Bousquet, subgen. nov. (in Osdara Walker, 1858), Spongesmia Bouchard & Bousquet, subgen. nov. (in Adesmia Fischer, 1822), and Zambesmia Bouchard & Bousquet, subgen. nov. (in Adesmia Fischer, 1822).

The names Adeps Gistel, 1857 and Adepsion Strand, 1917 syn. nov. [= Tetraphyllus Laporte & Brullé, 1831], Asyrmatus Canzoneri, 1959 syn. nov. [= Pystelops Gozis, 1910], Euzadenos Koch, 1956 syn. nov. [= Selenepistoma Dejean, 1834], Gondwanodilamus Kaszab, 1969 syn. nov. [= Conibius J.L. LeConte, 1851], Gyrinodes Fauvel, 1897 syn. nov. [= Nesotes Allard, 1876], Helopondrus Reitter, 1922 syn. nov. [= Horistelops Gozis, 1910], Hybonotus Dejean, 1834 syn. nov. [= Damatris Laporte, 1840], Iphthimera Reitter, 1916 syn. nov. [= Metriopus Solier, 1835], Lagriomima Pic, 1950 syn. nov. [= Neogria Borchmann, 1911], Orphelops Gozis, 1910 syn. nov. [= Nalassus Mulsant, 1854], Phymatium Billberg, 1820 syn. nov. [= Cryptochile Latreille, 1828], Prosoblapsia Skopin & Kaszab, 1978 syn. nov. [= Genoblaps Bauer, 1921], and Pseudopimelia Gebler, 1859 syn. nov. [= Lasiostola Dejean, 1834] are established as new synonyms (valid names in square brackets). Anachayus Bouchard & Bousquet, nom. nov. is proposed as a replacement name for Chatanayus Ardoin, 1957, Genateropa Bouchard & Bousquet, nom. nov. as a replacement name for Apterogena Ardoin, 1962, Hemipristula Bouchard & Bousquet, nom. nov. as a replacement name for Hemipristis Kolbe, 1903, Kochotella Bouchard & Bousquet, nom. nov. as a replacement name for Millotella Koch, 1962, Medvedevoblaps Bouchard & Bousquet, nom. nov. as a replacement name for Protoblaps G.S. Medvedev, 1998, and Subpterocoma Bouchard & Bousquet, nom. nov. is proposed as a replacement name for Pseudopimelia Motschulsky, 1860. Neoeutrapela Bousquet & Bouchard, 2013 is downgraded to a subgenus (stat. nov.) of Impressosora Pic, 1952. Anchomma J.L. LeConte, 1858 is placed in Stenosini: Dichillina (previously in Pimeliinae: Anepsiini); Entypodera Gerstaecker, 1871, Impressosora Pic, 1952 and Xanthalia Fairmaire, 1894 are placed in Lagriinae: Lagriini: Statirina (previously in Lagriinae: Lagriini: Lagriina); Loxostethus Triplehorn, 1962 is placed in Diaperinae: Diaperini: Diaperina (previously in Diaperinae: Diaperini: Adelinina); Periphanodes Gebien, 1943 is placed in Stenochiinae: Cnodalonini (previously in Tenebrioninae: Helopini); Zadenos Laporte, 1840 is downgraded to a subgenus (stat. nov.) of the older name Selenepistoma Dejean, 1834.

The type species [placed in square brackets] of the following available genus-group names are designated for the first Allostrongylium Kolbe, 1896 [Allostrongyliumsilvestre Kolbe, 1896], Auristira Borchmann, 1916 [Auristiraoctocostata Borchmann, 1916], Blapidocampsia Pic, 1919 [Campsiapallidipes Pic, 1918], Cerostena Solier, 1836 [Cerostenadeplanata Solier, 1836], Coracostira Fairmaire, 1899 [Coracostiraarmipes Fairmaire, 1899], Dischidus Kolbe, 1886 [Helopssinuatus Fabricius, 1801], Eccoptostoma Gebien, 1913 [Taraxidesruficrus Fairmaire, 1894], Ellaemus Pascoe, 1866 [Emcephalussubmaculatus Brême, 1842], Epeurycaulus Kolbe, 1902 [Epeurycaulusaldabricus Kolbe, 1902], Euschatia Solier, 1851 [Euschatiaproxima Solier, 1851], Heliocaes Bedel, 1906 [Blaps emarginata Fabricius, 1792], Hemipristis Kolbe, 1903 [Hemipristisukamia Kolbe, 1903], Iphthimera Reitter, 1916 [Stenocararuficornis Solier, 1835], Isopedus Stein, 1877 [Helopstenebrioides Germar, 1813], Malacova Fairmaire, 1898 [Malacovabicolor Fairmaire, 1898], Modicodisema Pic, 1917 [Disemasubopaca Pic, 1912], Peltadesmia Kuntzen, 1916 [Metriopusplatynotus Gerstaecker, 1854], Phymatium Billberg, 1820 [Pimeliamaculata Fabricius, 1781], Podoces Péringuey, 1886 [Podocesgranosula Péringuey, 1886], Pseuduroplatopsis Pic, 1913 [Borchmanniajavana Pic, 1913], Pteraulus Solier, 1848 [Pteraulussulcatipennis Solier, 1848], Sciaca Solier, 1835 [Hylithusdisctinctus Solier, 1835], Sterces Champion, 1891 [Stercesviolaceipennis Champion, 1891] and Teremenes Carter, 1914 [Tenebriolongipennis Hope, 1843].

Evidence suggests that some type species were misidentified. In these instances, information on the misidentification is provided and, in the following cases, the taxonomic species actually involved is fixed as the type species [placed in square brackets] following requirements in Article 70.3 of the International Code of Zoological Nomenclature: Accanthopus Dejean, 1821 [Tenebriovelikensis Piller & Mitterpacher, 1783], Becvaramarygmus Masumoto, 1999 [Dietysusnodicornis Gravely, 1915], Heterophaga Dejean, 1834 [Opatrumlaevigatum Fabricius, 1781], Laena Dejean, 1821, [Scaurusviennensis Sturm, 1807], Margus Dejean, 1834 [Colydiumcastaneum Herbst, 1797], Pachycera Eschscholtz, 1831 [Tenebriobuprestoides Fabricius, 1781], Saragus Erichson, 1842 [Celibecostata Solier, 1848], Stene Stephens, 1829 [Colydiumcastaneum Herbst, 1797], Stenosis Herbst, 1799 [Tageniaintermedia Solier, 1838] and Tentyriopsis Gebien, 1928 [Tentyriopsispertyi Gebien, 1940].

The following First Reviser actions are proposed to fix the precedence of names or nomenclatural acts (rejected name or act in square brackets): Stenosisciliaris Gebien, 1920 as the type species for Afronosis G.S. Medvedev, 1995 [Stenosisleontjevi G.S. Medvedev, 1995], Alienoplonyx Bremer, 2019 [Alienolonyx], Amblypteraca Mas-Peinado, Buckley, Ruiz & García-París, 2018 [Amplypteraca], Caenocrypticoides Kaszab, 1969 [Caenocripticoides], Deriles Motschulsky, 1872 [Derilis], Eccoptostira Borchmann, 1936 [Ecoptostira], †Eodromus Haupt, 1950 [†Edromus], Eutelus Solier, 1843 [Lutelus], Euthriptera Reitter, 1893 [Enthriptera], Meglyphus Motschulsky, 1872 [Megliphus], Microtelopsis Koch, 1940 [Extetranosis Koch, 1940, Hypermicrotelopsis Koch, 1940], Neandrosus Pic, 1921 [Neoandrosus], Nodosogylium Pic, 1951 [Nodosogilium], Notiolesthus Motschulsky, 1872 [Notiolosthus], Pseudeucyrtus Pic, 1916 [Pseudocyrtus], Pseudotrichoplatyscelis Kaszab, 1960 [Pseudotrichoplatynoscelis and Pseudotrichoplatycelis], Rhydimorpha Koch, 1943 [Rhytimorpha], Rhophobas Motschulsky, 1872 [Rophobas], Rhyssochiton Gray, 1831 [Ryssocheton and Ryssochiton], Sphaerotidius Kaszab, 1941 [Spaerotidius], Stira Agassiz, 1846 (Mollusca) [Stira Agassiz, 1846 (Coleoptera)], Sulpiusoma Ferrer, 2006 [Sulpiosoma] and Taenobates Motschulsky, 1872 [Taeniobates].

Supporting evidence is provided for the conservation of usage of Cyphaleus Westwood, 1841 nomen protectum over Chrysobalus Boisduval, 1835 nomen oblitum.

Introduction

The last world catalogue of darkling beetles in the family Tenebrionidae Latreille, 1802 (Insecta: Coleoptera) was published by Gebien (1937a, 1938a, 1939, 1940, 1941, 1942a, 1943, 1948) more than 70 years ago. At the time, entire lineages were treated by coleopterists as separate families (e.g., Alleculidae Laporte, 1840, Lagriidae Latreille, 1825 (1820), Nilionidae Oken, 1843, Cossyphodidae Wasmann, 1899) and were therefore excluded from Gebien’s “Katalog der Tenebrioniden”. Since then, important studies on the relationships of taxa classified within Tenebrionidae and their close relatives, based on morphological (e.g., Doyen and Tschinkel 1982) and molecular data (e.g., Kergoat et al. 2014b; Kamiński et al. 2021b), have led to significant improvements in our understanding of the limits of the family and the relationships of major clades within the family. In addition, several taxa traditionally included in Tenebrionidae have now been transferred into closely related families (Table 1).

With more than 30 000 described species worldwide (RLA, unpubl. data) and many new species described each year, the family Tenebrionidae has been described as “hyperdiverse” (Kergoat et al. 2014a). This overwhelming diversity, combined with significant changes in classification over the last decades, has hindered the production of recent catalogues at a world scale. Although some catalogues that cover large geographic areas have been published (e.g., Matthews and Bouchard 2008; Bousquet et al. 2018; Iwan et al. 2020), the nomenclatural information contained therein often differs in scope and major regional gaps of knowledge (e.g., fauna in the Afrotropical and Neotropical biogeographic realms) still remain.

The main objective of this publication is to provide a nomenclatural review of all genus-group names in the family Tenebrionidae. A table that includes an up-to-date synthesis of the classification of available genus-group names, along with the distribution of each valid genus and subgenus within the world’s major biogeographical realms, is included. Following bibliographic research, dating of references cited in this work is provided (whenever data is available) in order to establish the priority of genus-group names and therefore promote their nomenclatural stability in the future.

Methods

Nomenclatural data

All nomenclaturally available genus-group names in the family Tenebrionidae are listed alphabetically. For each name the author, year of publication, page number, gender (in square M = masculine, F = feminine, N = neuter), type species, typification, current status, and first synonymy (when the name is a synonym) are provided. The type species and type species fixation given for unjustified emendations and replacement names are identical to those of the available names they replace (International Commission on Zoological Nomenclature (henceforth ICZN) 1999, Article 67.8) and are listed following “Type species [automatic]:”. We follow previous authors (e.g., Alonso-Zarazaga and Lyal 2009; Bousquet et al. 2015, 2018) in considering type-species designations in R. Lucas (1920) potentially valid when a single species is listed under a particular genus-group name. According to ICZN (1999, Article 67.2.5) “a nominal species is deemed not to be originally included if it was doubtfully or conditionally included, or was cited as a species inquirenda, or as a species incertae sedis.” For this catalogue, any nominal species associated with a new genus with a question mark “?” was deemed not to be originally included. In accordance with Article 11.9.3.2 (ICZN 1999) we used the correct spelling of the species name (i.e., in agreement with the gender of the generic name) for all species-group name combinations. The author of each new synonym or new placement is given in square brackets (e.g., “[RLA]”) in the alphabetical list of available genus-group names. We included all genus-group names known to us up to the date of publication of this article.

While the identification of the type species is generally assumed to be correct (ICZN 1999, Article 70.1), evidence suggests that the type species of a small number of genus-group names were misidentified. In these cases, we include information about the misidentification and select the species (either the nominal species previously cited as the type species or the taxonomic species actually involved) that will best serve stability and universality in our opinion (ICZN 1999, Article 70.3). If we select the taxonomic species actually involved as type species, an action that could not have been taken under previous editions of the International Code of Zoological Nomenclature and that must be accompanied with a reference to Article 73 (ICZN 1999), it is given in the format “fixed herein (ICZN 1999, Article 70.3) as Tenebriovelikensis Piller and Mitterpacher, 1783, misidentified as Tenebriocaraboides Linnaeus, 1758 in the original designation by monotypy in Dejean (1821).”

In some cases, we have noted that the original combination of the accepted name of the type species is a primary homonym of an older species name (e.g., Helopstenebrioides Germar, 1813, the type species of Isopedus Stein, 1877, is a junior primary homonym of Helopstenebrioides Palisot de Beauvois, 1812). We refrain proposing replacement names for two first, the status of some senior homonyms, which in most cases involved pre-1820 names, is uncertain; second, in some cases the two homonyms belong to taxa in use but the names apply to taxa not considered congeneric after 1899, and the Commission (ICZN 1999: Article 23.9.5) mandates that the author(s) must not automatically replace the junior homonym but instead should refer to the Commission for a ruling. We hope that experts on the taxonomy of the taxa involved will look at all evidence and propose solutions to remove the homonymies as needed.

Genus-group names encountered in the literature that are not nomenclaturally available are listed in Appendix 1. Reasons for assigning the status of “nomenclaturally unavailable” for these names name proposed as an unmodified vernacular word, not appropriately latinized (ICZN 1999, Article 11.3, Recommendation 11A); name not used as valid when proposed (ICZN 1999, Article 11.5); name first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym (ICZN 1999, Article 11.6); name published before 1931 without a description, a definition or an indication (Article 12.1); name proposed as a replacement name for an unavailable name (ICZN 1999, Article 12.2.3); name published after 1930 without a description, a definition or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1); name published after 1930 without a type species (ICZN 1999, Article 13.3); replacement name published after 1930, for a name without valid typification, without designating a type species (ICZN 1999, Article 13.3.1); new nominal genus or subgenus published in a combined description with a new species not marked by “gen. nov., spec. nov.” or an equivalent expression (ICZN 1999, Article 13.4); name published after 1999 and not explicitly indicated as intentionally new (ICZN 1999, Article 16.1); rejected alternative original spelling (ICZN 1999, Article 24.2.3); original spelling corrected in the same work (ICZN 1999, Article 32.5.1.1); incorrect subsequent spelling that is not in prevailing usage (ICZN 1999, Article 33.3).

The symbol for fossil (†) and the acronym for plate (pl.) were used when relevant. The following initials were necessary to distinguish different authors with an identical family F.M. Brown, K.W. Brown, W. Kirby, W.F. Kirby, J.E. LeConte, J.L. LeConte, P.H. Lucas, R. Lucas, W.J. MacLeay, W.S. MacLeay, G.S. Medvedev, L.N. Medvedev, G.-A. Olivier, E. Olivier, F. Soldati, L. Soldati, C.G. Thomson, J. Thomson, C.O. Waterhouse, F.H. Waterhouse, G.R. Waterhouse. Note that the honorary title “Tjan-Schansky” was added by the Emperor of All Russia Nikolay II to the surname of A.P. Semenov’s father (and all his family members) in 1906 (M. Nabozhenko, pers. comm.). We use the author name “Semenov” for works published up to the end of 1906 and “Semenov-Tjan-Shansky” for those published after 1906. We also follow Bessudnova (2012: 102) and use the author’s surname “Fischer” for G. Fischer’s works up to the end of 1832 and use “Fischer von Waldheim” from 1833, the year he became a member of the Russian nobility.

Original literature for species-group names (i.e., type species) was verified; however, those works are not included in the “References” section because they are not the principal focus of this article, to conserve space and because they are, for the most part, available in recent publications. When the author(s) of a scientific name is different from the author(s) of the publication in which the name was proposed, the authorship of the scientific name is given in the format “Gray in Griffith and Pidgeon, 1832” (ICZN, Article 50.1.1 and Recommendation 51E) and only the reference pertaining to the whole work is given in the “References” section. Titles of references using a non-Latin alphabet were translated in English.

The name Xylotinus, as used by Sturm (1826: 59) to establish Xylotinusflabellicornis Sturm, 1826, the type species of Rhipidandrus J.L. LeConte, 1862, is considered an incorrect subsequent spelling for Xyletinus Latreille, 1810 [Coleoptera: Anobiidae] and is therefore not included here as an available genus-group name in Tenebrionidae (see Schwarz and Barber 1914: 175).

Two new synonymies were proposed in the fascicle on Coleoptera of the ‘Nomenclator Zoologicus’ by Agassiz (1846a). These synonymies are credited here to “Erichson in Agassiz” since Agassiz, who was not a beetle expert, acknowledged the contribution of the Coleoptera specialist W.F. Erichson, on the fascicle's title page, in the production of his catalogue.

Classification

The classification presented in Table 2 is a synthesis starting from works on Tenebrionidae family-group names (e.g., Bouchard et al. 2005, 2011; Bouchard and Bousquet 2020a) with additional data published in recent regional catalogues (e.g., Matthews and Bouchard 2008; Bousquet et al. 2018; Iwan et al. 2020), on fossils (Nabozhenko 2019), as well as on the diversity, taxonomy and phylogenetics of various clades (e.g., Bremer and Lillig 2014; Aalbu et al. 2017, 2018; Kamiński et al. 2019a, b, c, 2021b; Matthews and Lawrence 2019; Lumen et al. 2020; Lillig and Bremer 2021; Gearner et al. 2021). Only nomenclaturally available names are included in Table 2 wherein the valid genus-group names are listed in alphabetical order under each valid family-group name and the synonyms (preceded by “=”) are given in chronological order. Valid subgenera are given following the acronym “SG”.

Distribution

The world’s major biogeographical realms, as defined by Olson et al. (2001), were used to record the overall distribution of each valid genus and subgenus (Table 2). The maps showing the borders of each realm (Figs 1–5) were produced with the free and open source software QGIS (version 3.10) using the layer Terrestrial Ecoregions of the World (WWF 2012).

Figure 2.: Closeup map showing the boundary between the Nearctic and Neotropic biogeographic realms. Countries with two biogeographic realms are named.

Figure 3.: Closeup map showing the boundary between the Palaearctic and Afrotropic biogeographic realms. Countries with two biogeographic realms are named.

Figure 4.: Closeup map showing the boundary between the Palaearctic and Indo-Malay biogeographic realms. Countries with two biogeographic realms are named.

Figure 5.: Closeup map showing the boundary between the Indo-Malay and Australasia biogeographic realms. Countries with two biogeographic realms are named.

Publication dates

We tried to establish the date of publication of works listed in this catalogue as accurately as possible. In the References section, publication dates are listed after each reference in square brackets “[]” and preceded by DP (i.e., date of publication), unless only the year was found. These dates are either specific dates of publication (e.g., extracted from the works themselves or the journal wrappers, title pages, footers) or are the earliest known publication dates (e.g., extracted from recording journals or accounts of society meetings). In the latter case, the date is preceded by the word “by.” Sources of information for the dates of publication from societies, recording journals, and works are mentioned below.

Societies

Acad Nat Sci Phil: Academy of Natural Sciences, Philadelphia (Philadelphia, USA) [Proceedings];

Acad Sci Fr: Académie des Sciences (Paris, France) [Compte Rendus Hebdomadaires];

Acad Sci St. Peters: Académie Impériale des Sciences de St.-Pétersbourg (Saint Petersburg, Russia) [Bulletin Scientifique; Bulletin de la Classe historico-philologique];

Amer Ant Soc: American Antiquarian Society (Worcester, USA) [Proceedings];

Amer Ent Soc: American Entomological Society (Philadelphia, USA) [Proceedings];

Amer Phil Soc: American Philosophical Society (Philadelphia, USA) [Proceedings];

Bost Soc Nat Hist: Boston Society of Natural History (Boston, USA) [Proceedings];

Ent Soc Lond: Entomological Society of London (London, UK) [Transactions, Proceedings];

Ent Ver Stettin: Entomologischer Verein zu Stettin (Stettin, Germany [now Poland]) [Vereinsangelegenheiten];

Nederl Ent Ver: Nederlandsche Entomologische Vereniging (Leiden, Netherlands) [Inhouds-Opgave van Werken];

Roy Soc Queensl: Royal Society of Queensland (Brisbane, Australia) [Proceedings];

Soc Ent Belg: Société Entomologique de Belgique (Brussels, Belgium) [Annales, Comptes-Rendus];

Soc Ent Fr: Société Entomologique de France (Paris, France) [Annales, Bulletin];

Soc Imp Nat Mosc: Société Impériale des Naturalistes de Moscou (Moscow, Russia) [Bulletin].

Recording journals

Allg Bibl Deutsch: Allgemeine Bibliographie für Deutschland (Leipzig, Germany);

Bibl Belg: Bibliographie de la Belgique (Brussels, Belgium);

Bibl Fr: Bibliographie de la France (Paris, France);

Bull Nord: Bulletin du Nord, journal scientifique et littéraire, des mémoires et notices, des analyses et extraits d’ouvrages nouveaux; des variétés et mélanges, des annonces bibliographiques, etc., etc. (Moscow, Russia);

Ent Nachr: Entomologische Nachrichten (Berlin, Germany);

Lit Centrbl: Literarisches Centralblatt für Deutschland (Leipzig, Germany);

Lit Ztg: Literarische Zeitung (Berlin, Germany);

Nat Nov: Naturae Novitates (Berlin, Germany);

Naturaliste: Le Naturaliste (Paris, France);

Pet Nouv Ent: Petites Nouvelles Entomologiques (Paris, France);

Rev Coleopt: Revue Coléoptérologique (Brussels, Belgium);

Zool Rec: Zoological Record (London, UK).

Works reporting dates of publication

The following works included important information for dating the Kielmeyer and Jäger (1835), Guérin-Méneville (1838, 1858), Lefèvre (1885, 1895), Sharp (1892a), Sclater (1893), Oshanin (1910), Wheeler (1912), Sherborn (1934), Duncan (1937), Peavot (1937), F.H. Waterhouse (1937), ICZN (1957, 1958, 1999), F.M. Brown (1964), Raphael (1970), Cowan (1976), Kerzhner (1984), Hayek (1989), Baker (1996), Evenhuis (1997a, b, 2002, 2003, 2012, 2015, 2019a, b), Nagel and Schmidlin (2014), Merkl et al. (2008), Bouchard et al. (2011), Fujioka (2011), Bousquet (2016a, b, 2017), Williams (2017).

Results

Overall diversity and distribution

The family Tenebrionidae contains 4122 nomenclaturally available genus-group names (see Table 2 and List of available genus-group names in Tenebrionidae Latreille, 1802 below). Of the 2307 valid genera, 33 are extinct. A total of 761 subgenera are currently used as valid. The subfamily Pimeliinae contains the highest number of valid genera (n = 578) followed by Stenochiinae (n = 394), Tenebrioninae (n = 349), Blaptinae (n = 300), Lagriinae (n = 273), Alleculinae (n = 231) and Diaperinae (n = 128). The other four subfamilies contain few genera (Phrenapatinae = 28, Zolodininae = 3, Kuhitangiinae = 2 and Nilioninae = 1). Twenty valid genera could not be confidently placed in a subfamily and are included here as Tenebrionidae incertae sedis. The Afrotropic biogeographical realm is the most diverse with 1022 valid extant genus-group taxa (755 genera, 267 subgenera; see Table 2). The Palaearctic, Indo-Malay, and Neotropic realms follow with 925 (545 genera, 380 subgenera), 610 (501, 109) and 518 (428, 90) valid extant genus-group taxa respectively. The lowest number of valid extant genus-group taxa are in Australasia with 348 (310 genera, 38 subgenera), the Nearctic with 279 (206, 73) and Oceania with 62 (52, 10).

List of available genus-group names in Tenebrionidae Latreille, 1802

Abantiades Fairmaire, 395 [M]. Type species [automatic]: Abantisaenescens Fairmaire, 1892, by monotypy. Status: junior synonym of Diphyrrhynchus Fairmaire, 1849 in Blaptinae: Opatrini: Heterotarsina. Note: replacement name for Abantis Fairmaire, 1892; junior homonym of Abantiades Herrich-Schäffer, 1856 [Lepidoptera].

Abantis Fairmaire, 109 [F]. Type Abantisaenescens Fairmaire, 1892, by monotypy. Status: junior synonym of Diphyrrhynchus Fairmaire, 1849 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Gebien (1938a: 407). Note: junior homonym of Abantis Hoppfer, 1855 [Lepidoptera].

Aberlencus Iwan, 559 [M]. Type Aberlencusangolensis Iwan, 2002, by original designation. Status: junior synonym of Angolositus Koch, 1955 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 90).

Abiga Guérin-Méneville, clxxxix [F]. Type Abigahumilis Guérin-Méneville, 1860, by subsequent designation (Löbl et al. 2008a: 40). Status: junior synonym of Scelosodis Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Guérin-Méneville (1862: 376).

Abigopsis Escalera, 277 [F]. Type Scelosodisustus Fairmaire, 1879, by subsequent designation (Gebien 1937a: 612). Status: valid genus in Pimeliinae: Tentyriini.

Ablapsis Reitter, 368 [F]. Type Blaps compressipes Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Acanthioides Fairmaire, 320 [F]. Type Acanthioidesasperula Fairmaire, 1894, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Acanthobas Gebien, 168, 175 [M]. Type Acanthobasangusticollis Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Acanthoblaps Reitter, 687 [F]. Type Acanthoblapsdentitibia Reitter, 1889, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gebien (1910b: 226).

Acanthocamaria Gebien, 26, 31 [F]. Type Acanthocamariabrunneoopaca Gebien, 1919, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Acanthomera Latreille, 580 [F]. Type Pimeliadentipes Fabricius, 1787, by monotypy. Status: senior synonym of Psorodes Dejean, 1834 in Tenebrioninae: Amarygmini. Note: junior homonym of Acanthomera Wiedemann, 1821 [Diptera].

Acanthomerus Agassiz, 2 [M]. Type species [automatic]: Pimeliadentipes Fabricius, 1787, by monotypy. Status: junior synonym of Psorodes Dejean, 1834 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Acanthomera Latreille, 1828, not in prevailing usage.

Acanthosternus Montrouzier, 289 [M]. Type Acanthosternushalorageos Montrouzier, 1860, by monotypy. Status: junior synonym of Diphyrrhynchus Fairmaire, 1849 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Bates (1872a: 97).

Acantophorus Billberg, 32 [M]. Type Erodiusgibbus Fabricius, 1775, by subsequent designation (Bouchard and Bousquet 2020b: 7). Status: junior synonym of Erodius Fabricius, 1775 in Pimeliinae: Erodiini. Synonymy: Bouchard and Bousquet (2020b: 7).

Acastus Péringuey, 177 [M]. Type Acastusagrestis Péringuey, 1896, by subsequent designation (Ardoin 1963a: 87). Status: junior synonym of Gonocnemis J. Thomson, 1858 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1913: 76).

Accanthopus Dejean, 71 [M]. Type fixed herein (ICZN 1999, Article 70.3) as Tenebriovelikensis Piller & Mitterpacher, 1783, misidentified as Tenebriocaraboides Linnaeus, 1758 in the original designation by monotypy in Dejean (1821). Status: valid genus in Tenebrioninae: Helopini: Enoplopodina. Note: the type species “Blaps caraboides Germ.” was first established by monotypy; Seidlitz (1895: 681) first noted that Tenebriocaraboides Linnaeus of Germar (1817) was identical to Helopsdentipes Rossi, 1790, currently a junior synonym of Tenebriovelikensis Piller & Mitterpacher, 1783; we follow currently accepted concepts (e.g., Löbl et al. 2008b: 246) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Tenebriocaraboides Linnaeus, 1758 is a valid species in the genus Cychrus Fabricius, 1794 [Coleoptera: Carabidae].

Acerogria Borchmann, 19, 137 [F]. Type Cerogriadohrni Borchmann, 1911, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Acestophanus Koch, 67 [M]. Type species [automatic]: Acestuselongatus Haag-Rutenberg, 1875, by subsequent designation (R. Lucas 1920: 69). Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Acestus Haag-Rutenberg, 1875.

Acestus Haag-Rutenberg, 4, 56 [M]. Type Acestuselongatus Haag-Rutenberg, 1875, by subsequent designation (R. Lucas 1920: 69). Status: senior synonym of Acestophanus Koch, 1950 in Pimeliinae: Adelostomini. Note: junior homonym of Acestus Leidy, 1851 [Annelida].

Achaemenes Bogatchev, 39 [M]. Type Achaemenesvillosus Bogatchev, 1949 (= Thripterabogatchevi Kwieton, 1982), by original designation. Status: senior synonym of Bogatshevia G.S. Medvedev & Iwan, 2006 in Pimeliinae: Pimeliini. Note: junior homonym of Achaemenes Stål, 1866 [Hemiptera].

Achanius Erichson, 118 [M]. Type Achaniusanthicoides Erichson, 1847, by monotypy. Status: valid genus and subgenus in Pimeliinae: Evaniosomini. Note: transferred from Edrotini (where it was previously placed by Doyen (1994: 500)) by Flores and Aballay (2015: 167).

Achariotheca Kaszab, 273 [F]. Type Achariothecabaloghi Kaszab, 1970, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Achora Pascoe, 279 [F]. Type Asidaserricollis Hope, 1843 (= Cestrinusobscurus Erichson, 1842), by monotypy. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Gemminger in Gemminger and Harold (1870: 1929, with Prionotus Mulsant & Rey, 1859, a junior synonym of Isopteron Hope, 1841); Matthews (1998: 788).

Achrostus Fairmaire, 256 [M]. Type Achrostusrufonitens Fairmaire, 1891, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Achthosus Pascoe, 42 [M]. Type Achthosuswestwoodii Pascoe, 1863, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Acidia Illiger, 79 [F]. Type species [automatic]: Pimeliareflexa Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Akis Herbst, 1799 in Pimeliinae: Akidini. Note: unjustified emendation of Akis Herbst, 1799, not in prevailing usage.

Acis Billberg, 32 [F]. Type species [automatic]: Pimeliareflexa Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Akis Herbst, 1799 in Pimeliinae: Akidini. Note: unjustified emendation of Akis Herbst, 1799, not in prevailing usage.

Acisba Dejean, 185 [F]. Type Tentyriacribrosa Besser, 1832 (= Pimeliapunctata Fabricius, 1798), by subsequent designation (Löbl et al. 2008a: 40). Status: junior synonym of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Solier (1835b: 288). Note: the original combination of the accepted name of the type species, Pimeliapunctata Fabricius, 1798, is a junior primary homonym of Pimeliapunctata Thunberg, 1787.

Acmoeus Fåhraeus, 293 [M]. Type Anomalipuselephas Fåhraeus, 1870, by monotypy. Status: junior synonym of Anomalipus Guérin-Méneville, 1831 in Blaptinae: Platynotini: Platynotina. Synonymy: Gebien (1938a: 409).

Aconobius Casey, 617 [M]. Type Conibiosomalaciniata Casey, 1892, by original designation. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Acontodactylus Desbrochers des Loges, 5 [M]. Type Cossyphusminutissimus Laporte, 1840, by original designation. Status: valid subgenus of Cossyphus G.-A. Olivier, 1791 in Lagriinae: Cossyphini.

Acritolagria Borchmann, 48, 98 [F]. Type Lagriafulvopilosa Fairmaire, 1887, by subsequent designation (Borchmann 1936: 166). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Acromaticus Koch, 143 [M]. Type Sepidiumstriatum Quensel, 1806 (= Sepidiumstriatum Thunberg, 1787), by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Acropachia Mäklin, 656 [F]. Type Acropachiabifoveolata Mäklin, 1875, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Acropteron Perty, 64 [N]. Type Acropteronrufipes Perty, 1832 (= Acropteryxrufipes Gistel, 1831), by subsequent designation (Hope 1841: 133). Status: junior synonym of Acropteryx Gistel, 1831 in Tenebrioninae: Acropteronini. Synonymy: Gistel (1857: 551, synonymy of the type species), Martins and Pereira (1966: 159, as “Acropteroxysrufipes Distl., 1831”).

Acropterum Agassiz, 6 [N]. Type species [automatic]: Acropteronrufipes Perty, 1832 (= Acropteryxrufipes Gistel, 1831), by subsequent designation (Hope 1841: 133). Status: junior synonym of Acropteryx Gistel, 1831 in Tenebrioninae: Acropteronini. Note: unjustified emendation of Acropteron Perty, 1832, not in prevailing usage.

Acropteryx Gistel, 308 [F]. Type Acropteryxrufipes Gistel, 1831, by original designation. Status: valid genus in Tenebrioninae: Acropteronini. Note: genus previously included in Erotylidae: Languriinae (see Leschen and Węgrzynowicz 1998: 234); reversal of precedence cannot be used to conserve the usage of Acropteron Perty, 1832 because Acropteryx was used as valid after 1899 (e.g., Blackwelder 1945: 427).

Acroschatia Wilke, 269 [F]. Type Microschatiarobusta Horn, 1893, by original designation. Status: junior synonym of Microschatia Solier, 1836 in Pimeliinae: Asidini. Synonymy: K.W. Brown and Doyen (1992: 546).

Actanorie Bates, 289 [M]. Type Camariaundaticollis Fairmaire, 1875, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Actizeta Pascoe, 214 [F]. Type Actizetaammobioides Pascoe, 1875 (= Actizetaalbata Pascoe, 1875), by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Actizetina.

Acutogria Merkl, 137 [F]. Type Acutogriafalcata Merkl, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Acutoodescelis Kaszab, 941, 951 [F]. Type Platyscelispunctatissima Fairmaire, 1886, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Adamus Iwan, 255 [M]. Type Platynotusmicrositoides Kaszab, 1975, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Adavius Mulsant & Rey, 126, 138 [M]. Type Adaviusclavipes Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Addia Lewis, 465 [F]. Type Addiascatebrae Lewis, 1894, by monotypy. Status: junior synonym of Tetraphyllus Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini. Synonymy: Masumoto (1982a: 23, probable); Ando (1991: 64, confirmed).

Adelina Dejean, 315 [F]. Type Cucujusplanus Fabricius, 1801, by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Adelium W. Kirby, 420 [N]. Type Adeliumcalosomoides W. Kirby, 1819, by subsequent designation (Meyer 1844: 180). Status: valid genus in Lagriinae: Adeliini. Note: the type species previously recognized for this genus was Carabusporcatus Fabricius, 1775, by subsequent designation by Lacordaire (1859b: 438).

Adelodemus Haag-Rutenberg, 100 [M]. Type Adelodemusasperulus Haag-Rutenberg, 1878 (= Cestrinussqualidus W.J. MacLeay, 1872), by monotypy. Status: valid genus in Lagriinae: Adeliini.

Adelonia Laporte, 221 [F]. Type Ulomafiliforme Laporte, 1840, by monotypy. Status: valid genus and subgenus in Lagriinae: Belopini.

Adelostoma Duponchel, 342 [N]. Type Adelostomasulcatum Duponchel, 1827, by monotypy. Status: valid genus and subgenus in Pimeliinae: Adelostomini.

Adelostomoides Carl, 24 [M]. Type Adelostomagrande Haag-Rutenberg, 1879, by monotypy. Status: junior synonym of Zarudnionymus Semenov-Tjan-Shansky & Bogatchev, 1947 in Pimeliinae: Adelostomini. Synonymy: Purchart (2007: 240).

Adelozotypus Kaszab, 224 [M]. Type Adelozotypusnovaecaledoniae Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Adeliini.

Adelphinops Reitter, 168, 169 [M]. Type Adelphinusordubadensis Reitter, 1890, by monotypy. Status: valid subgenus of Adelphinus Fairmaire & Coquerel, 1866 in Tenebrioninae: Helopini: Helopina.

Adelphinus Fairmaire & Coquerel, 44 [M]. Type Eutrapelasuturalis P.H. Lucas, 1847, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina.

Adelphus Dejean, 208 [M]. Type Helopsmarginatus Fabricius, 1792, by subsequent designation (Bousquet and Bouchard 2013a: 58). Status: senior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Synonymy: Chevrolat (1847b: 458). Note: nomen oblitum (see Bouchard and Bousquet, 6).

Adeps Gistel, 63 [M]. Type Amarygmuspaykullii Dalman, 1823, by original designation. Status: junior synonym of Tetraphyllus Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini. Synonymy: new synonym [PB]. Note: Adeps Gistel, 1857, and its replacement name Adepsion Strand, 1917, have been forgotten in the literature; the type species of Adeps is currently included in the genus Tetraphyllus Laporte & Brullé, 1831 and for that reason Gistel’s name is considered a junior synonym of Tetraphyllus; junior homonym of Adeps Gistel, 1848 [Crustacea].

Adepsion Strand, 90 [N]. Type species [automatic]: Amarygmuspaykullii Dalman, 1823, by original designation. Status: junior synonym of Tetraphyllus Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini. Synonymy: new synonym [PB]. Note: replacement name for Adeps Gistel, 1857.

Ades Guérin-Méneville, 277 [M]. Type Adeshemisphericus Guérin-Méneville, 1857, by monotypy. Status: valid genus in Diaperinae: Leiochrinini.

Adesmia Fischer, 153 [F]. Type Adesmialongipes Fischer, 1822 (= Pimeliaanomala Fischer, 1820), by monotypy. Status: valid genus and subgenus in Pimeliinae: Adesmiini.

Adesmina Reitter, 5, 29 [F]. Type Adesmiaarabica Reitter, 1916, by subsequent designation (Löbl et al. 2008b: 122). Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Adisema Borchmann, pl. 7 [F]. Type Disemaambigua Mäklin, 1875, by monotypy. Status: junior synonym of Nemostiromorpha Pic, 1917 in Lagriinae: Lagriini: Statirina.

Adonicus Fairmaire, 258 [M]. Type Adonicuspurpuripennis Fairmaire, 1891, by monotypy. Status: junior synonym of Alcyonotus Pascoe, 1882 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1905: 255).

Adordanea Reitter, 229 [F]. Type Microderasubseriata Reitter, 1889, by monotypy. Status: junior synonym of Dordanea Reitter, 1887 in Pimeliinae: Tentyriini. Synonymy: Kaszab (1966: 304), G.S. Medvedev (1990: 112).

Adoryacus Koch, 34 [M]. Type Stizopusrotundicollis Fairmaire, 1897, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Adosogria Borchmann, 16, 63 [F]. Type Lagriabennigseni Borchmann, 1909, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Adynata Fåhraeus, 330 [F]. Type Adynatatricolor Fåhraeus, 1870, by subsequent designation (Borchmann 1936: 23). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Aeanes Champion, 566 [M]. Type Aeanesangusticollis Champion, 1893, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Aediatorix Bates, 315 [M]. Type Aediatorixjansoni Bates, 1868, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Aemymone Bates, 314 [F]. Type Goniaderacariosa Bates, 1868, by original designation. Status: valid subgenus of Goniadera Perty, 1832 in Lagriinae: Goniaderini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Aequigula Penrith, 20, 23 [F]. Type Alogeniushughesae Penrith, 1979, by original designation. Status: valid subgenus of Alogenius Gebien, 1910 in Pimeliinae: Adesmiini.

Aeschrocera Chen & Chou, 265 [F]. Type Cerogriabrunneocollis Chen & Chou, 1996 (= Cerogriadiversicornis Pic, 1933), by original designation. Status: junior synonym of Cerogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina. Synonymy: Merkl (2007: 262).

Aesthetus C.O. Waterhouse, 552 [M]. Type Aesthetustuberculatus C.O. Waterhouse, 1890, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Aesymnus Champion, 168 [M]. Type Aesymnusnitidus Champion, 1886, by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Aethales Dejean, 180 [M]. Type Epitragusbrunnicornis Latreille, 1811, by monotypy. Status: junior synonym of Epitragus Latreille, 1802 in Pimeliinae: Epitragini. Synonymy: Bousquet and Bouchard (2013a: 43). Note: as mentioned by Bousquet and Bouchard (2013a: 43) the type species Epitragusbrunnicornis Latreille was listed as a species incertae sedis in the genus Epitragus Latreille, 1802 by Freude (1967: 176).

Aethalides Bates, 50 [M]. Type Aethalidespunctipennis Bates, 1873, by monotypy. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 473).

Aethyssius Pascoe, 45 [M]. Type species [automatic]: Atractusviridis Boisduval, 1835, by subsequent designation (Duponchel and Chevrolat 1841: 312). Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324). Note: replacement name for Atractus Boisduval, 1835.

Afghanillus Kaszab, 1 [M]. Type Afghanillusklapperichi Kaszab, 1960, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Afghanopachys Kwieton, 29, 30 [M]. Type Pachyscelishaarlovi Gridelli, 1954, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Afrasida Wilke, 260 [F]. Type Asidacaryophyllea Wiedemann, 1823, by original designation. Status: valid genus and subgenus in Pimeliinae: Asidini.

Afrethas Koch, 23 [M]. Type Stenosispusillima Fairmaire, 1888, by original designation. Status: valid subgenus of Anethas Jakobson, 1924 in Pimeliinae: Stenosini: Stenosina.

Afrinus Fairmaire, 189 [M]. Type Afrinusgrandicornis Fairmaire, 1888, by subsequent designation (R. Lucas 1920: 79). Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: the designation of Afrinusstriolifrons Fairmaire, 1888 as the type species of Afrinus Fairmaire, 1888 by Koch (1950b: 329) is invalid (ICZN 1999, Article 69.1).

Afrobyrsax Ardoin, 894 [M]. Type Afrobyrsaxgirardi Ardoin, 1973, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Afrohelops Schawaller, 75 [M]. Type Afrohelopskenyaensis Schawaller, 2012, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: removed from the tribe Helopini and placed as Tenebrionidae incertae sedis by Nabozhenko (2018: 183).

Afrolaena Endrödy-Younga & Schawaller, 9, 11 [F]. Type Afrolaenatibialis Endrödy-Younga & Schawaller, 2002, by original designation. Status: valid genus in Lagriinae: Laenini.

Afronosis G.S. Medvedev, 849, 859 [F]. Type Stenosisciliaris Gebien, 1920, by original designation. Status: valid subgenus of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina. Note: Stenosisleontjevi G.S. Medvedev, 1995 was listed as a second type species of this genus in the original publication (G.S. Medvedev, 859); we act as First Revisers and select Stenosisciliaris Gebien, 1920 as the type species for Afronosis G.S. Medvedev, 1995.

Afrostrongylium Robiche, 83 [N]. Type Strongyliumfrancoisi Robiche, 2019, by original designation. Status: valid subgenus of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini.

Afrotagalus Gebien, 111, 120 [M]. Type Afrotagaluseidmanni Gebien, 1942, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Afrotenebrio Gridelli, 223, 230 [M]. Type Tenebrioguineensis Imhoff, 1843, by original designation. Status: valid subgenus of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini.

Agastenes R. Lucas, 79 [M]. Type species [automatic]: Agastheneswestwoodi Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: unjustified emendation of Agasthenes Bates, 1873.

Agasthenes Bates, 352 [M]. Type Agastheneswestwoodi Bates, 1873, by monotypy. Status: senior synonym of Agastenes R. Lucas, 1920 in Tenebrioninae: Heleini: Cyphaleina. Note: junior homonym of Agasthenes Förster, 1869 [Hymenoptera].

Agelarches Gistel, x [M]. Type species [automatic]: Pimeliaangulata Fabricius, 1775, by subsequent designation (Hope 1841: 118). Status: junior synonym of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Pimelia Fabricius, 1775.

Agissopterus Fairmaire, 513 [M]. Type Agissopterussemipunctatus Fairmaire, 1884, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Aglypta Gebien, 329 [F]. Type Aglyptaoctocostata Gebien, 1908, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Agnaptoria Reitter, 364, 372 [F]. Type Agnaptoriarubripes Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Agonopus Gebien, 110 [M]. Type Gonopuspuncticollis Solier, 1848, by subsequent designation (Endrödy-Younga 2000: 7). Status: valid subgenus of Gonopus Latreille, 1828 in Blaptinae: Platynotini: Platynotina.

Agraecus Fairmaire, 21 [M]. Type Agraecuschalcoides Fairmaire, 1900, by monotypy. Status: junior synonym of Carabelops Fairmaire, 1899 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1956b: 90).

Agroblaps Motschulsky, 531 [F]. Type Blaps fatidica Sturm, 1807 (= Blaps lethifera Marsham, 1802), by subsequent designation (Nabozhenko 2008: 35). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860).

Agymnonyx Gebien, 325, 328 [M]. Type Agymnonyxsulciventris Gebien, 1921, by subsequent designation (Gebien 1942a: 334). Status: valid genus in Stenochiinae: Cnodalonini.

Ahexaroptrum Kaszab, 291 [N]. Type Ahexaroptrumhumeridens Kaszab, 1960, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Ainu Lewis, 479 [M]. Type Ainutenuicornis Lewis, 1894, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Akis Herbst, 124 [F]. Type Pimeliareflexa Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: valid genus in Pimeliinae: Akidini.

Alaephus Horn, 344, 346 [M]. Type Alaephuspallidus Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Vacronini.

Alaetrinus Iwan, 14, 24 [M]. Type Tenebriopullus Sahlberg, 1823, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Alagria Borchmann, 50, 183 [F]. Type species [automatic]: Lagriostirahispida Kolbe, 1902 (= Lagriasubseriata Reitter, 1880), by subsequent designation (Borchmann 1936: 224). Status: valid genus in Lagriinae: Lagriini: Lagriina. Note: replacement name for Lagriostira Kolbe, 1902.

Alaudes Horn, 361 [M]. Type Alaudessingularis Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Cnemeplatiini: Alaudina.

Alcinoe Ménétriés, 230 [F]. Type Alcinoehelopioides Ménétriés, 1849, by monotypy. Status: senior synonym of Alcinoeta Strand, 1929 in Pimeliinae: Tentyriini. Note: junior homonym of Alcinoe Rang, 1828 [Ctenophora].

Alcinoeta Strand, 23 [F]. Type species [automatic]: Alcinoehelopioides Ménétriés, 1849, by monotypy. Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Alcinoe Ménétriés, 1849.

Alcmeonis Bates, 270 [F]. Type Alcmeonispulchra Bates, 1868, by monotypy. Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324, with Aethyssius Pascoe, 1863, a junior synonym of Lepturidea Fauvel, 1862).

Alcyonotus Pascoe, 35 [M]. Type Alcyonotusiridescens Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Alegoria Laporte, 221 [F]. Type Alegoriadilatata Laporte, 1840, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Alethia Champion, 417 [F]. Type Alethiasallaei Champion, 1888, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Alhuena Kulzer, 912 [F]. Type Alhuenapenai Kulzer, 1956 (= Peltolobuswaterhousei Bates, 1873), by original designation. Status: junior synonym of Peltolobus Lacordaire, 1859 in Pimeliinae: Trilobocarini. Synonymy: Kulzer (1962: 100).

Alienophloeus Bremer, 45 [M]. Type Corticeuspumilio Bremer, 2018, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Alienoplonyx Bremer, 59 [M]. Type Alienoplonyxalleni Bremer, 2019, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: we act as First Revisers and reject the alternative original spelling Alienolonyx, used by Bremer (2019: 60).

Allardius Ragusa, 130 [M]. Type Parablopsoculatus Baudi di Selve, 1876, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Allecula Fabricius, 21 [F]. Type Cistelamorio Fabricius, 1787, by subsequent designation (Duponchel 1840: 283). Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Alleculina Pic, 246 [F]. Type Synalleculamacroceps Pic, 1922, by original designation. Status: valid subgenus of Allecula Fabricius, 1801 in Alleculinae: Alleculini: Alleculina.

Alleculodes Borchmann, 335 [F]. Type Alleculodesdiscrepans Borchmann, 1925, by original designation. Status: junior synonym of Bolbostetha Fairmaire, 1896 in Alleculinae: Alleculini: Alleculina. Synonymy: Borchmann (1941a: 23).

Alleculopsis Semenov, 609 [F]. Type Alleculopsisdeserticola Semenov, 1894, by original designation. Status: valid subgenus of Mycetocharina Seidlitz, 1890 in Alleculinae: Alleculini: Alleculina.

Allocera Borchmann, 49, 175 [F]. Type Porrolagriasubaenea Borchmann, 1908, by subsequent designation (Borchmann 1936: 227). Status: senior synonym of Cerostira Borchmann, 1942 in Lagriinae: Lagriini: Lagriina. Note: junior homonym of Allocera Sichel, 1866 [Hymenoptera].

Allocossyphodes Andreae, 204, 215 [M]. Type Cossyphodesarnoldi Brauns, 1925, by original designation. Status: junior synonym of Cossyphodes Westwood, 1851 in Pimeliinae: Cossyphodini: Cossyphodina. Synonymy: Schawaller (2013c: 362, implied by inclusion of Cossyphodesarnoldi Brauns, 1925 in Cossyphodes Westwood, 1851 without use of a subgenus rank).

Allodengitha Bogatchev, 57 [F]. Type Allodengithadeserta Bogatchev, 1963, by original designation. Status: junior synonym of Alcinoeta Strand, 1929 in Pimeliinae: Tentyriini. Synonymy: G.S. Medvedev (1989: 798).

Allogria Borchmann, 48, 100 [F]. Type Allogriaspinosa Borchmann, 1916, by subsequent designation (Borchmann 1936: 74). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Allomyladion Bogatchev, 626 [N]. Type Penthicusporicollis Reitter, 1896, by original designation. Status: valid subgenus of Penthicus Faldermann, 1836 in Tenebrioninae: Opatrini: Opatrina.

Allopezus Gebien, 504 [M]. Type Allopezusmiritarsis Gebien, 1922, by monotypy. Status: junior synonym of Asbolodomimus Pic, 1921 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1948: 543).

Allophasia Pascoe, 351 [F]. Type Allophasiafryi Pascoe, 1871, by monotypy. Status: junior synonym of Diaperis Geoffroy, 1762 in Diaperinae: Diaperini: Diaperina. Synonymy: Triplehorn and Brendell (1985: 14).

Allophylax Bedel, 177 [M]. Type Phylaxlittoralis Mulsant, 1854 (= Opatrumpicipes G.-A. Olivier, 1812), by subsequent designation (Gebien 1938a: 412). Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina. Note: replacement name for Neophylax Bedel, 1906.

Allostrongylium Kolbe, 364 [N]. Type Allostrongyliumsilvestre Kolbe, 1896, by present designation. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 539, with Anomoearthrum Mäklin, 1867, a junior synonym of Strongylium W. Kirby, 1819).

Allotadzhikistania Bogatchev, 43 [F]. Type Allotadzhikistaniacomata Bogatchev, 1960, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Allotriocochabambia Faúndez, Rider & Carvajal, 595 [F]. Type species [automatic]: Cochabambiakulzeri Marcuzzi, 1985, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: replacement name for Cochabambia Marcuzzi, 1985; Marcuzzi (1985: 185) mentioned that his new genus Cochabambia could not be placed in “any given Neotropical tribe so far known” and is therefore included here in Tenebrionidae incertae sedis under its replacement name.

Alobates Motschulsky, 25 [M]. Type Tenebriopensylvanicus DeGeer, 1775, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Alogenius Gebien, 91 [M]. Type species [automatic]: Metriopusfavosus Erichson, 1843, by subsequent designation (Gebien 1937a: 661). Status: valid genus and subgenus in Pimeliinae: Adesmiini. Note: replacement name for Pedionomus Haag-Rutenberg, 1875.

Alogista Fåhraeus, 318 [F]. Type Alogistaabnormis Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Alogistopsis Borchmann, 53 [F]. Type Alogistopsispilistriata Borchmann, 1943, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Alphasida Escalera, 380 [F]. Type Asidagaditana Escalera, 1905 (= Alphasidatypica Gebien 1937), by subsequent designation (Casey 1912: 78). Status: valid genus and subgenus in Pimeliinae: Asidini.

Alphitobius Stephens, 19 [M]. Type Helopspicipes Panzer, 1794 (= Opatrumlaevigatum Fabricius, 1781), by monotypy (see ICZN 1975, Opinion 1039). Status: valid genus in Tenebrioninae: Alphitobiini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1975, Opinion 1039).

Alphitophagus Stephens, 12 [M]. Type Alphitophagusquadripustulatus Stephens, 1832 (= Diaperisbifasciata Say, 1824), by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

†Alphitopsis Kirejtshuk, Nabozhenko & Nel, 549 [F]. Type Alphitopsisinitialis Kirejtshuk, Nabozhenko & Nel, 2011, by original designation. Status: valid genus in Tenebrioninae: Alphitobiini. Note: described from Lower Cretaceous deposits (China).

Altes Pascoe, 288, 290 [M]. Type Chartopteryxbinodosus Pascoe, 1862, by original designation. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Gemminger in Gemminger and Harold (1870: 1994, with Chartopteryx Westwood, 1841, a synonym of Cyphaleus Westwood, 1841); Matthews (1992: 490).

Altiprosodes G.S. Medvedev, 580 [M]. Type Prosodeskuhistanica G.S. Medvedev, 1996, by original designation. Status: junior synonym of Megaprosodes Reitter, 1909 in Blaptinae: Blaptini: Prosodina. Synonymy: G.S. Medvedev (2001: 83). Note: originally described as a section within a subgenus.

Alymon Pascoe, 484 [M]. Type Alymonprolatus Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Amachla Koch, 117 [F]. Type Machlasulcicollis Fåhraeus, 1871, by original designation. Status: valid genus in Pimeliinae: Asidini.

Amarantha Motschulsky, 141 [F]. Type Amaranthaviridis Motschulsky, 1860, by monotypy. Status: senior synonym of Metaclisa Jacquelin du Val, 1861 in Tenebrioninae: Metaclisini. Synonymy: Lewis (1891: 70). Note: nomen oblitum (see Bouchard et al. 2007: 393).

Amaropsis Champion, 567 [F]. Type Amaropsisannulicornis Champion, 1893, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Amarosoma Redtenbacher, 131 [N]. Type Amarosomasimulans Redtenbacher, 1868, by monotypy. Status: junior synonym of Pheloneis Pascoe, 1866 in Lagriinae: Adeliini. Synonymy: Pascoe (1876: 52).

Amarsenes Bates, 297 [M]. Type Tetraphyllusoblongocamelus Fairmaire, 1877, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Amarygmimus Bates, 354 [M]. Type Amarygmimusduboulayi Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Amarygmomimus Rye, 290 [M]. Type species [automatic]: Amarygmimusduboulayi Bates, 1873, by monotypy. Status: junior synonym of Amarygmimus Bates, 1873 in Tenebrioninae: Heleini: Cyphaleina. Note: unjustified emendation of Amarygmimus Bates, 1873, not in prevailing usage.

Amarygmus Dalman, 60 [M]. Type Chrysomelamicans Fabricius, 1794, by subsequent designation (Gebien 1921a: 410). Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Amathobius Gebien, 129 [M]. Type Amathobiusglyptopterus Gebien, 1920, by subsequent designation (Koch 1963: 42). Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Amatodes Dejean, 189 [F]. Type Pimeliagemmata Fabricius, 1801, by monotypy. Status: valid genus and subgenus in Blaptinae: Pedinini: Helopinina. Note: the original combination of the name of the type species, Pimeliagemmata Fabricius, 1801, is a junior primary homonym of Pimeliagemmata Herbst, 1799.

†Amberophlus Novák & Háva, 128 [M]. Type Amberophlusniger Novák & Háva, 2019, by original designation. Status: valid genus in Alleculinae: Cteniopodini. Note: described from Eocene Baltic amber.

Ambigatus Fairmaire, 246 [M]. Type Ambigatusrufonitens Fairmaire, 1892, by subsequent designation (R. Lucas 1920: 87). Status: valid subgenus of Achanius Erichson, 1847 in Pimeliinae: Evaniosomini.

Amblycara Fairmaire, cxlvii [N]. Type Amblycarabiskrense Fairmaire, 1893 (= Melancrusalutaceum Fairmaire, 1875), by monotypy. Status: senior synonym of Amblycarenum Gebien, 1910 in Pimeliinae: Tentyriini. Note: junior homonym of Amblycara Bergroth, 1891 [Hemiptera].

Amblycarenum Gebien, 77 [N]. Type species [automatic]: Amblycarabiskrense Fairmaire, 1893 (= Melancrusalutaceum Fairmaire, 1875), by monotypy. Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Amblycara Fairmaire, 1893.

Amblychirus Koch, 87 [M]. Type Trigonopusbrevior Fairmaire, 1897, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Amblycyphus Motschulsky, 401 [M]. Type Amblycyphusasperatus Motschulsky, 1870 (= Centriopterapectoralis Blaisdell, 1921), by monotypy. Status: junior synonym of Cryptoglossa Solier, 1837 in Pimeliinae: Cryptoglossini. Synonymy: Aalbu et al. (1995: 483).

Amblyptera Solier, 195 [F]. Type Pimeliascabrosa Solier, 1836, by subsequent designation (Viñolas and Cartagena 2005: 267). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Amblypteraca Mas-Peinado, Buckley, Ruiz & García-París, 543 [F]. Type Pimeliarugosa Fabricius, 1792 (= Pimeliafairmairii Kraatz, 1865), by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Note: we act as First Revisers and reject the alternative original spelling Amplypteraca, used by Mas-Peinado et al. (2018: 531, 543).

Amblysphagus Fairmaire, 16 [M]. Type Amblysphaguspachyderus Fairmaire, 1896, by monotypy. Status: valid genus in Blaptinae: Opatrini: Neopachypterina.

Amenophis J. Thomson, 93 [M]. Type Amenophisfairmairei J. Thomson, 1858, by subsequent designation (Gebien 1941: 342). Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Ametrocera Fåhraeus, 260 [F]. Type Ametroceraaurita Fåhraeus, 1870, by subsequent designation (R. Lucas 1920: 88). Status: valid genus in Blaptinae: Pedinini: Helopinina.

Amiantus Fåhraeus, 279 [M]. Type Amiantusgibbosus Fåhraeus, 1870, by subsequent designation (Haag-Rutenberg 1871: 45). Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Amicrodera Kaszab, 282, 292 [F]. Type Microderalindbergi Kaszab, 1966, by original designation. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Ammidanemia Reitter, 132 [F]. Type Anemiafausti Solsky, 1882 (= Cheirodesbrevicollis Wollaston, 1864), by subsequent designation (G.S. Medvedev 1990: 212). Status: junior synonym of Pseudanemia Wollaston, 1864 in Tenebrioninae: Melanimonini. Synonymy: Ardoin (1971: 359, through placement of the type species in the subgenusPseudanemia Wollaston, 1864).

Ammidium Erichson, 250 [N]. Type Ammidiumciliatum Erichson, 1843, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Ammobius Guérin-Méneville, 121 [M]. Type Ammobiusrufus Guérin-Méneville, 1844, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6); the original combination of the name of the type species has been given as “Trachyscelisrufus Guérin-Méneville, 1844” in recent literature; however, we use the combination Ammobiusrufus here as it was used by Guérin-Méneville (1844: 560) in his ‘Table alphabétique des genres, sous-genres et espèces figurés ou décrits dans l’iconographie du règne animal. Insectes’.

Ammocera Borchmann, 5 [F]. Type Ammoceraunicolor Borchmann, 1941, by original designation. Status: valid subgenus of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina.

Ammodoides Lesne, 227, 233 [M]. Type Arthrodeislateripunctatus Fairmaire, 1890, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Ammodonus Mulsant & Rey, 141, 143 [M]. Type Opatrumfossor J.L. LeConte, 1847, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina. Note: transferred from Ammobiina by Lumen et al. (2020: 341).

Ammogiton Peyerimhoff, 326 [M]. Type Ammogitonviberti Peyerimhoff, 1920, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Ammophorus Guérin-Méneville, pl. 4 [M]. Type Ammophorusperuvianus Guérin-Méneville, 1831, by monotypy. Status: valid genus in Tenebrioninae: Scotobiini.

Ammophtorus Lacordaire, 284 [M]. Type species [automatic]: Ammobiusrufus Guérin-Méneville, 1844, by monotypy. Status: junior synonym of Ammobius Guérin-Méneville, 1844 in Blaptinae: Opatrini: Ammobiina. Note: unnecessary replacement name for Ammobius Guérin-Méneville, 1844.

Ammotrypes Fairmaire, 194 [M]. Type Ammotrypescrenulicollis Fairmaire, 1879, by monotypy. Status: junior synonym of Eurycaulus Fairmaire, 1868 in Blaptinae: Opatrini: Sclerina. Synonymy: Grimm (2005: 9); Löbl and Smetana (2010: 33).

Ammozoides Kaszab, 91 [M]. Type Ammozoumvalidicorne Reitter, 1900, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Ammozoum Semenov, 352 [N]. Type Ammozoumhyalinum Semenov, 1891, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Amnodeis Miller, 117 [M]. Type Amnodeisgrandis Miller, 1858, by subsequent designation (R. Lucas 1920: 90). Status: valid genus in Pimeliinae: Erodiini.

Amorphochirus Gebien, 339 [M]. Type Pycnocerushercules Fairmaire, 1884, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Amorphopoda Fåhraeus, 320 [F]. Type Amorphopodaelateroides Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Amphianax Bates, 350 [M]. Type Amphianaxsubcoriaceus Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Amphidora Eschscholtz, 9 [F]. Type Amphidoralittoralis Eschscholtz, 1829, by monotypy. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini. Note: Doyen and Lawrence (1979: 367) synonymized this genus with Eleodes Eschscholtz, 1829; however, Smith and Johnston in Bousquet et al. (2018: 143) used Amphidora as a valid subgenus of Eleodes.

Amphithrix Español, 306 [F]. Type Amphithrixpeyerimhoffi Español, 1952, by original designation. Status: senior synonym of Amphithrixoides Bouchard & Löbl, 2008 in Blaptinae: Opatrini: Ammobiina. Note: junior homonym of Amphithrix Ragonot, 1893 [Lepidoptera].

Amphithrixoides Bouchard & Löbl, 39 [M]. Type species [automatic]: Amphithrixpeyerimhoffi Español, 1952, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: replacement name for Amphithrix Español, 1952.

Anacardiosis Endrödy-Younga, 210, 221 [F]. Type Cardiosishamiltonuli Koch, 1969, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Anachayus Bouchard & Bousquet, new replacement name [M]. Type species [automatic]: Chemolanusvillosipes Fairmaire, 1884, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Chatanayus Ardoin, 1957.

Anacycus Fairmaire, 33 [M]. Type Dietysusnavicularis Fairmaire, 1894, by subsequent designation (Löbl et al. 215). Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Blair (1929a: 245; with Elixota Pascoe, 1866, a junior synonym of Amarygmus Dalman, 1823).

Anadischidus Kolbe, 241 [M]. Type Nyctobatesiphthinoides Quedenfeldt, 1885, by monotypy. Status: junior synonym of Deriles Motschulsky, 1872 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1911a: 441).

Anaedes Agassiz, 20 [M]. Type species [automatic]: Anaeduspunctatissimus Blanchard, 1842, by monotypy. Status: junior synonym of Anaedus Blanchard, 1842 in Lagriinae: Goniaderini. Note: unjustified emendation of Anaedus Blanchard, 1842, not in prevailing usage.

Anaedus Blanchard, pl. 14 [M]. Type Anaeduspunctatissimus Blanchard, 1842, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Anamenederes Koch, 68 [M]. Type Menederesdigitatus Koch, 1954, by original designation. Status: valid subgenus of Menederes Solier, 1848 in Blaptinae: Platynotini: Eurynotina.

Anamphidora Casey, 330 [F]. Type Anamphidoraparvula Casey, 1924, by original designation. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Anarmostodera Fairmaire, 114 [F]. Type Anarmostoderacrassicornis Fairmaire, 1897, by monotypy. Status: valid genus in Tenebrioninae: Praeugenini.

Anatolica Eschscholtz, 5, 7 [F]. Type Tentyriasubquadrata Tauscher, 1812, by subsequent designation (Gebien 1937a: 599). Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Anatrum Reichardt, 84, 209 [N]. Type Anatrumsongoricum Reichardt, 1936, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Anausis Bates, 355 [F]. Type Anausismacleayi Bates, 1873, by monotypy. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 490).

Anaxius Fåhraeus, 307 [M]. Type Anaxiusobesus Fåhraeus, 1870, by monotypy. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Anaxo Bates, 272 [F]. Type Anaxobrevicornis Bates, 1868, by monotypy. Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324, with Aethyssius Pascoe, 1863, a junior synonym of Lepturidea Fauvel, 1862).

Anchomma J.L. LeConte, 63 [N]. Type Anchommacostatum J.L. LeConte, 1858, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina. Note: new placement [RLA], previously included in Pimeliinae: Anepsiini; genus originally described in Tenebrionoidea: Zopheridae: Colydiinae.

Anchophthalmops Koch, 173 [M]. Type Anchophthalmopsbrevipleurum Koch, 1956, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Anchophthalmus Gerstaecker, 533 [M]. Type Anchophthalmussilphoides Gerstaecker, 1854, by subsequent designation (Gebien 1938a: 298). Status: valid genus and subgenus in Blaptinae: Platynotini: Platynotina.

Ancylopoma Pascoe, 354 [N]. Type Ancylopomapunctigerum Pascoe, 1871, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: see Bouchard and Bousquet (2020a: 102) for comments regarding the placement of this genus and its associated family-group name.

Andocamaria Masumoto, 230, 232 [F]. Type Campsiomorphaformosana Pic, 1930, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Andremiopsis Chatanay, 406 [F]. Type Andremiopsiscostata Chatanay, 1913, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Andremius Fairmaire, 364 [M]. Type Andremiuscrispatus Fairmaire, 1903, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Andrimus Casey, 155 [M]. Type Cteniopusmurrayi J.L. LeConte, 1866, by subsequent designation (R. Lucas 1920: 96). Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Androchirus J.L. LeConte, 244 [M]. Type Cistelafuscipes Melsheimer, 1846 (= Cistelaerythropa W. Kirby, 1837), by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Androsus Gebien, 325, 386 [M]. Type Chariothecaviolacea Pascoe, 1887, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Anebacis Peyerimhoff, 4, 15 [F]. Type Tentyriacribricollis Fairmaire, 1875, by subsequent designation (Löbl et al. 2008a: 40). Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Anectus Horn, 399 [M]. Type Anectusvestitus Horn, 1867, by monotypy. Status: valid genus in Pimeliinae: Branchini.

Anemia Laporte, 218 [F]. Type Anemiagranulata Laporte, 1840, by monotypy. Status: junior synonym of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini. Synonymy: Marseul (1863: 183); Spilman (1973: 41).

Anemiadena Bouchard & Bousquet, new subgenus [F]. Type Anemiaconvexa Gestro, 1881, by present designation. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini. Note: Ardoin (1971: 362, 402) introduced the new subgenus name Anemiadena for two nominal species, but unfortunately did not designate a type species; the fact that Anemiaconvexa Gestro, 1881 was listed as the “type” of Anemiadena in the Zoological Record for the year 1971 (Anonymous in Commonwealth Institute of Entomology 1975) does not represent a valid type species designation since the nomenclatural act is anonymous (ICZN 1999, Article 14); the subgenusAnemiadena, which has been treated as valid since 1971, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Anemiaconvexa Gestro, 1881 as type species and referring to Ardoin (1971: 362) for the character states that characterise and differentiate Anemiadena.

Anephyctus Fairmaire, 257 [M]. Type Anephyctushirtulus Fairmaire, 1891, by monotypy. Status: junior synonym of Miltoprepes Gerstaecker, 1871 in Tenebrioninae: Praeugenini. Synonymy: Gridelli (1939: 78, through placement of the type species in Miltoprepes Gerstaecker, 1871).

Anepsius J.L. LeConte, 147 [M]. Type Anepsiusdelicatulus J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Anepsiini.

Anethas Jakobson, 242 [M]. Type species [automatic]: Pseudethaslongiceps Fairmaire, 1898, by monotypy. Status: valid genus and subgenus in Pimeliinae: Stenosini: Stenosina. Note: replacement name for Pseudethas Fairmaire, 1898.

Angoleantus Koch, 128, 133 [M]. Type Rhammatodesstriatulus Koch, 1941, by original designation. Status: valid subgenus of Rhammatodes Haag-Rutenberg, 1876 in Pimeliinae: Tentyriini.

Angolositus Koch, 448 [M]. Type Angolositussadabandeirus Koch, 1955, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Aniara Melsheimer, 139 [F]. Type Ulomapiceum Melsheimer, 1846, by monotypy. Status: senior synonym of Eutochia J.L. LeConte, 1862 in Tenebrioninae: Ulomini. Note: name previously attributed to “Lacordaire, 1859” in the literature; junior homonym of Aniara Hope, 1838 [Coleoptera: Carabidae].

Aniarus Gemminger in Gemminger and Harold, 1964 [M]. Type species [automatic]: Ulomapiceum Melsheimer, 1846, by monotypy. Status: junior synonym of Eutochia J.L. LeConte, 1862 in Tenebrioninae: Ulomini. Note: unjustified emendation of Aniara Melsheimer, 1853, not in prevailing usage.

Anisocara Gebien, 101 [N]. Type Anisocaragynandromorphum Gebien, 1925, by monotypy. Status: junior synonym of Platydema Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Synonymy: Schawaller (2004: 5).

Anisocerus Faldermann, 39 [M]. Type Anisocerustristis Faldermann, 1837, by monotypy. Status: senior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Note: junior homonym of Anisocerus Audinet-Serville, 1835 [Coleoptera: Cerambycidae].

Anisophaedis Ando, 107 [M]. Type Anisophaedisohkurai Ando, 1993, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Anisosis Deyrolle, 81, 232 [F]. Type Anisosiscaudata Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Note: the alternative original spelling Urosis, used by Deyrolle (1867: 81), was rejected by Dallas (1868: 265–266) who acted as First Reviser.

Anisostira Borchmann, 296 [F]. Type Anisostiravaricolor Borchmann, 1915 (= Macrolagriarugipennis Lewis, 1896), by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Annamosdara Kaszab, 3, 30 [F]. Type Annamosdaramultidentata Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Anobriomaia Kaszab, 67 [F]. Type Anobriomaiasulcata Kaszab, 1941, by original designation. Status: junior synonym of Foochounus Pic, 1921 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 134).

Anodesis Solier, 508, 594 [F]. Type Anodesiscleryi Solier, 1834, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Anognathena Ando in Ando et al., 2017: 148 [F]. Type Anognathenaneraida Ando, 2017, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Anomalipus Guérin-Méneville, pl. 29 [M]. Type Blapsdentipes Fabricius, 1794, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina. Note: nomen protectum (see Bouchard and Bousquet 2020a: 101, 132); unjustified emendation of the original spelling Anomalipes, introduced by Lacordaire (1859a: 257), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1), see Bouchard et al. (2005: 511).

Anommabates Koch, 84 [M]. Type Anommabatespauliani Koch, 1956, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Anomoearthrum Mäklin, 482 [N]. Type Anomoearthrumdebile Mäklin, 1867, by subsequent designation (Gebien 1948: 540). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gemminger in Gemminger and Harold (1870: 2028).

Anophthalmolamus Ferrer, 122 [M]. Type Anophthalmolamusfuerteventurae Ferrer, 1993, by original designation. Status: valid genus in Tenebrioninae: incertae sedis.

Anopidium Jeannel & Paulian, 63 [N]. Type Anopidiumelgonicum Jeannel & Paulian, 1945, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.Note: originally described in Tenebrionoidea: Zopheridae: Colydiinae.

Anotoma Borchmann, 18, 81 [F]. Type Chrysolagriaheynei Borchmann, 1915, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Antennoluprops Schawaller, 30 [M]. Type Antennolupropsbremeri Schawaller, 2007, by original designation. Status: valid genus in Lagriinae: Lupropini.

Anteros Laporte, 235 [M]. Type Helopschalibeus Rossi, 1790 (= Tenebriocaeruleus Linnaeus, 1758), by subsequent designation (Iablokoff-Khnzorian 1964: 303). Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Erichson in Agassiz (1846a: 12). Note: junior homonym of Anteros Hübner, 1819 [Lepidoptera].

Anthracias Dejean, 205 [M]. Type Ulomacornutum Fischer, 1823, by monotypy. Status: junior synonym of Cryphaeus Klug, 1833 in Tenebrioninae: Toxicini: Toxicina. Synonymy: Gebien (1921b: 238).

†Anthracohelops Haupt, 114, 128 [M]. Type Anthracohelopsgigas Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Anthracosomus Agassiz, 26 [M]. Type species [automatic]: Anthrasomuschevrolatii Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Anthrasomus Guérin-Méneville, 1834 in Pimeliinae: Praociini. Note: unjustified emendation of Anthrasomus Guérin-Méneville, 1834, not in prevailing usage.

Anthracula Fairmaire, 236 [F]. Type Anthraculalatifrons Fairmaire, 1897, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Anthrasomus Guérin-Méneville, 32 [M]. Type Anthrasomuschevrolatii Guérin-Méneville, 1834, by monotypy. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini.

Anthrenopsis Koch, 74 [F]. Type Platydemascriptipennis Fairmaire, 1875 (= Basidesziczac Motschulsky, 1873), by original designation. Status: valid subgenus of Ellipsodes Wollaston, 1854 in Diaperinae: Crypticini.

Anticlia Gistel, x, 125 [F]. Type species [automatic]: Opatrumorientale Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: junior synonym of Sclerum Dejean, 1834 in Blaptinae: Opatrini: Sclerina. Note: unnecessary replacement name for Sclerum Dejean, 1834.

Antimachus Gistel, 1055 [M]. Type Phalariafurcifera Dalman, 1821, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Antofagapraocis Flores, 62, 68 [M]. Type Falsopraocissubnudus Kulzer, 1959, by original designation. Status: valid genus in Pimeliinae: Praociini.

Antoineius Koch, 418 [M]. Type Micrositusjeanneli Koch, 1945 (= Melambiusinermis Antoine, 1942), by original designation. Status: valid subgenus of Otinia Antoine, 1942 in Blaptinae: Dendarini: Melambiina.

Aoupinia Matthews, 441 [F]. Type Aoupiniapseudohelea Matthews, 2003, by original designation. Status: valid genus in Lagriinae: Adeliini.

Apalmia Fairmaire, 60 [F]. Type Apalmiacerambycina Fairmaire, 1896, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Apasis Pascoe, 139 [F]. Type Apasishowittii Pascoe, 1869, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Apatelus Mulsant & Rey, 88, 91 [M]. Type Apatelushopei Mulsant & Rey, 1859, by monotypy. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Carter (1926: 123).

Apatopsis Semenov, 368 [F]. Type Apatopsisgrombczewskii Semenov, 1891, by subsequent designation (R. Lucas 1920: 107). Status: valid genus in Pimeliinae: Pimeliini.

Apelina Saha, 429 [F]. Type Apelinakeralaensis Saha, 1988, by original designation. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer and Lillig (2014: 12).

Apellatus Pascoe, 45 [M]. Type Apellatuslateralis Pascoe, 1863 (= Apellatusamoenus Pascoe, 1866), by monotypy. Status: junior synonym of Euomma Boheman, 1858 in Alleculinae: Alleculini: Alleculina. Synonymy: Pascoe (1866a: 491).

Apentanes Reitter, 45, 51 [M]. Type Arthrodeisoccidentalis Fairmaire, 1868, by subsequent designation (Gebien 1937a: 536). Status: valid subgenus of Arthrodeis Solier, 1834 in Pimeliinae: Erodiini.

Apentanodes Reitter, 46, 53 [M]. Type Arthrodeisglobosus Reiche & Saulcy, 1857, by subsequent designation (Gebien 1937a: 539). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Aphaleria Reitter, 235 [F]. Type Aphaleriacapnisoides Reitter, 1896 (= Erodiuspygmaeus Fischer, 1821), by monotypy. Status: junior synonym of Bradyus Dejean, 1834 in Tenebrioninae: Dissonomini. Synonymy: Löbl et al. (2008b: 240).

Aphanaspis Wollaston, 478 [F]. Type Pimeliagranulicollis Wollaston, 1864, by subsequent designation (Löbl et al. 2008a: 40). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Aphanotus J.L. LeConte, 233 [M]. Type Eulabisbrevicornis J.L. LeConte, 1859, by original designation. Status: valid subgenus of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini.

Aphectus Carter, 127, 128 [M]. Type species [automatic]: Hectusanthracinus Pascoe, 1869, by monotypy. Status: junior synonym of Olisthaena Erichson, 1842 in Tenebrioninae: Heleini: Cyphaleina. Note: unnecessary replacement name for Hectus Pascoe, 1869.

Aphelus Gebien, 62, 78 [M]. Type Aphelussimplicicollis Gebien, 1921, by subsequent designation (Gebien 1941: 341). Status: junior synonym of Argobrachium Fairmaire, 1899 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1961b: 31).

Aphrotus Péringuey, 252 [M]. Type species [automatic]: Xenustricorniger Péringuey, 1899, by monotypy. Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Xenus Péringuey, 1899.

Aphtora Bates, 265 [F]. Type Aphtorarufipes Bates, 1872, by monotypy. Status: valid genus in Phrenapatinae: incertae sedis. Note: Matthews and Lawrence (2019: 623) placed this genus in the subfamily Phrenapatinae (tribal position uncertain).

Aphyllocerus Fairmaire, 348 [M]. Type Aphyllocerusdecipiens Fairmaire, 1881, by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1921a: 409).

Apistocerus Fairmaire, 78 [M]. Type Apistoceruswasmanni Fairmaire, 1899, by monotypy. Status: valid subgenus of Rhyzodina Chevrolat, 1873 in Tenebrioninae: Rhysopaussini.

Apithesis C.O. Waterhouse, 476 [F]. Type Apithesisobesa C.O. Waterhouse, 1881, by monotypy. Status: junior synonym of Clitobius Mulsant & Rey, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Purchart and Kamiński (2017: 143).

Aplanasida Reitter, 11, 30 [F]. Type Asidabrevicosta Solier, 1836, by monotypy. Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Apocrypha Eschscholtz, 13 [F]. Type Apocryphaanthicoides Eschscholtz, 1831, by monotypy. Status: valid genus in Tenebrioninae: Apocryphini.

Apocryphodes Matthews, 704, 765 [M]. Type Apocryphodesthompsoni Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Apodemus Fåhraeus, 293 [M]. Type Anomalipusplanus Fåhraeus, 1870, by subsequent designation (Iwan 2002c: 234). Status: junior synonym of Anomalipus Guérin-Méneville, 1831 in Blaptinae: Platynotini: Platynotina. Synonymy: Gebien (1938a: 409). Note: junior homonym of Apodemus Kaup, 1829 [Mammalia].

Apolites Jacquelin du Val, 324 [M]. Type Helopsmucoreus Waltl, 1838, by monotypy. Status: senior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Synonymy: Allard (1876b: ciii). Note: junior homonym of Apolites Sundevall, 1835 [Aves].

Apomestris Bates, 357 [M]. Type Apomestriswestwoodi Bates, 1873, by monotypy. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1926: 119, with Altes Pascoe, 1869, a junior synonym of Cyphaleus Westwood, 1841).

Apostethus Pascoe, 27 [M]. Type Apostethusterrenus Pascoe, 1882, by monotypy. Status: junior synonym of Adelodemus Haag-Rutenberg, 1878 in Lagriinae: Adeliini. Synonymy: Carter (1926: 131).

Apristopus Kolbe, 167 [M]. Type Priosceliscrassicornis Westwood, 1842, by monotypy. Status: junior synonym of Calostegia Westwood, 1843 in Lagriinae: Pycnocerini. Synonymy: Gebien (1904a: 169).

Aprosphaena Reitter, 140, 142 [F]. Type Aprosphaenaadriani Reitter, 1916 (= Tageniastriatopunctata Wiedemann, 1821), by subsequent designation (Löbl et al. 2008a: 40). Status: junior synonym of Stenosida Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Blair (1935a: 103).

Apsena J.L. LeConte, 228 [F]. Type Eulabispubescens J.L. LeConte, 1851, by original designation. Status: valid genus in Tenebrioninae: Eulabini.

Apsheronellus Bogatchev, 157 [M]. Type Apsheronellusarenarius Bogatchev, 1967, by original designation. Status: valid genus in Blaptinae: Pedinini: Leichenina.

Apsida Lacordaire, 309 [F]. Type Apsidachrysomelina Lacordaire, 1859, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Aptereucyrtus Gebien, 477 [M]. Type Aptereucyrtushemichalceus Gebien, 1922, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Aptereutochia Kaszab, 190 [F]. Type Eutochiaaptera Kaszab, 1980, by original designation. Status: valid subgenus of Cenoscelis Wollaston, 1868 in Tenebrioninae: Ulomini. Note: this name was first published by Kaszab (1979a: 92) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article (13.1) and is therefore unavailable from that date.

Aptericula Borchmann, 219 [F]. Type Aptericulanyassensis Borchmann, 1937, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Apterobrachys Kaszab, 295 [M]. Type Apterobrachyswilhelminae Kaszab, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Apteroclitobius Koch, 391, 404 [M]. Type Clitobiusgrandis Fairmaire, 1896, by original designation. Status: junior synonym of Clitobius Mulsant & Rey, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Purchart and Kamiński (2017: 143).

Apterocyphostethe Kaszab, 302 [F]. Type Himatismuskoltzei Reitter, 1895, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini.

Apterogena Ardoin, 67 [F]. Type Apterogenacanonnei Ardoin, 1962, by original designation. Status: senior synonym of Genateropa Bouchard & Bousquet, nom. nov. in Stenochiinae: Stenochiini. Note: junior homonym of Apterogena Amyot, 1847 [Hemiptera].

Apteroleprocaulus Kaszab, 182 [M]. Type Leprocaulusmontanus Kaszab, 1982, by original designation. Status: junior synonym of Hexarhopalus Fairmaire, 1891 in Stenochiinae: Cnodalonini. Synonymy: Bečvář and Purchart (2008: 39).

Apteromaia Kulzer, 719 [F]. Type Eucyrtusovipennis Gebien, 1913, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Apteromerus Blair, 72 [M]. Type Opatrinusconvexus Fairmaire, 1849, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Apteromira Weise, 123 [F]. Type Cistelaovulum Kiesenwetter, 1863, by original designation. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Apteronympha Seidlitz, 336, 338 [F]. Type Lagriarubida Graells, 1858, by subsequent designation (Merkl 2004: 284). Status: valid subgenus of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina.

Apterophenus Gebien, 325, 342 [M]. Type Apterophenusevanescens Gebien, 1921, by subsequent designation (Gebien 1941: 1142). Status: valid genus in Stenochiinae: Cnodalonini.

Apteroseriscius Koch, 64 [M]. Type Pseudoserisciusespanoli Koch, 1950, by original designation. Status: valid genus in Diaperinae: Crypticini.

Apterosphaeria Koch, 298 [F]. Type Derosphaeriusrugifrons Fairmaire, 1888, by original designation. Status: valid subgenus of Derosphaerius Westwood, 1881 in Pimeliinae: Tentyriini.

Apterotarpela Kaszab, 262 [F]. Type Apterotarpelaklapperichi Kaszab, 1954, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Apterotheca Gebien, 348 [F]. Type Chariothecabesti Blackburn, 1894, by subsequent designation (Gebien 1942a: 338). Status: valid genus in Stenochiinae: Cnodalonini.

Apterozidalus Ardoin, 1315 [M]. Type Apterozidalusroyi Ardoin, 1965, by original designation. Status: junior synonym of Zidalus Mulsant & Rey, 1853 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (1995b: 362).

Apteruleda Gebien, 134 [F]. Type Apteruledauncipes Gebien, 1928, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Apteruloma Gebien, 149 [N]. Type Apterulomamagnum Gebien, 1928, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Apterulomoides Kaszab, 279 [M]. Type Ulomarotundipenne Kaszab, 1982, by monotypy. Status: valid subgenus of Uloma Dejean, 1821 in Tenebrioninae: Ulomini.

Aptila Fåhraeus, 258 [F]. Type Aptilacostata Fåhraeus, 1870, by subsequent designation (R. Lucas 1920: 115). Status: valid genus in Blaptinae: Pedinini: Helopinina.

Arabammobius Grimm & Lillig, 307 [M]. Type Ammobiustarsalis Grimm, 2012, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Arabcynaeus Schawaller, 164 [M]. Type Arabcynaeusbremeri Schawaller, 2009, by original designation. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Araeopselaphus Gebien, 10 [M]. Type Araeopselaphusmyrmecophilus Gebien, 1921, by monotypy. Status: valid genus in Diaperinae: Crypticini.

Araeoschizus J.L. LeConte, 138 [M]. Type Araeoschizuscostipennis J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Araeoschizina.

Araucaricola Lea, 218 [F]. Type Araucaricolaebenina Lea, 1929, by monotypy. Status: junior synonym of Iscanus Fauvel, 1904 in Lagriinae: Lupropini. Synonymy: Kaszab (1982a: 59).

Archaeoglenes Broun, 188 [M]. Type Archaeoglenescostipennis Broun, 1893, by monotypy. Status: valid genus in Phrenapatinae: Archaeoglenini.

Archasida Wilke, 261 [F]. Type Afrasidainnotata Wilke, 1922, by original designation. Status: valid subgenus of Afrasida Wilke, 1922 in Pimeliinae: Asidini.

Archeophthora Kaszab, 166 [F]. Type Brachycilibetasmanicum Carter, 1919, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Archinamaqua Schawaller, 80 [F]. Type Archinamaqualyleae Schawaller, 2012 (= Menederopsisconstricta Koch, 1954), by original designation. Status: junior synonym of Menederopsis Koch, 1954 in Blaptinae: Platynotini: Eurynotina. Synonymy: Kamiński and Schawaller (2019: 293).

Archinamibia Koch, 157 [F]. Type Archinamibiapeezi Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Arcothymus Pascoe, 476 [M]. Type Arcothymuscoenosus Pascoe, 1866 (= Adeliumtriste Montrouzier, 1860), by monotypy. Status: valid genus in Lagriinae: Adeliini. Note: as mentioned by Matthews and Bouchard (2008: 350) the type species was originally described from Australia in error; the genus Arcothymus Pascoe, 1866 is endemic to New Caledonia.

Arctylus Dejean, 180 [M]. Type Praocispentagonus Lacordaire, 1830, by monotypy. Status: junior synonym of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Synonymy: Bousquet and Bouchard (2013a: 43).

Ardamimicus Smith, 601 [M]. Type Ardamimicuscognatoi Smith, 2013, by original designation. Status: valid genus in Pimeliinae: Asidini.

Ardeleodes Blaisdell, 128 [M]. Type Eleodestibialis Blaisdell, 1909, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini. Note: the alternative original spelling Arpeleodes, used by Blaisdell (1937: 128), was rejected by Gebien (1938a: 63) who acted as First Reviser.

Ardelio Gistel, xi [M]. Type species [automatic]: Ulomacornutum Fischer, 1823, by monotypy. Status: junior synonym of Cryphaeus Klug, 1833 in Tenebrioninae: Toxicini: Toxicina. Note: unnecessary replacement name for Anthracias Dejean, 1834.

Ardoinia Kaszab, 249 [F]. Type Ardoiniadiaclinoides Kaszab, 1969, by original designation. Status: valid genus in Tenebrioninae: Alphitobiini.

Ardoinia Özdikmen, 202 [F]. Type species [automatic]: Orghidaniatorrei Ardoin, 1977, by monotypy. Status: senior synonym of Spelaebiosis Bousquet & Bouchard, 2018 in Tenebrioninae: Triboliini. Note: replacement name for Orghidania Ardoin, 1977; junior homonym of Ardoinia Kaszab, 1969 [Coleoptera: Tenebrionidae: Tenebrioninae: Alphitobiini].

Ardoiniellus Schawaller, 138 [M]. Type Ardoiniellusmontanus Schawaller, 2013, by original designation. Status: valid genus in Lagriinae: Lupropini.

Arenacara Penrith, 43, 46 [N]. Type Stenocarabrunnipes Haag-Rutenberg, 1877, by original designation. Status: valid subgenus of Stenocara Solier, 1835 in Pimeliinae: Adesmiini.

Arenoblaps G.S. Medvedev, 400 [F]. Type Blaps hiemalis Semenov-Tjan-Shansky & Bogatchev, 1940, by original designation. Status: valid subgenus of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina.

Argasidus Péringuey, 251 [M]. Type Argasidussquamosus Péringuey, 1899, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Argenticrinis Louw, 99, 100 [M]. Type Argenticrinishaackei Louw, 1979 (= Psammodeslossowi Koch, 1952), by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Argobrachium Fairmaire, 216 [N]. Type Argobrachiumimpressifrons Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Argoporis Horn, 325 [F]. Type Cerenopuscostipennis J.L. LeConte, 1851, by subsequent designation (Gebien 1937a: 797). Status: valid genus in Tenebrioninae: Cerenopini. Note: as pointed out by Bousquet et al. (2018: 97, 179), evidence shows that Threnus Motschulsky, 1870 was published before the currently accepted valid name Argoporis Horn, 1870; as Threnus Motschulsky, 1870 was used as valid after 1899 (e.g., Leng 224), reversal of precedence cannot be used to conserve usage of Argoporis Horn, 1870; an application to the ICZN is necessary to conserve usage of Argoporis Horn, 1870.

Argutiolana Robiche, 191 [F]. Type Argutiolanamaguini Robiche, 2001, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: transferred from Tenebrioninae: Tenebrionini by Robiche (2019a: 97).

Argyradelpha G.S. Medvedev, 306 [F]. Type Argyradelphalopatini G.S. Medvedev, 2005, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Argyrophana Semenov, 222, 224 [F]. Type Argyrophanadeserti Semenov, 1889, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Ariarathus Fairmaire, ccxi [M]. Type Ariarathusulomoides Fairmaire, 1891 (= Tenebrioatronitens Fairmaire, 1891), by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Armalia Casey, 289, 330 [F]. Type Emmenastustexanus J.L. LeConte, 1866, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Armenohelops Nabozhenko, 42 [M]. Type Armenohelopsarmeniacus Nabozhenko, 2002, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Armigena Bouchard & Bousquet, new subgenus [F]. Type Nesogenatestaceipes Fairmaire, 1868, by present designation. Status: valid subgenus of Nesogena Mäklin, 1863 in Tenebrioninae: Praeugenini. Note: first proposed by Froussart (1961: 60) without type species designation; the subgenusArmigena, which is currently used as valid, is therefore unavailable (ICZN 1999, Articles 13.3, 16.1); we hereby make the name available by selecting Nesogenatestaceipes Fairmaire, 1868 as type species and referring to Froussart (1961: 60) for the character states that characterise and differentiate Armigena.

Arnoldiola Semenov-Tjan-Shansky & Bogatchev, 201 [F]. Type Arnoldiolapeculiaris Semenov-Tjan-Shansky & Bogatchev, 1940 (= Stalagmopteraruginota Reitter, 1896), by original designation. Status: junior synonym of Stalagmoptera Solsky, 1876 in Pimeliinae: Pimeliini. Synonymy: Löbl et al. (2008b: 168). Note: junior homonym of Arnoldiola Strand, 1928 [Diptera].

Arrhabaeus Champion, 144 [M]. Type Arrhabaeusconvexus Champion, 1886, by monotypy. Status: junior synonym of Dioedus J.L. LeConte, 1862 in Phrenapatinae: Penetini. Synonymy: Kaszab (1977a: 314).

Arrhenoplita W. Kirby, 235 [F]. Type Ips haemorrhoidalis Fabricius, 1787, by original designation. Status: junior synonym of Neomida Latreille, 1829 in Diaperinae: Diaperini: Diaperina. Synonymy: Duponchel and Chevrolat (1841: 157).

Artactes Pascoe, xii [M]. Type Artactesnigritarsis Pascoe, 1868, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Arthrochora Gebien, 75 [F]. Type Arthrochoraarenicola Gebien, 1938, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Arthroconus Solier, 238 [M]. Type Arthroconuspiceus Solier, 1851 (= Gymnognathusfuscus Solier, 1851), by subsequent designation (Lacordaire 1859a: 67). Status: valid genus in Pimeliinae: Edrotini. Note: the First Reviser (Arthroconus Solier, 1851 versus Gymnognathus Solier, 1851) is Lacordaire (1859a: 67).

Arthrodeis Solier, 508, 513 [M]. Type Arthrodeisrotundatus Solier, 1834, by subsequent designation (Hope 1841: 114). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Arthrodes Agassiz, 35 [M]. Type species [automatic]: Arthrodeisrotundatus Solier, 1834, by subsequent designation (Hope 1841: 114). Status: junior synonym of Arthrodeis Solier, 1834 in Pimeliinae: Erodiini. Note: unjustified emendation of Arthrodeis Solier, 1834, not in prevailing usage.

Arthrodibius Lesne, 227, 235 [M]. Type Arthrodeislaxepunctatus Fairmaire, 1884, by subsequent designation (Sharp 1919: 119). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Arthrodinus Reitter, 299 [M]. Type Erodiusobesus Brullé, 1839, by subsequent designation (Reitter 1914a: 46). Status: valid subgenus of Arthrodeis Solier, 1834 in Pimeliinae: Erodiini.

Arthrodion Lesne, 227, 234 [N]. Type Spyrathusafricanus Fairmaire, 1882, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Arthrodosis Reitter, 299 [F]. Type Erodiusglobosus Faldermann, 1837, by subsequent designation (R. Lucas 1920: 119). Status: valid genus in Pimeliinae: Erodiini. Note: the original combination of the accepted name of the type species, Erodiusglobosus Faldermann, 1837, is a junior primary homonym of Erodiusglobosus Thunberg, 1787.

Arthrodygmus Reitter, 46 [M]. Type Arthrodygmusfieberi Reitter, 1914, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Arthrohyalosis Kaszab, 75 [F]. Type Arthrodosismostofii Pierre, 1975, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Arthrohyalus Koch, 490, 503 [M]. Type Arthrohyalussarcinipennis Koch, 1943, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Arthromacra W. Kirby, 238 [F]. Type Arthromacradonacioides W. Kirby, 1837 (= Lagriaaenea Say, 1824), by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Arthroplatus Solier, 246 [M]. Type Arthroplatuspallipes Solier, 1851, by monotypy. Status: junior synonym of Acropteryx Gistel, 1831 in Tenebrioninae: Acropteronini. Synonymy: Schaum (1852: 183, with Acropteron Perty, 1832, a junior synonym of Acropteryx Gistel, 1831).

Arturium Koch, 83 [N]. Type Melanolophusater C.O. Waterhouse, 1885, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Artystona Bates, 104 [F]. Type Titaenaerichsonii White, 1846, by subsequent designation (R. Lucas 1920: 120). Status: valid genus in Tenebrioninae: Titaenini. Note: this name was introduced one year earlier by Bates (1873f: 473) but it is unavailable from that date since there is no description nor originally included species (ICZN 1999, Article 12.1).

Arunogria Merkl, 14 [F]. Type Arunogriapubescens Merkl, 1991, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Aryenis Bates, 309 [F]. Type Aryenisrufescens Bates, 1868 (= Statiraunicolor Blanchard, 1843), by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Asbolius Fairmaire, 134 [M]. Type Asboliusquadricollis Fairmaire, 1902, by monotypy. Status: junior synonym of Rhammatodes Haag-Rutenberg, 1876 in Pimeliinae: Tentyriini. Synonymy: Blair (1935a: 103).

Asbolodes Fairmaire, 52 [M]. Type Asbolodeshumerosus Fairmaire, 1892, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asbolodomimus Pic, 20 [M]. Type Asbolodomimussubcarinatus Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asbolus J.L. LeConte, 129 [M]. Type Asbolusverrucosus J.L. LeConte, 1851, by subsequent designation (Aalbu 2005: 721). Status: valid genus in Pimeliinae: Cryptoglossini. Note: the older name Asbolus Voet, 1793 [Coleoptera: Silphidae] was published in a work that did not include consistent application of binominal nomenclature and is therefore unavailable (ICZN 1999, Article 11.4).

Ascalabus Fairmaire, 30 [M]. Type Ascalabuspedinoides Fairmaire, 1893, by monotypy. Status: senior synonym of Calabosca Fairmaire, 1894 in Stenochiinae: Cnodalonini. Note: junior homonym of Ascalabus Agassiz, 1846 [Pisces].

Ascelosodis Redtenbacher, 117 [F]. Type Ascelosodisserripes Redtenbacher, 1868, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Asemogena Péringuey, 281 [F]. Type Asemogenasimplex Péringuey, 1904, by subsequent designation (R. Lucas 1920: 121). Status: valid genus in Stenochiinae: Stenochiini.

Asialassus Nabozhenko & Ando, 311 [M]. Type Helopscordicollis Marseul, 1876, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Asida Latreille, 167 [F]. Type Opatrumgriseum Fabricius, 1781, by monotypy. Status: valid genus and subgenus in Pimeliinae: Asidini.

Asidelia Fairmaire, 296 [F]. Type Asideliacontracta Fairmaire, 1905, by monotypy. Status: valid genus in Pimeliinae: Praociini.

Asidesthes Fairmaire, 246 [F]. Type Asidesthesperrieri Fairmaire, 1900, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Asididius Fairmaire, 236 [M]. Type Asididiuscoquerelii Fairmaire, 1869, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asidina Casey, 76, 169 [F]. Type Pelecyphorusparallelus J.L. LeConte, 1851, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Asidoblaps Fairmaire, 342 [F]. Type Asidoblapsdavidis Fairmaire, 1886, by subsequent designation (R. Lucas 1920: 122). Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Asidobothris Fairmaire, 72 [F]. Type Asidobothrisclathrata Fairmaire, 1886, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asidodema Koch, 139 [F]. Type Oncosomaalternicoste Gebien, 1910, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Asidomachla Wilke, 262 [F]. Type Asidabicostata Fåhraeus, 1870, by original designation. Status: junior synonym of Machlomorpha Péringuey, 1899 in Pimeliinae: Asidini. Synonymy: Koch (1962a: 125).

Asidomorpha Koch, 121 [F]. Type Afrasidaprona Wilke, 1922, by original designation. Status: valid genus in Pimeliinae: Asidini.

Asidopsis Casey, 77, 185 [F]. Type Asidaopaca Say, 1824, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Asiobirus G.S. Medvedev, 170, 179 [M]. Type Cabirusvalidipes Reitter, 1891, by original designation. Status: junior synonym of Cabirutus Strand, 1929 in Blaptinae: Pedinini: Pedinina. Synonymy: Kamiński and Iwan (2017: 595).

Asiocaedius G.S. Medvedev & Nepesova, 138 [M]. Type species [automatic]: Pseudocaediuskiseritzkii G.S. Medvedev, 1966, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: replacement name for Pseudocaedius G.S. Medvedev, 1966.

Asiomira Dubrovina, 367 [F]. Type Isomiraophthalmica Seidlitz, 1896, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: changed from subgenus of Isomira Mulsant, 1856 to valid genus by Novák (2016a: 179).

Asiopus Sharp, 43 [M]. Type Asiopusopatroides Sharp, 1892, by monotypy. Status: valid genus in Lagiinae: incertae sedis. Note: this little-known genus has been treated as a member of the subfamily Lagriinae (e.g., Fery 2013: 66).

Asiris Motschulsky, 24 [F]. Type Asirisangulicollis Motschulsky, 1872 (= Tenebrioaustralis Boisduval, 1835), by original designation. Status: junior synonym of Meneristes Pascoe, 1869 in Tenebrioninae: Heleini: Asphalina. Synonymy: C.O. Waterhouse (1876: 288).

Asopidiopsis Kaszab, 511, 515 [F]. Type Asopidiopsisovalis Kaszab, 1955, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Asopis Haag-Rutenberg, 104 [F]. Type Asopissuavis Haag-Rutenberg, 1878, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asphaltesthes Kraatz, 80, 181 [F]. Type Mesostenacostata Erichson, 1843, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Asphalus Pascoe, xii [M]. Type Asphalusebeninus Pascoe, 1868, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Asphalina.

Asphena Semenov, 218 [F]. Type Asphenakomarowi Semenov, 1889, by monotypy. Status: junior synonym of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini. Synonymy: Reitter (1916c: 141).

Aspidius Mulsant & Rey, 123 [M]. Type Blapspunctata Fabricius, 1792, by subsequent designation (Bousquet et al. 2018: 195). Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Champion (1885: 124).

Aspidocephalus Motschulsky, 63 [M]. Type Aspidocephalusdesertus Motschulsky, 1839, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina. Note: unjustified emendation of the original spelling Aspicephalus, introduced by Lacordaire (1859a: 107), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Aspidolobus Redtenbacher, 118 [M]. Type Aspidolobuspiliger Redtenbacher, 1868, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Aspidosoma Agassiz, 36, 37 [N]. Type species [automatic]: Aspisomafulvipenne Duponchel & Chevrolat, 1841, by original designation. Status: junior synonym of Anaedus Blanchard, 1842 in Lagriinae: Goniaderini. Note: unjustified emendation of Aspisoma Duponchel & Chevrolat, 1841, not in prevailing usage.

Aspidosternum Mäklin, 500 [N]. Type Tenebriocyaneus Fabricius, 1794, by monotypy. Status: junior synonym of Metallonotus Gray, 1832 in Lagriinae: Pycnocerini. Synonymy: Alluaud (1889: xlvi).

Aspila Fåhraeus, 251 [F]. Type Aspilabicostata Fåhraeus, 1870, by monotypy. Status: senior synonym of Prunaspila Koch, 1950 in Pimeliinae: Adelostomini. Note: junior homonym of Aspila Stephens, 1834 [Lepidoptera].

Aspilomorpha Koch, 33, 101 [F]. Type Aspilomorphamediolobata Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Aspisoma Duponchel & Chevrolat, 240 [N]. Type Aspisomafulvipenne Duponchel & Chevrolat, 1841, by original designation. Status: senior synonym of Anaedus Blanchard, 1842 in Lagriinae: Goniaderini. Synonymy: Lacordaire (1859a: 396). Note: junior homonym of Aspisoma Laporte, 1833 [Coleoptera: Lampyridae].

Astalbus Fairmaire, 484 [M]. Type Astalbusscrobicollis Fairmaire, 1900, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Astathmetus Bates, 23 [M]. Type Astathmetusalienus Bates, 1874, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: this name was introduced one year earlier by Bates (1873f: 472) but it is unavailable from that date since there is no description nor originally included species (ICZN 1999, Article 12.1).

Astatira Borchmann, 218, 322 [F]. Type Astatirahumeralis Borchmann, 1921, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Asthenochirus Fairmaire, viii [M]. Type Asthenochirusnigropunctatus Fairmaire, 1885, by subsequent designation (R. Lucas 1920: 123). Status: valid genus in Tenebrioninae: Amarygmini. Note: redescribed as new by Fairmaire (1887b: 298).

Asthenopoda Chatanay, 527, 544 [F]. Type Asthenopodafragilis Chatanay, 1915, by original designation. Status: junior synonym of Psilonesogena Bates, 1879 in Stenochiinae: Stenochiini. Synonymy: De Moor (1970: 9).

Asticostena Fairmaire, 228 [F]. Type Asticostenaalternata Fairmaire, 1897, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Astorthocnemis Lillig & Pavlíček, 98 [F]. Type Storthocnemissaudita Koch, 1965, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Astrotus J.L. LeConte, 19 [M]. Type Microschatiacontorta J.L. LeConte, 1853, by original designation. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Asyleptus Péringuey, 179 [M]. Type Asyleptusfumosus Péringuey, 1896, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Asyrmatus Canzoneri, 149 [M]. Type Helopspiceus Sturm, 1826, by original designation. Status: junior synonym of Pystelops Gozis, 1910 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: new synonym [YB]. Note: Pystelops Gozis, 1910 has been forgotten in the literature; its type species is currently included in the subgenusAsyrmatus Canzoneri, 1959 and for that reason Canzoneri’s name is considered a junior synonym of Pystelops; the original combination of the accepted name of the type species, Helopspiceus Sturm, 1826, is a junior primary homonym of Helopspiceus G.-A. Olivier, 1792.

Atahualpina Español, 113 [F]. Type Atahualpinaperuviana Español, 1960 (= Phaleriasubparalella Chevrolat, 1878), by original designation. Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Synonymy: Triplehorn (1991: 258).

Atasthalomorpha Miyatake, 68, 77 [F]. Type Atasthalusdentifrons Lewis, 1894, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Atasthalus Pascoe, 348 [M]. Type Atasthalusspectrum Pascoe, 1871, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Athrodactyla Klug, 90 [F]. Type Athrodactylaelongata Klug, 1833, by subsequent designation (Hope 1841: 126). Status: valid genus in Tenebrioninae: Tenebrionini.

Atlasion Koch, 427 [N]. Type Micrositusbedeli Escalera, 1914, by original designation. Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina. Note: the First Reviser (Atlasion Koch, 1948 versus Megatlasion Koch, 1948) is Antoine (1957: 351).

Atlasotaurus Bouyon, 464 [M]. Type Omophlusmaroccanus P.H. Lucas, 1846, by original designation. Status: valid subgenus of Heliotaurus Mulsant, 1856 in Alleculinae: Cteniopodini.

Atoichus Carter, 72 [M]. Type Licymniusbicolor Blackburn, 1893, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Atoreuma Gebien, 1132 [N]. Type species [automatic]: Toreumacupreum Carter, 1913, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: replacement name for Eutoreuma Carter, 1914.

Atrachyderma Skopin, 228 [N]. Type Pimeliasetosa Faldermann, 1832, by original designation. Status: valid subgenus of Trachyderma Latreille, 1828 in Pimeliinae: Pimeliini. Note: Pimeliasetosa, the type species, has been attributed to Fischer in the literature; the species was described as “Pimeliasetosa Fald.” which, from the context, indicates that Faldermann was responsible for the description of the species.

Atractus Boisduval, 283 [M]. Type Atractusviridis Boisduval, 1835, by subsequent designation (Duponchel and Chevrolat 1841: 312). Status: senior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324). Note: junior homonym of Atractus Wagler, 1828 [Reptilia].

Atrocrates Koch, 82 [M]. Type Trigonopusplatyderus Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Atrocrypticanus Iwan, 73 [M]. Type Atrocrypticanusfraternus Iwan, 1999, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Atropsorodes Ardoin, 88 [M]. Type Atropsorodesbarnardi Ardoin, 1963, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: name published earlier by Ardoin (1962b: 969) without fixation of a type species in the original publication (ICZN 1999, Article 13.3) and is therefore unavailable from that date.

Atryphodes Pascoe, 478 [M]. Type species [automatic]: Thoracophoruswalckenaerii Hope, 1841, by original designation. Status: junior synonym of Cardiothorax Motschulsky, 1860 in Lagriinae: Adeliini. Note: unnecessary replacement name for Thoracophorus Hope, 1841.

Auchmobius J.L. LeConte, 139 [M]. Type Auchmobiussublaevis J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Augolesthus Motschulsky, 34 [M]. Type Augolesthuspurpureofasciatus Motschulsky, 1872, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Aulacodera Agassiz, 40, 41 [F]. Type species [automatic]: Nycteliacrenicosta Guérin-Méneville, 1834, by subsequent designation (Gebien 1937a: 752). Status: junior synonym of Auladera Solier, 1836 in Pimeliinae: Nycteliini. Note: unjustified emendation of Auladera Solier, 1836, not in prevailing usage.

Aulacus Gray in Griffith and Pidgeon, 783 [M]. Type Aulacuschilensis Gray, 1832, by monotypy. Status: senior synonym of Orthogonoderes Solier, 1841 in Pimeliinae: Praociini. Synonymy: Kulzer (1958a: 81). Note: junior homonym of Aulacus Jurine, 1807 [Hymenoptera].

Auladera Solier, 307, 331 [F]. Type Nycteliacrenicosta Guérin-Méneville, 1834, by subsequent designation (Gebien 1937a: 752). Status: valid genus in Pimeliinae: Nycteliini.

Aulonasida Reitter, 9, 19 [F]. Type Asidachauveneti Solier, 1836, by subsequent designation (F. Soldati 2008: 32). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Aulonogria Borchmann, 9 [F]. Type Lagriarugosa Fabricius, 1801, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina. Note: also published as new in Borchmann (1930a: 405, 424).

Aulonolcus Reitter, 168 [M]. Type Pedinusaltaicus Gebler, 1830, by monotypy. Status: valid subgenus of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina.

Aulonoscelis Reitter, 173 [F]. Type Platyscelishauseri Reitter, 1895 (= Dilaphilacoides Fischer von Waldheim, 1844), by original designation. Status: junior synonym of Gebleria Motschulsky, 1846 in Blaptinae: Blaptini: Prosodina. Synonymy: G.S. Medvedev (2005b: 99).

Auristira Borchmann, 47, 129 [F]. Type Auristiraoctocostata Borchmann, 1916, by present designation. Status: junior synonym of Costiferolagria Pic, 1915 in Lagriinae: Lagriini: Lagriina. Synonymy: Borchmann (1936: 176).

Australoseriscius Koch, 65 [M]. Type Crypticusexplorator Gebien, 1920, by original designation. Status: valid subgenus of Pseudoseriscius Español, 1950 in Diaperinae: Crypticini.

Austrocaribius Marcuzzi, 18 [M]. Type Austrocaribiusvenezuelensis Marcuzzi, 1954, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Austropalorus Halstead, 72, 129 [M]. Type Austropalorusplanatus Halstead, 1967, by original designation. Status: valid genus in Tenebrioninae: Palorini.

Austropeus Carter, 543 [M]. Type Austropeuspustulosus Carter, 1924, by monotypy. Status: junior synonym of Apterotheca Gebien, 1921 in Stenochiinae: Cnodalonini. Synonymy: Bouchard (2002: 452).

Austroptorina Bai, Li & Ren, 166 [F]. Type Gnaptorinalongicornis Li & Ren, 2004, by original designation. Status: valid subgenus of Gnaptorina Reitter, 1887 in Blaptinae: Blaptini: Gnaptorinina.

Autocera Wollaston, 154 [F]. Type Autoceralaticeps Wollaston, 1857, by monotypy. Status: junior synonym of Cnemeplatia Costa, 1847 in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Synonymy: Kraatz (1859: 75).

Axumia Reiche, pl. 22 [F]. Type Axumiapraelonga Reiche, 1850, by monotypy. Status: junior synonym of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Kraatz (1865: 170).

Axynaon Blackburn, 34 [N]. Type Axynaonchampioni Blackburn, 1897, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Azarelius Fairmaire, vii [M]. Type Azareliussculpticollis Fairmaire, 1892, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: transferred from Rhysopaussini to Amarygmini by Löbl and Smetana (2010: 34).

Azonoderus Harold, 125 [M]. Type Azonoderustristis Harold, 1879, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Balachowskya Peyerimhoff, 61 [F]. Type Balachowskyaportentosa Peyerimhoff, 1928, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Balassogloa Semenov, 372 [F]. Type Balassogloasphenarioides Semenov, 1891, by subsequent designation (R. Lucas 1920: 131). Status: valid genus in Alleculinae: Cteniopodini.

Balius Gistel, xi, xiv [M]. Type species [automatic]: Pimeliasimplex Solier, 1836, by monotypy. Status: junior synonym of Melanostola Solier, 1836 in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Melanostola Solier, 1836.

Bancocistela Pic, 151 [F]. Type Bancocistelaivoirensis Pic, 1947, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Bantodemus Koch, 428 [M]. Type Trigonopuslethaeus Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Baratus Fairmaire, 233 [M]. Type Baratuscrenulatus Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Barbora Novák, 462 [F]. Type Barboracastanea Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Barlacus Fairmaire, 45 [M]. Type Barlacuscostulatus Fairmaire, 1900 (= Asyleptusfumosus Péringuey, 1896), by monotypy. Status: junior synonym of Asyleptus Péringuey, 1896 in Tenebrioninae: Amarygmini. Synonymy: Ardoin (1962b: 957); Bremer (2013a: 77).

Barsenis Pascoe, 17 [F]. Type Barsenisfulvipes Pascoe, 1887, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Bartolozzia Ferrer, 369 [F]. Type Bartolozziahispida Ferrer, 1998, by original designation. Status: valid genus in Pimeliinae: Asidini.

Barycistela Blackburn, 327 [F]. Type Barycistelarobusta Blackburn, 1891, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Baryscelis Boisduval, 253 [F]. Type Baryscelislaticollis Boisduval, 1835, by subsequent designation (Hope 1841: 126). Status: valid genus in Tenebrionidae: incertae sedis. Note: the type material of the two species included in this genus has not been located (Matthews and Bouchard 2008: 350) and therefore the identity of the genus is uncertain.

Barytipha Pascoe, 288, 292 [F]. Type Barytiphasocialis Pascoe, 1869, by monotypy. Status: junior synonym of Pterohelaeus Brême, 1842 in Tenebrioninae: Heleini: Heleina. Synonymy: Matthews (1993: 1041).

Basanaedus Pic, 11 [M]. Type Basanaedusluteomaculatus Pic, 1917, by subsequent designation (Gebien 1941: 820). Status: junior synonym of Lyprochelyda Fairmaire, 1899 in Lagriinae: Goniaderini. Synonymy: Ardoin (1969a: 166).

Basanopsis Gebien, 21 [F]. Type Basanopsiscurvipes Gebien, 1914, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Basanus Lacordaire, 306 [M]. Type Basanusforticornis Lacordaire, 1859, by monotypy. Status: valid genus in Diaperinae: Scaphidemini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Basides Motschulsky, 471 [M]. Type Basidesbifasciatus Motschulsky, 1873, by subsequent designation (Gebien 1925d: 560). Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: Basides Motschulsky, 1873 was used as valid after 1899 (e.g., Gebien, 381) and therefore reversal of precedence cannot be used to conserve usage of Ischnodactylus Chevrolat, 1877.

Basilewskyum Koch, 30, 94 [N]. Type Basilewskyumstenosinoide Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Bassianus Matthews & Doyen, 44 [M]. Type Tenebriocolydioides Erichson, 1842, by original designation. Status: valid genus in Tenebrioninae: Heleini: Asphalina.

Batessia Ponting, 131 [F]. Type species [automatic]: Agastheneswestwoodi Bates, 1873, by monotypy. Status: junior synonym of Agastenes R. Lucas, 1920 in Tenebrioninae: Heleini: Cyphaleina. Note: unnecessary replacement name for Agasthenes Bates, 1873.

Batuliodes Casey, 499 [M]. Type Batuliusrotundicollis J.L. LeConte, 1851, by original designation. Status: valid genus in Pimeliinae: Anepsiini.

Batuliomorpha Doyen, 359 [F]. Type Batuliomorphacomata Doyen, 1987, by original designation. Status: valid genus in Pimeliinae: Anepsiini.

Batulius J.L. LeConte, 148 [M]. Type Batuliussetosus J.L. LeConte, 1851, by subsequent designation (Casey 1907: 497). Status: valid genus in Pimeliinae: Anepsiini.

Bearnicistela Pic in E. Olivier and Pic, 139 [F]. Type Bearnicistelaluteicolor Pic, 1909, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Bechuanitis Koch, 93 [F]. Type Trachynotusbrucki Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Becvaramarygmus Masumoto, 369 [M]. Type fixed herein (ICZN 1999, Article 70.3) as Dietysusnodicornis Gravely, 1915, misidentified as Dietysusatricolor Pic, 1922 in the original designation in Masumoto (1999). Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Note: the type species “Dietysusatricolor Pic” was first established by original designation; Bremer and Lillig (2014: 12, 49) first noted that Dietysusatricolor Pic of Masumoto (1999) was identical to Dietysusnodicornis Gravely, 1915; we follow the concept of Bremer and Lillig (2014: 12, 49) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Dietysusatricolor Pic, 1922 is a junior synonym of Amarygmusfilicornis (Gravely, 1915); Iwan et al. (2020: 262) erroneously placed Becvaramarygmus Masumoto, 1999 as a junior synonym of the subgenusAmarygmus Dalman, 1823 (M. Lillig and H. Bremer, pers. comm. 2020).

Becvarius Masumoto, 207 [M]. Type Becvariusstanislavi Masumoto, 1998, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bellendenum Matthews, 704, 794 [N]. Type Bellendenumgonyxuthum Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Belopomerus Reitter, 3 [M]. Type Calcarzoufali Reitter, 1915, by monotypy. Status: valid subgenus of Centorus Mulsant, 1854 in Lagriinae: Belopini.

Belopus Gebien, 459 [M]. Type species [automatic]: Tenebrioelongatus Herbst, 1797, by monotypy. Status: valid subgenus of Centorus Mulsant, 1854 in Lagriinae: Belopini. Note: replacement name for Calcar Dejean, 1821; nomen protectum (see Bouchard and Bousquet 2020b: 6).

Belousovia G.S. Medvedev, 157 [F]. Type Belousoviahelenae G.S. Medvedev, 2007, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Belutschistanops Löbl, Bouchard, Merkl & Bousquet, 3 [M]. Type Hyperopsschusteri Koch, 1940, by original designation. Status: valid subgenus of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: name first proposed by Koch (1943a: 523, 552) without a type species originally included (ICZN 1999, Article 13.3); Löbl et al. (2008a: 40) designated Hyperopsschusteri Koch, 1940 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Bequaertiella Pic, 486 [F]. Type Bequaertiellacoerulescens Pic, 1914, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Bermejoina Español, 12 [F]. Type Bermejoinaaiunica Español, 1944, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Betasida Reitter, 8, 11 [F]. Type Asidaluctuosa Boisduval, 1835, by subsequent designation (Viñolas and Cartagena 2005: 110). Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini. Note: some authors (e.g., F. Soldati 2008: 32) have used Asidaargenteolimbata Escalera, 1901 as type species for this genus; however, the older designation by Viñolas and Cartagena (2005: 110) should stand since Asidaluctuosa Boisduval, 1835 was one of the two originally included nominal species in Betasida.

Betschia Dajoz, 135 [F]. Type Betschiaminuta Dajoz, 1980, by original designation. Status: valid genus in Diaperinae: Gnathidiini: incertae sedis. Note: transferred from Tenebrionoidea: Zopheridae (as “Colydiidae”) by Ivie and Ślipiński (1990: 18); placed in “Gnathidiini incertae sedis” by Schawaller and Purchart (2012: 312).

Bia Hope, 132 [F]. Type species [automatic]: Trogossitathoracica Fabricius, 1792, by monotypy. Status: junior synonym of Bius Dejean, 1834 in Tenebrioninae: Tenebrionini. Note: unjustified emendation of Bius Dejean, 1834, not in prevailing usage.

Bielawskia Marcuzzi, 179 [F]. Type Bielawskiacubana Marcuzzi, 1985 (= Trimytantrondecui Ardoin, 1977), by monotypy. Status: junior synonym of Trimytantron Ardoin, 1977 in Pimeliinae: Edrotini. Synonymy: Marcuzzi (1998: 153).

Biolus Mulsant & Rey, 25 [M]. Type Eurynotusasperipennis Mulsant & Rey, 1854 (= Helopsgranulatus Fabricius, 1787), by subsequent designation (Koch 1953a: 276). Status: valid subgenus of Eurynotus W. Kirby, 1819 in Blaptinae: Platynotini: Eurynotina.

Biomorphus Motschulsky, 38 [M]. Type Biomorphustuberculatus Motschulsky, 1872 (= Amphidoraattenuata J.L. LeConte, 1851), by original designation. Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Aalbu et al. (1995: 485).

Bionesus Fairmaire, 70 [M]. Type Bionesuscinereosparsus Fairmaire, 1879, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Bioplanes Mulsant, 144 [M]. Type Bioplanesmeridionalis Mulsant, 1854, by monotypy. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Bioramix Bates, 478 [M]. Type Bioramixovalis Bates, 1879, by subsequent designation (Kaszab 1940a: 175). Status: valid genus and subgenus in Blaptinae: Platyscelidini. Note: nomenclatural stability in this genus is threatened by the discovery of an older type species designation (Bioramixpamirensis Bates, 1879, by subsequent designation by R. Lucas (1920: 138), which is currently included in the valid subgenusPlanoplatyscelis Kaszab, 1940); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Kaszab (1940a: 175); taxon also described as a new taxon with the same species by Bates (1890: 69).

Birolagria Pic, 86 [F]. Type Birolagriacicatricosa Pic, 1956 (= Cerogriaoriunda Borchmann, 1924), by monotypy. Status: junior synonym of Acerogria Borchmann, 1936 in Lagriinae: Lagriini: Lagriina. Synonymy: Merkl (1987: 124).

Biroum Kaszab, 104 [N]. Type Biroumparadoxum Kaszab, 1956, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bius Dejean, 205 [M]. Type Trogossitathoracicus Fabricius, 1792, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Blacodatus Koch, 42 [M]. Type Blacodesvertagus Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Blacodes Duponchel, 590 [M]. Type Pedinussulcatus Laporte, 1840, by monotypy. Status: junior synonym of Blenosia Laporte, 1840 in Blaptinae: Opatrini: Stizopodina. Synonymy: Erichson (1843: 245). Note: name previously attributed to Blanchard (1845: 13) in the literature.

Blapida Perty, 58 [F]. Type Blapidaokeni Perty, 1830, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Blapidium Bauer, 231 [N]. Type Blaps ocreata Allard, 1880, by subsequent designation (Nabozhenko and Chigray 2020: 10). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10). Note: originally proposed without included nominal species; Kolbe (1928: 200), by including four species in association with this name, was the first author to subsequently and expressly included nominal species in Blapidium (ICZN 1999, Article 67.2.2).

Blapidocamaria Pic, 7 [F]. Type Pseudoblapidaincostata Pic, 1919, by monotypy. Status: valid subgenus of Pseudoblapida Pic, 1917 in Stenochiinae: Cnodalonini.

Blapidocampsia Pic, 3 [F]. Type Campsiapallidipes Pic, 1918, by present designation. Status: valid subgenus of Campsia Lepeletier & Audinet-Serville, 1828 in Stenochiinae: Cnodalonini.

Blapidurus Fairmaire, xcvi [M]. Type Blapiduruscrassicornis Fairmaire, 1891, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Champion (1895: 48).

Blapimorpha Motschulsky, 531 [F]. Type Blapsreflexa Gebler, 1832, by subsequent designation (Nabozhenko 2008: 35). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860).

Blapisa Motschulsky, 530 [F]. Type Blaps jaegeri Hummel, 1827 (= Tenebriomortisagus Linnaeus, 1758), by subsequent designation (Nabozhenko 2008: 35). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860).

Blaposodes Skopin, 47 [M]. Type Blaps baerii Fischer von Waldheim, 1842, by original designation. Status: junior synonym of Peltarium Fischer von Waldheim, 1844 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Blaps Fabricius, 254 [F]. Type Tenebriomortisagus Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Blaptinae: Blaptini: Blaptina.

Blapstinus Dejean, 66 [M]. Type Blapspunctata Fabricius, 1792, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Blaptogonia G.S. Medvedev, 186 [F]. Type Tagonoidescostulata Fairmaire, 1901, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Blaptoprosodes Reitter, 120 [M]. Type Prosodesmucronata Reitter, 1893, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: G.S. Medvedev (2001: 54).

Blapylis Horn, 304, 315 [F]. Type Eleodescordata Eschscholtz, 1829, by subsequent designation (Bousquet et al. 2018: 143). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Blastarnodes Koch, 154 [M]. Type Blastarnodesherero Koch, 1958, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Blastarnus Fairmaire, 132 [M]. Type Blastarnusgrallator Fairmaire, 1897, by subsequent designation (Oustalet 1898: 972). Status: junior synonym of Diestecopus Solier, 1848 in Blaptinae: Pedinini: Helopinina. Synonymy: Koch (1958: 152).

Blatticephalus Heller, 377 [M]. Type Blatticephalusadelotopus Heller, 1918, by monotypy. Status: valid genus in Tenebrioninae: Falsocossyphini.

Blenosia Laporte, 209 [F]. Type Pedinussulcatus Laporte, 1840, by subsequent designation (R. Lucas 1920: 140). Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Blepegenes Pascoe, xii [M]. Type Blepegenesaruspex Pascoe, 1868, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Blepusa Westwood, 69 [F]. Type Blepusacostata Westwood, 1842, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Blindus Mulsant & Rey, 206 [M]. Type Pedinusstrigosus Faldermann, 1835, by monotypy. Status: valid subgenus of Pedinus Latreille, 1797 in Blaptinae: Pedinini: Pedinina.

Bluops Carter, 227 [M]. Type Bluopsverrucosus Carter, 1914, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Bobina Novák, 125 [F]. Type Bobinajendeki Novák, 2015, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Bobisthes Novák, 178 [M]. Type Bobisthesbellator Novák, 2019, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Bogatshevia G.S. Medvedev & Iwan, 618 [F]. Type species [automatic]: Achaemenesvillosus Bogatchev, 1949 (= Thripterabogatchevi Kwieton, 1982), by original designation. Status: valid genus in Pimeliinae: Pimeliini. Note: replacement name for Achaemenes Bogatchev, 1949 (as “Achaemenus”).

Bolbophanes Carter, 86 [M]. Type Paraphanesdumbrelli Lea, 1895, by subsequent designation (Gebien 1941: 1133). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Bolbostetha Fairmaire, 117 [F]. Type Bolbostethasoleata Fairmaire, 1896, by subsequent designation (Novák and Pettersson 2008: 320). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Boletophagus Agassiz, 48 [M]. Type species [automatic]: Silphareticulata Linnaeus, 1767, by subsequent designation (C.G. Thomson 1859: 115). Status: junior synonym of Bolitophagus Illiger, 1798 in Tenebrioninae: Bolitophagini. Note: unjustified emendation of Bolitophagus Illiger, 1798, not in prevailing usage.

Boletoxenus Motschulsky, 63 [M]. Type Boletoxenusgibber Motschulsky, 1858, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolithophilus Gistel, 137 [M]. Type species [automatic]: Cistelascapularis Illiger, 1805 (= Cistelahumeralis Fabricius, 1787), by subsequent designation (Westwood 1838: 32). Status: junior synonym of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: replacement name for Mycetophila Gyllenhal, 1810.

Bolitolaemus Gebien, 23 [M]. Type Bolitolaemuscatenulatus Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolitonaeus Lewis, 387 [M]. Type Bolitonaeusmergae Lewis, 1894, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolitopertha Gebien, 379 [F]. Type Bolitoperthanovemcostata Gebien, 1910 (= Bolitophagusborbonicus Fairmaire, 1880), by monotypy. Status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: Gebien (1922b: 268, with Cherostus C.O. Waterhouse, 1894, a junior synonym of Rhipidandrus J.L. LeConte, 1862).

Bolitophagiella Miyatake, 68, 71 [F]. Type Bolitophaguspannosus Lewis, 1894, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolitophagus Illiger, 100 [M]. Type Silphareticulata Linnaeus, 1767, by subsequent designation (C.G. Thomson 1859: 115). Status: valid genus in Tenebrioninae: Bolitophagini. Note: as mentioned by Bouchard and Bousquet (2020b: 8) nomenclatural stability in this genus is threatened by the discovery of an older type species designation (Opatrumagricola Herbst, 1783, by subsequent designation by Curtis (1836: pl. 586), which is currently the type species of the valid genus Eledona Latreille, 1797); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by C.G. Thomson (1859: 115).

Bolitotherus Candèze, 367 [M]. Type Bolitophaguscornutus Fabricius, 1801, by subsequent designation (J.L. LeConte 1862: 236). Status: valid genus in Tenebrioninae: Bolitophagini. Note: the younger species name Bolitophaguscornutus Fabricius, 1801 was given priority over the older synonym Opatrumbifurcum Fabricius, 1798 by the ICZN (2019, Opinion 2438).

Bolitotrogus Miyatake, 68, 80 [M]. Type Bolitotroguskurosonis Miyatake, 1964, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolitoxenus Gemminger in Gemminger and Harold, 1946 [M]. Type species [automatic]: Boletoxenusgibber Motschulsky, 1858, by monotypy. Status: junior synonym of Boletoxenus Motschulsky, 1858 in Tenebrioninae: Bolitophagini. Note: unjustified emendation of Boletoxenus Motschulsky, 1858, not in prevailing usage.

Bolitrium Gebien, 390 [N]. Type Bolitriumcrenulicolle Gebien, 1914, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Bolusculus Matthews, 704, 798 [M]. Type Bolusculusarcanus Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Bombocnodulus Koch, 36 [M]. Type Psammodescrinicollis Haag- Rutenberg, 1879, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Borbochara Novák, 259 [F]. Type Borbocharabicolor Novák, 2009, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Borbonalia Novák, 136 [F]. Type Borbonaliabrancuccii Novák, 2014, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Borborella Novák, 196 [F]. Type Borborellahergovitsi Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Borboresthes Fairmaire, 253 [F]. Type Alleculacruralis Marseul, 1876, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: see Löbl and Smetana (2013: 39) for comments on the gender of this name.

Borchmannia Pic in Borchmann and Pic, 35 [F]. Type Borchmannialineaticeps Pic, 1912, by subsequent designation (R. Lucas 1920: 144). Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Borchmannius Bousquet & Bouchard in Bousquet et al., 2015: 142 [M]. Type species [automatic]: Glyptothoraxpilosus Borchmann, 1937, by original designation. Status: valid genus in Alleculinae: incertae sedis. Note: replacement name for Glyptothorax Borchmann, 1937.

Boreoptorina G.S. Medvedev, 417 [F]. Type Gnaptorinacordicollis G.S. Medvedev, 1998, by original designation. Status: junior synonym of Hesperoptorina G.S. Medvedev, 2009 in Blaptinae: Blaptini: Gnaptorinina. Synonymy and First Reviser action (Boreoptorina G.S. Medvedev, 2009 versus Hesperoptorina G.S. Medvedev, 2009) by Li et al. (2021: 245).

Boreosaragus Matthews, 1040, 1067 [M]. Type Saraguslugubris Lea, 1897, by original designation. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Borneocamaria Pic, 17 [F]. Type Borneocamariaatra Pic, 1917, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Borneocistela Pic, 18 [F]. Type Borneocisteladiversicornis Pic, 1922, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Borneogonocnemis Pic, 16 [F]. Type Borneogonocnemisruficolor Pic, 1936, by monotypy. Status: valid subgenus of Paragonocnemis Kraatz, 1899 in Tenebrioninae: Amarygmini.

Borneolaena Schawaller, 2 [F]. Type Borneolaenariedeli Schawaller, 1998, by original designation. Status: valid genus in Lagriinae: Laenini.

Borneosphaerotus Grimm, 218 [M]. Type Borneosphaerotussantubongicus Grimm, 2015, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Borneosphena Purchart & Grimm, 522 [F]. Type Borneosphenafouquei Purchart & Grimm, 2016, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Borneostira Pic, 53 [F]. Type Rouyerusbrevilineatus Pic, 1912, by monotypy. Status: valid subgenus of Rouyerus Pic, 1911 in Lagriinae: Lagriini: Statirina.

Borneosynopticus Grimm, 219 [M]. Type Borneosynopticustubericollis Grimm, 2015, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Boromorphus Wollaston, 492 [M]. Type Boromorphusmaderae Wollaston, 1854 (= Borostagenioides P.H. Lucas, 1846), by monotypy. Status: valid genus in Pimeliinae: Boromorphini.

Bothrasida Casey, 76, 122 [F]. Type Asidaclathrata Champion, 1884, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Bothrichara Borchmann, 48, 104 [F]. Type Lagriapulchella Guérin-Méneville, 1830, by subsequent designation (Borchmann 1936: 68). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Bothrionota Borchmann, 16, 66 [F]. Type Bothrionotafoveata Borchmann, 1930, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Bothriostira Borchmann, 239, 473 [F]. Type Bothriostiracylindracea Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Bothrotes Casey, 379 [M]. Type Epitraguscanaliculatus Say, 1824, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Bothynocara Gebien, 167, 170 [N]. Type Bothynocaratenuepunctatum Gebien, 1928, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bothynocephalus Doyen, 315 [M]. Type Bothynocephaluscristatus Doyen, 1988, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bothynogria Borchmann, 128 [F]. Type Bothynogriacalcarata Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Botiras Fairmaire, xcviii [M]. Type Botirasstriatellus Fairmaire, 1891, by subsequent designation (R. Lucas 1920: 145). Status: junior synonym of Chianalus Bates, 1879 in Blaptinae: Platyscelidini. Synonymy: Blair (1923a: 283).

Bouchardandrus Steiner, 543 [M]. Type Haplandrusconcolor J.L. LeConte, 1866, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini.

Bovius Gistel, ix [M]. Type species [automatic]: Tenebrioserratus Fabricius, 1775, by monotypy. Status: junior synonym of Prioscelis Hope, 1841 in Lagriinae: Pycnocerini. Note: unnecessary replacement name for Iphius Dejean, 1834.

Brachycilibe Carter, 207 [F]. Type Brachycilibeantennata Carter, 1911, by monotypy. Status: junior synonym of Dioedus J.L. LeConte, 1862 in Phrenapatinae: Penetini. Synonymy: Kaszab (1978a: 164).

Brachycryptus Quedenfeldt, 129 [M]. Type Brachycryptustripolitanus Quedenfeldt, 1891, by monotypy. Status: senior synonym of Tripolicryptus Strand, 1929 in Alleculinae: Cteniopodini. Note: junior homonym of Brachycryptus C.G. Thomson, 1873 [Hymenoptera].

Brachycula Fairmaire, 278 [F]. Type Brachyculaquadrivittata Fairmaire, 1906, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: the alternative original spelling Brachicula, used by Fairmaire (1906: 278), was rejected by Bousquet et al. (2015: 143) who acted as the First Revisers.

Brachycyphus Gebler, 473. Type Pimeliaimbricata Fischer, 1820, by monotypy. Status: junior synonym of Ocnera Fischer, 1822 in Pimeliinae: Pimeliini. Synonymy: Gemminger in Gemminger and Harold (1870: 1888).

Brachyesthes Fairmaire, 490 [F]. Type Melanesthespilosellus Marseul, 1866, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Brachygenius Dejean, 206 [M]. Type species [automatic]: Nyctelialuczotii Guérin-Méneville, 1831, by subsequent designation (Duponchel 1845b: 449). Status: junior synonym of Gyriosomus Guérin-Méneville, 1834 in Pimeliinae: Nycteliini. Note: unnecessary replacement name for Gyriosomus Guérin-Méneville, 1834.

Brachyidium Fairmaire, 33 [N]. Type Brachyidiumbreviusculum Fairmaire, 1883, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Brachymoschium Fairmaire, 348 [N]. Type Brachymoschiumparvitarse Fairmaire, 1896, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Brachyontis Casey, 82, 141 [F]. Type Coniontisglobulina Casey, 1895, by monotypy. Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Aalbu et al. (2002: 487).

Brachyphrynus Fairmaire in Fairmaire et al., 1882: 71 [M]. Type Brachyphrynusspissicornis Fairmaire, 1882, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Brachypilium Fairmaire, 23 [N]. Type Brachypiliumsculpturatum Fairmaire, 1896, by monotypy. Status: junior synonym of Stenochinus Motschulsky, 1860 in Stenochiinae: Cnodalonini. Synonymy: Löbl et al. (2008b: 349).

Brachypophlaeus Fairmaire, 113 [M]. Type Hypophlaeusdimidiatipennis Fairmaire, 1880, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Brachyscelis Dejean, 179 [F]. Type Pimeliamusiva Ménétriés, 1832, by subsequent designation (Bouchard et al. 2007: 388). Status: senior synonym of Pachyscelis Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Bouchard et al. (2007: 388). Note: the date of publication of the name Brachyscelis Germar, 1834 (Coleoptera: Chrysomelidae) is unknown whereas Brachyscelis Dejean was published by 30 June 1834; nomen oblitum (see Bouchard et al. 2007: 388).

Bradygena Fairmaire, 211 [F]. Type Nesogenaviolacea Fairmaire, 1903 (= Nesogenaepiscopalis Fairmaire, 1875), by subsequent designation (Froussart 1961: 91). Status: junior synonym of Nesogena Mäklin, 1863 in Tenebrioninae: Praeugenini. Synonymy: Gebien (1948: 549).

Bradymerus Perroud & Montrouzier, 110 [M]. Type Bradymerustuberculatus Perroud & Montrouzier, 1865, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Bradynocerus Fairmaire, 36 [M]. Type Bradynocerusaulacopterus Fairmaire, 1883, by monotypy. Status: junior synonym of Bradymerus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1939: 751).

Bradysphaerotus Kaszab, 293 [M]. Type Bradysphaerotuspapuanus Kaszab, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bradyus Dejean, 190 [M]. Type Erodiuspygmaeus Fischer, 1821, by monotypy. Status: valid genus in Tenebrioninae: Dissonomini.

Branchus J.L. LeConte, 222 [M]. Type Branchusfloridanus J.L. LeConte, 1862, by monotypy. Status: valid genus in Pimeliinae: Branchini.

Brasilius Gebien, 169, 178 [M]. Type Nyctobatesexaratus Fairmaire, 1889, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bratyna Westwood, 228 [F]. Type Bratynaapicalis Westwood, 1875, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Bremerianus Masumoto & Bečvář, 418 [M]. Type Bremerianuscameronensis Masumoto & Bečvář, 2005, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Bremerus Ferrer, 234 [M]. Type Bremerusfrankkochi Ferrer, 2004, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini.

Brinckia Koch, 117 [F]. Type Psammodesdebilis Péringuey, 1899, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Brises Pascoe, 145 [M]. Type Brisestrachynotoides Pascoe, 1869, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina. Note: the First Reviser (Brises Pascoe, 1869 versus Ephidonius Pascoe, 1869) is Carter (1914b: 46).

Brittona G.S. Medvedev & Lawrence, 574 [F]. Type Brittonaminuta G.S. Medvedev & Lawrence, 1986, by original designation. Status: valid genus in Diaperinae: Hyociini: Brittonina.

Broomium Koch, 338, 342 [N]. Type Broomiumnudum Koch, 1950, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Brosimapsida Ferrer & Ødegaard, 640 [F]. Type Brosimapsidagonospoides Ferrer & Ødegaard, 2005, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Brycopia Pascoe, 141 [F]. Type Brycopiapilosella Pascoe, 1869, by monotypy. Status: valid genus in Lagriinae: Adeliini. Note: the First Reviser (Brycopia Pascoe, 1869 versus Dinoria Pascoe, 1869) is Carter (1920a: 238).

Bulbulus Lesne, 227, 240 [M]. Type Arthrodeisbyrrhiformis Fairmaire, 1892, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Bunamarygmus Masumoto, 127 [M]. Type Bunamarygmusthailandicus Masumoto, 1988, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Burmanosis G.S. Medvedev, 845, 859 [F]. Type Stenosiskaszabi G.S. Medvedev, 1995, by original designation. Status: valid subgenus of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina.

Buxela Fairmaire, 28 [F]. Type Buxelasordescens Fairmaire, 1894, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Byallius Pascoe, 42 [M]. Type Byalliusreticulatus Pascoe, 1869, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Bycrea Pascoe, xii [F]. Type Bycreavillosa Pascoe, 1868, by monotypy. Status: junior synonym of Trichoton Hope, 1841 in Blaptinae: Opatrini: Blapstinina. Synonymy: Kamiński et al. (2019c: 360).

Byrrhoncus Koch, 49 [M]. Type Byrrhoncusmonticola Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Byrsax Pascoe, 42 [M]. Type Byrsaxcoenosus Pascoe, 1860 (= Boletophagusgibbifer Wesmael, 1836), by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini. Note: genus originally described in Tenebrionoidea: Zopheridae: Colydiinae.

Byzacnus Pascoe, 469 [M]. Type Byzacnuspicticollis Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cabirus Mulsant & Rey, 148, 223 [M]. Type Cabirusminutissimus Mulsant & Rey, 1853, by subsequent designation (Gebien 1938a: 314). Status: senior synonym of Cabirutus Strand, 1929 in Blaptinae: Pedinini: Pedinina. Note: junior homonym of Cabirus Hübner, 1819 [Lepidoptera].

Cabirutus Strand, 24 [M]. Type species [automatic]: Cabirusminutissimus Mulsant & Rey, 1853, by subsequent designation (Gebien 1938a: 314). Status: valid genus and subgenus in Blaptinae: Pedinini: Pedinina. Note: replacement name for Cabirus Mulsant & Rey, 1853.

Cacicus Dejean, 182 [M]. Type Elenophorusamericanus Lacordaire, 1830, by monotypy. Status: senior synonym of Megelenophorus Gebien, 1910 in Pimeliinae: Elenophorini: Megelenophorina. Note: junior homonym of Cacicus Lacépède, 1799 [Aves].

Cacoplesia Fairmaire, 237 [F]. Type species [automatic]: Plesiamelanura Klug, 1833, by subsequent designation (Bousquet et al. 2015: 137). Status: junior synonym of Eubalia Laporte, 1840 in Alleculinae: Alleculini: Gonoderina. Synonymy: Borchmann (1909a: 713). Note: replacement name for Plesia Klug, 1833.

Caecochares Koch, 91 [M]. Type Caecocharesgrjebinei Koch, 1956, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Caecomenimopsis Kaszab, 198 [F]. Type Caecomenimopsisleleupi Kaszab, 1970, by original designation. Status: junior synonym of Menimopsis Champion, 1896 in Diaperinae: Gnathidiini: Anopidiina. Synonymy: Peck (1990: 370).

Caecophloeus Dajoz, 278 [M]. Type Caecophloeusfranzi Dajoz, 1972, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Caediexis Lebedev, 125 [F]. Type Caediexisarenicola Lebedev, 1932, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Caediomorpha Blackburn, 272 [F]. Type Caediomorphaaustralis Blackburn, 1888 (= Morychusheteromerus King, 1869), by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: moved from the subtribe Ammobiina to Opatrina by Lumen et al. (2020: 346).

Caedius Blanchard, 13 [M]. Type Opatrumsphaeroides Hope, 1843, by subsequent designation (Lacordaire 1859a: 262). Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: Caedius is an incorrect subsequent spelling of the original spelling Coedius, first used by Lacordaire (1859a: 261), and in prevailing usage; Caedius is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Caenoblaps König, 24 [F]. Type Caenoblapsdifformis König, 1906, by monotypy. Status: junior synonym of Dila Fischer von Waldheim, 1844 in Blaptinae: Blaptini: Blaptina. Synonymy: Chigray et al. (2019: 3).

Caenocapicus Endrödy-Younga, 15, 19 [M]. Type Caenocrypticuscapensis Endrödy-Younga, 1996, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini.

Caenocorse C.G. Thomson, 117 [F]. Type Hypophlaeusdepressus Fabricius, 1790, by original designation. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Bedel (1906a: 92).

Caenocrypticoides Kaszab, 322 [M]. Type Caenocrypticoidesloksai Kaszab, 1969, by original designation. Status: valid genus in Pimeliinae: Caenocrypticini. Note: we act as First Revisers and reject the alternative original spelling Caenocripticoides, used by Kaszab (1969b: 324).

Caenocrypticus Gebien, 139 [M]. Type Caenocrypticusuncinatus Gebien, 1920, by monotypy. Status: valid genus and subgenus in Pimeliinae: Caenocrypticini.

Calabosca Fairmaire, 395 [F]. Type species [automatic]: Ascalabuspedinoides Fairmaire, 1893, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Ascalabus Fairmaire, 1893.

Calaharena Koch, 64 [F]. Type Calaharenadutoiti Koch, 1963, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Calcar Dejean, 67 [N]. Type Tenebrioelongatus Herbst, 1797, by monotypy. Status: senior synonym of Belopus Gebien, 1911 in Lagriinae: Belopini. Note: junior homonym of Calcar de Montfort, 1810 [Mollusca].

†Calcarocistela Nabozhenko in Nabozhenko et al., 2016a: 1421 [F]. Type Calcarocistelakirejtshuki Nabozhenko, 2016, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: described from Lower Cretaceous deposits (China).

Calcarosis Penrith, 21, 98 [F]. Type Zophosismichaelis Penrith, 1977, by original designation. Status: junior synonym of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Synonymy: Penrith (1981a: 22).

Callicomus Motschulsky, 138 [M]. Type Diaperisriederii Faldermann, 1833, by subsequent designation (Jakobson 1924: 243). Status: senior synonym of Emypsara Pascoe, 1866 in Diaperinae: Phaleriini. Synonymy: Jakobson (1924: 243). Note: junior homonym of Callicomus Agassiz, 1846 [Coleoptera: Cerambycidae].

Callignathus Agassiz, 58, 60 [M]. Type species [automatic]: Calognathuschevrolatii Guérin-Méneville, 1836, by monotypy. Status: junior synonym of Calognathus Guérin-Méneville, 1836 in Pimeliinae: Cryptochilini: Calognathina. Note: unjustified emendation of Calognathus Guérin-Méneville, 1836, not in prevailing usage.

Callimaria Fairmaire, 12 [F]. Type Callimariaimpressipennis Fairmaire, 1888, by monotypy. Status: junior synonym of Actanorie Bates, 1879 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1956b: 90).

Callismilax Bates, 105 [F]. Type Leptomorphaaenea Montrouzier, 1860, by subsequent designation (Gebien 1942a: 333). Status: valid genus in Tenebrioninae: Titaenini. Note: Matthews and Lawrence (2019: 637) mentioned that this genus is not distinguishable from Titaena Erichson, 1842.

Callyntra Solier, 307, 335 [F]. Type Nycteliamulticosta Guérin-Méneville, 1834, by subsequent designation (Duponchel 1842b: 63). Status: valid genus in Pimeliinae: Nycteliini.

Calobamon Kraatz, 80, 105 [M]. Type Calobamonleptoderus Kraatz, 1865, by monotypy. Status: senior synonym of Thraustocolus Kraatz, 1866 in Pimeliinae: Tentyriini. Note: junior homonym of Calobamon Loew, 1850 [Diptera].

Calognathus Guérin-Méneville, pl. 172 [M]. Type Calognathuschevrolatii Guérin-Méneville, 1836, by monotypy. Status: valid genus in Pimeliinae: Cryptochilini: Calognathina.

Calogria Borchmann, 48, 107 [F]. Type Calogriacostata Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Calosis Deyrolle, 81, 222 [F]. Type Calosisamabilis Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Note: the alternative original spelling Calliosis (p. 81) was corrected to Calosis in the “Errata” of the same work (p. 248), Calosis is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Calostega Gemminger in Gemminger and Harold, 1991 [F]. Type species [automatic]: Calostegiapurpuripennis Westwood, 1843, by monotypy. Status: junior synonym of Calostegia Westwood, 1843 in Lagriinae: Pycnocerini. Note: unjustified emendation of Calostegia Westwood, 1843, not in prevailing usage.

Calostegia Westwood, 117 [F]. Type Calostegiapurpuripennis Westwood, 1843, by monotypy. Status: valid genus in Lagriinae: Pycnocerini. Note: redescribed as a new genus by Westwood (1844: 221, as “Calostega”).

Calous Koch, 151 [M]. Type Blastarnusmichaelseni Gebien, 1920, by original designation. Status: valid subgenus of Nicandra Fairmaire, 1888 in Blaptinae: Pedinini: Helopinina.

Calydonella Doyen, 8 [F]. Type Calydonellalisa Doyen, 1995, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Calydoniomorpha Pic, 19 [F]. Type Calydoniomorphabrevicornis Pic, 1917, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Calydonis Pascoe, 31 [F]. Type Calydonisrefulgens Pascoe, 1882, by subsequent designation (Gebien 1919: 141). Status: valid genus in Stenochiinae: Cnodalonini.

Calymmatophorus Gemminger in Gemminger and Harold, 1904 [M]. Type species [automatic]: Praociscucullatus Guérin-Méneville, 1834, by subsequent designation (R. Lucas 1920: 161). Status: junior synonym of Calymmophorus Solier, 1841 in Pimeliinae: Praociini. Note: unjustified emendation of Calymmophorus Solier, 1841 (as “Calymmaphorus”), not in prevailing usage.

Calymmophorus Solier, 209, 245 [M]. Type Praociscucullata Guérin-Méneville, 1834, by subsequent designation (R. Lucas 1920: 161). Status: valid genus in Pimeliinae: Praociini. Note: two spellings were originally used, Calymmaphorus (pp. 209, 245, 246, 247, 250, 370) and Galymmaphorus (p. 247, in note); however, the incorrect subsequent spelling Calymmophorus, first introduced by Burmeister (1875: 494), is in prevailing usage and deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Calymmus Montrouzier, 289 [M]. Type Toxicumberardi Montrouzier, 1860, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina. Note: the name Calymmus was listed as a synonym of Toxicum Latreille, 1802 by Montrouzier (1860: 289); it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Fauvel 1862: 148); therefore, Calymmus was made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Calyptopsis Solier, 253, 269 [F]. Type Calyptopsisemondi Solier, 1835 (= Hegetercaraboides Brullé, 1832), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Camaria Lepeletier & Audinet-Serville, 454 [F]. Type Camarianitida Lepeletier & Audinet-Serville, 1828 (= Tenebrionitens G.-A. Olivier, 1795), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camarimena Motschulsky, 473 [F]. Type Camarimenaovicauda Motschulsky, 1863, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camariocropterum Pic, 16 [N]. Type Camariocropterumlaticeps Pic, 1920, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camariodes Fairmaire, 232 [M]. Type Camariodescoquerelii Fairmaire, 1869 (= Camariahelopioides Klug, 1833), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: the First Reviser (Camariodes Fairmaire, 1869 versus Tinophyllus Fairmaire, 1869) is Fairmaire (1886c: 75).

Camariomorpha Pic, 7 [F]. Type Camariomorphasingularis Pic, 1915, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camarothelops Kolbe, 30 [M]. Type Camarothelopsbraueri Kolbe, 1910, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: removed from the tribe Helopini and placed as Tenebrionidae incertae sedis by Nabozhenko (2018: 183).

Camphonota Solier, 195 [F]. Type Tenebriosubglobosus Pallas, 1781, by subsequent designation (Lacordaire 1859a: 188). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Campolene Pascoe, 44 [F]. Type Campolenenitida Pascoe, 1863, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camponotiphilus Lea, 257 [M]. Type Camponotiphilusfimbricollis Lea, 1914, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Campsia Lepeletier & Audinet-Serville, 455 [F]. Type Cnodalonirroratum Dalman, 1823, by subsequent designation (Hope 1841: 133). Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Campsiomorpha Pic, 19 [F]. Type Camarialata Pic, 1915, by subsequent designation (Gebien 1942a: 323). Status: valid genus in Stenochiinae: Cnodalonini.

Camptobrachys Kaszab, 4, 24 [M]. Type Camptobrachyssulcatus Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Canariella Uyttenboogaart, 341, 342 [F]. Type Canariellaarenapta Uyttenboogaart, 1929 (= Philhammussericans Fairmaire, 1871), by monotypy. Status: junior synonym of Philhammus Fairmaire, 1871 in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Synonymy: Uyttenboogaart (1940: 68). Note: junior homonym of Canariella Hesse, 1918 [Mollusca].

Cantaloubeus Ardoin, 203 [M]. Type Cantaloubeusviridis Ardoin, 1959, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Cantopipleurus Koch, 578 [M]. Type Tentyriamesostenoides Baudi di Selve, 1881, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Capeluprops Schawaller, 271, 273 [M]. Type Capelupropslaenoides Schawaller, 2011, by original designation. Status: valid genus in Lagriinae: Lupropini.

Capicrypticus Koch, 55 [M]. Type Lamprocrypticuscapensis Koch, 1950, by original designation. Status: valid genus in Diaperinae: Crypticini.

Capidium Koch, 44 [N]. Type Oncotustardus Solier, 1848, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Capnisa Dejean, 197 [F]. Type Bradyuskarelini Faldermann, 1836, by monotypy. Status: junior synonym of Gnathosia Fischer, 1821 in Pimeliinae: Tentyriini. Synonymy: Reitter (1896d: 129).

Capnisiceps Chatanay, 215 [M]. Type Capnisicepsmaindroni Chatanay, 1914, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Capnochroa J.L. LeConte, 244 [F]. Type Cistelafuliginosa Melsheimer, 1846, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Capricephalius Koch, 492, 508 [M]. Type Arthrodibiusbazmanicus Schuster, 1938, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Carabelops Fairmaire, 534 [M]. Type Carabelopsaenescens Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Caraboblaps Bauer, 231 [F]. Type none designated. Status: undetermined taxon in Blaptinae: Blaptini: Blaptina. Note: this genus was described before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Caraboblaps and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Caracasa Pic, 21 [F]. Type Caracasaaeneipennis Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Carchares Pascoe, 12 [M]. Type Carcharesmacer Pascoe, 1887, by monotypy. Status: valid genus in Tenebrioninae: Scaurini.

Cardigenius Solier, 407, 492 [M]. Type Cardigeniuslaticollis Solier, 1836, by subsequent designation (Wilke 1922: 276). Status: valid genus and subgenus in Pimeliinae: Asidini.

Cardiobioramix Kaszab, 154, 183 [M]. Type Bioramixasidioides Bates, 1879, by original designation. Status: junior synonym of Euryhelops Reitter, 1902 in Blaptinae: Platyscelidini. Synonymy: Nabozhenko and Egorov (2020: 266).

Cardiochianalus Kaszab, 150, 202 [M]. Type Botirassculptipennis Fairmaire, 1891, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini. Note: the alternative original spelling Chardiochianalus, used by Kaszab (1940a: 202), was rejected by Kaszab (1975a: 4) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Cardiogenius Agassiz, 65, 66 [M]. Type species [automatic]: Cardigeniuslaticollis Solier, 1836, by subsequent designation (Wilke 1922: 276). Status: junior synonym of Cardigenius Solier, 1836 in Pimeliinae: Asidini. Note: unjustified emendation of Cardigenius Solier, 1836, not in prevailing usage.

Cardiosis Deyrolle, 81, 235 [F]. Type Cardiosismouffleti Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Cardiothorax Motschulsky, 67 [M]. Type species [automatic]: Thoracophoruswalckenaerii Hope, 1841, by original designation. Status: valid genus in Lagriinae: Adeliini. Note: replacement name for Thoracophorus Hope, 1841.

Caribanosis Nabozhenko, Kirejtshuk, Merkl, Varela, Aalbu & Smith, 568 [F]. Type Rhypasmaquisqueyanus Garrido & Varela, 2011, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Carinosella Purchart, 254 [F]. Type Carinosellamaasaiorum Purchart, 2010, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Caristela Fairmaire, 311 [F]. Type Caristelamegalops Fairmaire, 1894, by monotypy. Status: junior synonym of Mycetocharina Seidlitz, 1890 in Alleculinae: Alleculini: Alleculina. Synonymy: Bedel (1897: 36).

Caroliphosis Penrith, 114, 115 [F]. Type Hesseosisadamantina Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Carpiella Koch, 148 [F]. Type Carpiellalatisternum Koch, 1962, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

†Caryosoma Haupt, 114, 125 [N]. Type Caryosomarugosum Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Casnonidea Fairmaire, 264 [F]. Type Casnonideaholomelaena Fairmaire, 1882, by subsequent designation (Borchmann 1930a: 444). Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Catamerus Fairmaire, 290 [M]. Type Catamerusrevoili Fairmaire, 1887, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Cataphanus Gebien, 325, 346 [M]. Type Cataphanusquadraticollis Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cataphronetis P.H. Lucas, pl. 30 [F]. Type Cataphronetislevaillantii P.H. Lucas, 1846 (= Phtoracrenata Germar, 1836), by monotypy. Status: junior synonym of Phtora Germar, 1836 in Diaperinae: Phaleriini. Synonymy: Lacordaire (1859a: 335).

Catapiestus Perty, xxxviii [M]. Type Catapiestuspiceus Perty, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Catobleps Blair, 149 [M]. Type Catoblepsblattoides Blair, 1918 (= Blatticephalusadelotopus Heller, 1918), by original designation. Status: junior synonym of Blatticephalus Heller, 1918 in Tenebrioninae: Falsocossyphini. Synonymy: Blair (1919c: 102).

Catolaena Reitter, 282 [F]. Type Laenaturkestanica Reitter, 1897, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Laena Dejean, 1821 in Lagriinae: Laenini. Synonymy: Schawaller (2001b: 4).

Catomidius Seidlitz, 791, 792 [M]. Type Hedyphanesrhynchophorus Seidlitz, 1895, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Catomus Allard, 1876 in Tenebrioninae: Helopini: Helopina. Synonymy: Reitter (1922a: 7).

Catomodontus Löbl & Merkl in Löbl et al., 2020: 2 [M]. Type Catomuscoronatus Koch, 1935, by original designation. Status: valid subgenus of Catomus Allard, 1876 in Tenebrioninae: Helopini: Helopina. Note: the name was first proposed by Koch (1935: 107, 108) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 251) designated Catomuscoronatus Koch, 1935 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Catomulus Reitter, 302 [M]. Type Catomulussubaeneus Reitter, 1897 (= Oxycaraolcesii Fairmaire, 1883), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Catomus Allard, 4 [M]. Type Catomuspersicus Allard, 1876, by subsequent designation (Gebien 1943: 409). Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina.

Catopherus Carter, 713 [M]. Type Catopheruscorpulentus Carter, 1918 (= Axynaonchampioni Blackburn, 1897), by monotypy. Status: junior synonym of Axynaon Blackburn, 1897 in Tenebrioninae: Amarygmini. Synonymy: Carter (1920a: 249).

Caucasohelops Nabozhenko, 816 [M]. Type Eustenomacidiussvetlanae Nabozhenko, 2006, by original designation. Status: valid subgenus of Eustenomacidius Nabozhenko, 2006 in Tenebrioninae: Helopini: Cylindrinotina.

Caucasonotus Nabozhenko, 107 [M]. Type Cylindronotusdombaicus Nabozhenko, 2000, by original designation. Status: valid subgenus of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina.

Caudamarygmus Bremer, 88, 94 [M]. Type Caudamarygmusnotabilis Bremer, 2001, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Caulostena Fairmaire, 355 [F]. Type Caulostenafoveicollis Fairmaire, 1896, by monotypy. Status: valid genus in Alleculinae: Alleculini: Mycetocharina.

Cauricara Penrith, 7, 42 [N]. Type Stenocaravelox Péringuey, 1886, by original designation. Status: valid subgenus of Stenocara Solier, 1835 in Pimeliinae: Adesmiini.

Caverneleodes Triplehorn, 39 [M]. Type Eleodeseasterlai Triplehorn, 1975, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Caxtonana Buck, 224 [F]. Type Caxtonanacostata Buck, 1960, by monotypy. Status: junior synonym of Apterotheca Gebien, 1921 in Stenochiinae: Cnodalonini. Synonymy: Bouchard (2002: 452).

Cechenosternum Gebien, 9 [N]. Type Cechenosternumnigromaculatum Gebien, 1921, by subsequent designation (Gebien 1939: 748). Status: valid genus in Diaperinae: Crypticini.

Cecrops Gistel, 21 [M]. Type Tenebriogigas Linnaeus, 1763, by subsequent designation (Bousquet and Bouchard 2017: 131). Status: junior synonym of Mylaris Pallas, 1781 in Stenochiinae: Cnodalonini. Synonymy: Bousquet and Bouchard (2017: 131).

Cedrosius Fairmaire, 332 [M]. Type Cedrosiuscalosomoides Fairmaire, 1902, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Celebesa Pic, 23 [F]. Type Celebesaelongata Pic, 1921, by subsequent designation (Riley 1923: 126). Status: valid genus in Stenochiinae: Cnodalonini.

Celibe Boisduval, 262 [F]. Type Celibeaustralis Boisduval, 1835, by subsequent designation (Watt 1968: 36). Status: valid genus in Tenebrioninae: Heleini: Heleina.

Celox Gistel, 126 [F]. Type species [automatic]: Cnodalonirroratum Dalman, 1823, by subsequent designation (Hope 1841: 133). Status: junior synonym of Campsia Lepeletier & Audinet-Serville, 1828 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Campsia Lepeletier & Audinet-Serville, 1828.

Cenophorus Mulsant & Rey, 113 [M]. Type Cenophorusviduus Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Cenoscelis Wollaston, 179 [F]. Type Cenoscelistibialis Wollaston, 1868 (= Ulomapullum Erichson, 1843), by monotypy. Status: valid genus and subgenus in Tenebrioninae: Ulomini.

Centorus Mulsant, 272 [M]. Type Calcarprocerum Mulsant, 1854, by monotypy. Status: valid genus and subgenus in Lagriinae: Belopini.

Centrioptera Mannerheim, 279 [F]. Type Centriopteracaraboides Mannerheim, 1843, by monotypy. Status: junior synonym of Cryptoglossa Solier, 1837 in Pimeliinae: Cryptoglossini. Synonymy: Aalbu et al. (2002: 486).

Centrocnemis Kraatz in Heyden and Kraatz, 330 [F]. Type Centrocnemismollis Kraatz, 1882, by monotypy. Status: senior synonym of Centrocnemita Strand, 1935 in Pimeliinae: Pimeliini. Note: junior homonym of Centrocnemis Signoret, 1852 [Hemiptera].

Centrocnemita Strand, 284 [F]. Type species [automatic]: Centrocnemismollis Kraatz, 1882, by monotypy. Status: valid subgenus of Lasiostola Dejean, 1834 in Pimeliinae: Pimeliini. Note: replacement name for Centrocnemis Kraatz, 1882.

Centronopus Solier, 153, 154, 258 [M]. Type Centronopusextensicollis Solier, 1848 (= Tenebriosuppressus Say, 1835), by original designation. Status: valid genus and subgenus in Tenebrioninae: Centronopini.

Centropus Jakobson, 528 [M]. Type species [automatic]: Centronopusextensicollis Solier, 1848 (= Tenebriosuppressus Say, 1835), by original designation. Status: junior synonym of Centronopus Solier, 1848 in Tenebrioninae: Centronopini. Note: unjustified emendation of Centronopus Solier, 1848, not in prevailing usage; junior homonym of Centropus Illiger, 1811 [Aves].

Cephacerus Rafinesque, 113 [M]. Type species [automatic]: Erodiusgibbus Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Erodius Fabricius, 1775 in Pimeliinae: Erodiini. Note: unnecessary replacement name for Erodius Fabricius, 1775.

Cephaladesmia Gebien, 49, 61 [F]. Type Cephaladesmiathomseni Gebien, 1920, by monotypy. Status: junior synonym of Stenodesia Reitter, 1916 in Pimeliinae: Adesmiini. Synonymy: Penrith (1979: 70, 85).

Cephalamarygmus Bremer, 92, 95 [M]. Type Amarygmuspreangerensis Pic, 1952, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Cephaleucyrtus Pic, 22 [M]. Type Cephaleucyrtusviridicollis Pic, 1923, by monotypy. Status: junior synonym of Gauromaia Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1941: 1144).

Cephaloplonyx Pic, 12 [M]. Type Cephaloplonyxopacus Pic, 1922 (= Helopsdentipes Fabricius, 1781), by monotypy. Status: junior synonym of Hoplobrachium Fairmaire, 1886 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1943: 922).

Cephalostenus Solier, 160, 184 [M]. Type Cephalostenusdejeanii Solier, 1838 (= Scauruselegans Brullé, 1832), by subsequent designation (Hope 1841: 115). Status: valid genus in Tenebrioninae: Scaurini.

Cephalothydemus Pic, 24 [M]. Type Cephalothydemustheresae Pic, 1923, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ceradelium Preudhomme de Borre, 126 [N]. Type Ceradeliumarmatum Preudhomme de Borre, 1868 (= Blepegenesaruspex Pascoe, 1868), by monotypy. Status: junior synonym of Blepegenes Pascoe, 1868 in Lagriinae: Adeliini. Synonymy: Pascoe (1869: 41).

Ceradesmia Gebien, 49, 63 [F]. Type Stenocaraalbicolle Haag-Rutenberg, 1878, by original designation. Status: valid subgenus of Metriopus Solier, 1835 in Pimeliinae: Adesmiini.

Ceramba Fauvel, 206 [F]. Type Cerambahydrovatina Fauvel, 1904, by monotypy. Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Gebien (1940: 429).

Cerandria Dejean, 200 [F]. Type Trogossitacornuta Fabricius, 1798, by subsequent designation (Duponchel 1842b: 285). Status: junior synonym of Gnatocerus Thunberg, 1814 in Diaperinae: Diaperini: Adelinina. Synonymy: Erichson (1847a: 119).

Cerandrosus Gebien, 325, 394 [M]. Type Cerandrosusnodipennis Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cerasoma Endrödy-Younga, 113 [N]. Type Cerasomacerasus Endrödy-Younga, 1989, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Ceratanisus Gemminger in Gemminger and Harold, 1818 [M]. Type species [automatic]: Anisocerustristis Faldermann, 1837, by monotypy. Status: valid genus in Pimeliinae: Ceratanisini. Note: replacement name for Anisocerus Faldermann, 1837.

Ceratoma Borchmann, 49 [F]. Type none designated. Status: undetermined taxon in Lagriinae: Lagriini: Lagriina. Note: this genus was included in a key, which fulfils the criteria of availability for new genus-group names proposed before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Ceratoma and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Ceratopelius Antoine, 52, 54 [M]. Type Ceratopeliusmussardi Antoine, 1963, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Ceratupis Perty, 57 [F]. Type Ceratupisnigerrima Perty, 1830, by monotypy. Status: junior synonym of Antimachus Gistel, 1829 in Tenebrioninae: Ulomini. Synonymy: Lacordaire (1859a: 330).

Cerenopus J.L. LeConte, 143 [M]. Type Cerenopusconcolor J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 173). Status: valid genus in Tenebrioninae: Cerenopini.

Cerocamptus Gebien, 28, 151 [M]. Type Camariamalayana Fairmaire, 1893, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cerodolus Sharp, 410 [M]. Type Cerodoluschrysomeloides Sharp, 1886, by monotypy. Status: valid genus in Tenebrioninae: Heleini: incertae sedis. Note: transferred from Titaenini by Matthews and Lawrence (2019: 628).

Cerogria Borchmann, 210 [F]. Type Lagriaanisocera Wiedemann, 1823, by subsequent designation (Borchmann 1936: 119). Status: valid genus and subgenus in Lagriinae: Lagriini: Lagriina. Note: as mentioned by Löbl et al. (2020: 5), discovery of the earlier designation of Cerogriadohrni Borchmann, 1911 as the type species of Cerogria (by subsequent designation by R. Lucas (1920: 174)) threatens the nomenclatural stability of this genus since C.dohrni is the type species of the valid genus Acerogria Borchmann, 1936; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Borchmann (1936: 119).

Cerogriodes Borchmann, 25 [M]. Type Cerogriaklapperichi Borchmann, 1941, by monotypy. Status: junior synonym of Cerogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina. Synonymy: Merkl (2007: 262).

Ceromelaephus Koch, 87 [M]. Type Trachynotusbadeni Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Ceropria Laporte & Brullé, 332, 396 [F]. Type Helopsindutus Wiedemann, 1819, by subsequent designation (Duponchel 1844b: 359). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cerosis Gebien, 33, 35 [F]. Type Cerosishereroensis Gebien, 1920, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Cerostena Solier, 307, 325 [F]. Type Cerostenadeplanata Solier, 1836, by present designation. Status: junior synonym of Psectrascelis Solier, 1836 in Pimeliinae: Nycteliini. Synonymy: Fairmaire (1876: 356).

Cerostira Borchmann, 36 [F]. Type species [automatic]: Porrolagriasubaenea Borchmann, 1908, by subsequent designation (Borchmann 1936: 227). Status: valid genus in Lagriinae: Lagriina. Note: replacement name for Allocera Borchmann, 1916.

Cerysia Bremer, 68 [F]. Type Elixotalaevicornis Blair, 1929, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Cestrinus Erichson, 172 [M]. Type Cestrinustrivialis Erichson, 1842, by subsequent designation (Blair 1919a: 529). Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Champion (1894a: 354), Doyen et al. (1990: 231).

Chaerodes White, 12 [M]. Type Chaerodestrachyscelides White, 1846, by monotypy. Status: valid genus in Lagriinae: Chaerodini.

Chaeroplonyx Bremer, 36 [M]. Type Plesiophthalmuskimanisensis Masumoto, 2001, by original designation. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Chaetopsia Gebien, 567 [F]. Type Chaetopsiaangusticollis Gebien, 1925, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Chaetotoma Motschulsky, 533 [F]. Type Tenebriocephalotes Pallas, 1781, by subsequent designation (Löbl et al. 2008a: 41). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Chaetyllus Pascoe, 122 [M]. Type Chaetyllusanthicoides Pascoe, 1860, by monotypy. Status: valid genus in Lagriinae: Laenini. Note: placed in the tribe Laenini by Kanda (2016: 562).

Chalcocyclus Fairmaire, 74 [M]. Type Chalcocyclusspeculifer Fairmaire, 1884, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Chalcopauliana Ardoin, 209 [F]. Type Chalcopaulianavinsoni Ardoin, 1961, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Chalcoplonyx Ardoin, 308, 333 [M]. Type Gonocnemisviridis Kraatz, 1899, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Chalcopteroides Gebien, 497 [M]. Type species [automatic]: Chalcopterusiridicolor Blessig, 1861, by subsequent designation (Gebien 1948: 497). Status: valid genus in Tenebrioninae: Amarygmini. Note: replacement name for Chalcopterus Blessig, 1861; Chalcopteroides was proposed earlier by Strand (1935b: 302); however, this name is unavailable because Strand did not designate a type species for the nominal taxon, a mandatory requirement for replacement name without valid typification proposed after 1930 (ICZN 1999; Article 13.3.1).

Chalcopterus Blessig, 103 [M]. Type Chalcopterusiridicolor Blessig, 1861, by subsequent designation (Gebien 1948: 497). Status: senior synonym of Chalcopteroides Gebien, 1948 in Tenebrioninae: Amarygmini. Note: junior homonym of Chalcopterus Reichenbach, 1852 [Aves].

Chalcostylus Fairmaire, 409 [M]. Type Chalcostylusperrieri Fairmaire, 1898, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Nycteropina.

Charianus Bates, 297 [M]. Type Tetraphylluspurpuratus Coquerel, 1851, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Chariophenus Blair, 239 [M]. Type Chariophenuswasmanni Blair, 1929, by original designation. Status: junior synonym of Foochounus Pic, 1921 in Stenochiinae: Cnodalonini. Synonymy: Schawaller and Ando (2009: 260).

Chariotheca Pascoe, 125 [F]. Type Chariothecacoruscans Pascoe, 1860, by subsequent designation (Gebien 1942a: 335). Status: valid genus in Stenochiinae: Cnodalonini.

Chariothes Carter, 78 [M]. Type species [automatic]: Chariothecacoruscans Pascoe, 1860, by subsequent designation (Gebien 1942a: 335). Status: junior synonym of Chariotheca Pascoe, 1860 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Chariotheca Pascoe, 1860.

Charisius Champion, 421 [M]. Type Charisiusfasciatus Champion, 1888, by subsequent designation (R. Lucas 1920: 178). Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: the First Reviser (Charisius Champion, 1888 versus Narses Champion, 1888) is Campbell (2014: 271).

Chartopteryx Westwood, 43 [F]. Type Chartopteryxchildrenii Westwood, 1841, by monotypy. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy and First Reviser action (Chartopteryx Westwood, 1841 versus Cyphaleus Westwood, 1841): Matthews (1992: 490).

Chaseleodes Thomas, 122 [F]. Type Eleodescurta Champion, 1884, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Chatanayus Ardoin, 61 [M]. Type Chemolanusvillosipes Fairmaire, 1884, by original designation. Status: senior synonym of Anachayus Bouchard & Bousquet, nom. nov. in Stenochiinae: Cnodalonini. Note: junior homonym of Chatanayus Fleutiaux 1940 [Coleoptera: Elateridae].

Cheilopoma Murray, 20 [N]. Type Cheilopomacastaneum Murray, 1867, by monotypy. Status: valid genus in Diaperinae: Hypophlaeini.

Cheirodes Gené, 73 [M]. Type Cheirodessardous Gené, 1839, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Melanimonini.

Cheiroplus Ardoin, 129 [M]. Type Cheiroplusfreyi Ardoin, 1963, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: the earlier usage of Cheiroplus by Ardoin (1962b: 970) is unavailable since it was published after 1930 without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Cheirosis Deyrolle, 81, 220 [F]. Type Zophosisovata Faldermann, 1837 (= Pedinusacuminatus Fischer, 1832), by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Chemolanus Bates, 296 [M]. Type Tetraphyllusconsobrinus Fairmaire, 1868 (= Camariaobscura Klug, 1833), by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cherostus C.O. Waterhouse, 68 [M]. Type Cherostuswalkeri C.O. Waterhouse, 1894, by subsequent designation (Barber 1914: 191). Status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: Gebien (1939: 762).

Chianalus Bates, 479 [M]. Type Chianaluscostipennis Bates, 1879, by monotypy. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Chilenolagria Pic, 28 [F]. Type Chilenolagrialimbata Pic, 1936, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Chileone Bates, 264 [F]. Type Chileonedeyrollei Bates, 1868, by monotypy. Status: junior synonym of Hypaulax Bates, 1868 in Stenochiinae: Cnodalonini. Synonymy: Carter (1914b: 46).

Chiliarchum Koch, 263 [N]. Type Molurisbertolonii Guérin-Méneville, 1844, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina. Note: elevated to rank of genus (from subgenus of Ocnodes Fåhraeus, 1870) by Kamiński et al. (2020: 9, 17)

Chilometopon Horn, 31 [N]. Type Trimytisabnormis Horn, 1870, by subsequent designation (Casey 1907: 367). Status: valid genus in Pimeliinae: Edrotini.

Chinotrigon Skopin, 170 [M]. Type Trigonoscelissublaevigata Reitter, 1887, by original designation. Status: valid subgenus of Trigonoscelis Dejean, 1834 in Pimeliinae: Pimeliini.

Chirocharis Kolbe, 166, 179 [F]. Type Chiroscelisaustralis Westwood, 1845, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Chirodes Agassiz, 78, 81 [M]. Type species [automatic]: Cheirodessardous Gené, 1839, by monotypy. Status: junior synonym of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini. Note: unjustified emendation of Cheirodes Gené, 1839, not in prevailing usage.

Chiroscelis Lamarck, 261 [F]. Type Chiroscelisbifenestra Lamarck, 1804 (= Tenebriodigitatus Fabricius, 1801), by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Chirosis Gemminger in Gemminger and Harold, 1806 [F]. Type species [automatic]: Zophosisovata Faldermann, 1837 (= Pedinusacuminatus Fischer, 1832), by monotypy. Status: junior synonym of Cheirosis Deyrolle, 1867 in Pimeliinae: Zophosini. Note: unjustified emendation of Cheirosis Deyrolle, 1867, not in prevailing usage.

Chitwania Novák, 91 [F]. Type Chitwaniakejvali Novák, 2015, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Chlamydion Gistel, x [N]. Type species [automatic]: Opatrumorientale Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: junior synonym of Sclerum Dejean, 1834 in Blaptinae: Opatrini: Sclerina. Note: unnecessary replacement name for Sclerum Dejean, 1834.

Chlorocamma Bates, 371 [N]. Type Chlorocammacarenipenne Bates, 1873 (= Leptomorphasulcata Montrouzier, 1860), by subsequent designation (Gebien 1942a: 755). Status: valid genus in Stenochiinae: Cnodalonini.

Chlorophanes Matthews, 451, 461 [M]. Type Platyphanespunctipennis Carter, 1911, by original designation. Status: senior synonym of Phanechloros Matthews & Bouchard, 2008 in Tenebrioninae: Heleini: Cyphaleina. Note: junior homonym of Chlorophanes Reichenbach, 1853 [Aves].

Chlorophila Semenov, 374 [F]. Type Lagriaportschinskii Semenov, 1891, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Choaspes Champion, 118 [M]. Type Choaspespurpureus Champion, 1885, by subsequent designation (Gebien 1941: 338). Status: senior synonym of Choastes Champion, 1893 in Stenochiinae: Cnodalonini. Note: junior homonym of Choaspes Moore, 1881 [Lepidoptera].

Choastes Champion, 526 [M]. Type species [automatic]: Choaspespurpureus Champion, 1885, by subsequent designation (Gebien 1941: 338). Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Choaspes Champion, 1885.

Choerodes Gemminger in Gemminger and Harold, 1944 [M]. Type species [automatic]: Chaerodestrachyscelides White, 1846, by monotypy. Status: junior synonym of Chaerodes White, 1846 in Lagriinae: Chaerodini. Note: unjustified emendation of Chaerodes White, 1846, not in prevailing usage; junior homonym of Choerodes Leidy, 1852 [Mammalia] and Choerodes Guenée, 1857 [Lepidoptera].

Cholipus Pascoe, 471 [M]. Type Cholipusbrevicornis Pascoe, 1866, by original designation. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Mäklin (1863a: 552, with Encyalesthus Motschulsky, 1860, a junior synonym of Derosphaerus J. Thomson, 1858).

Chorasmius Bates, 310 [M]. Type Evaniosomusprocerus Erichson, 1847, by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Choresmolamus G.S. Medvedev, 150 [M]. Type Dilamusoxianus G.S. Medvedev, 1978, by original designation. Status: valid subgenus of Dilamus Jacquelin du Val, 1861 in Blaptinae: Opatrini: Ammobiina.

Choristopsis Kraatz, 81, 227 [F]. Type Choristopsiscaucasica Kraatz, 1865, by monotypy. Status: junior synonym of Calyptopsis Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Reitter (1900c: 130).

Chromatia J.L. LeConte, 244 [F]. Type Cistelaamoena Say, 1824, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Chromomaea Pascoe, 490 [F]. Type Chromomaeapicta Pascoe, 1866, by monotypy. Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Fauvel (1905: 225).

Chrysobalus Boisduval, 267 [M]. Type Chrysobalusfulgidipennis Boisduval, 1835, by monotypy. Status: senior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Lacordaire (1859b: 409); although the identity of the type species is uncertain (Matthews 1992: 439, Matthews and Bouchard 2008: 350), it is probably a species of Cyphaleus Westwood, 1841. Note: nomen oblitum; we provide references to support the conservation of Cyphaleus Westwood, 1841 as the valid name for this genus (ICZN 1999, Article 23.9.1) in Appendix 2.

Chrysolagria Seidlitz, 336, 339 [F]. Type Lagriaviridipennis Fabricius, 1798, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Chrysolinoides Jolivet, 2 [M]. Type Chrysolinoidesphilippinensis Jolivet, 1951 (= Hemicerairidicolor Gebien, 1921), by monotypy. Status: junior synonym of Hemicera Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini. Synonymy: Kulzer (1952: 764). Note: genus originally described in Chrysomeloidea: Chrysomelidae.

Chrysomaia Kulzer, 755, 756 [F]. Type Eucyrtuscarbunculus Fairmaire, 1885, by original designation. Status: junior synonym of Augolesthus Motschulsky, 1872 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 134).

Chrysopeplus Gebien, 755 [M]. Type species [automatic]: Helopsexpolitus Broun, 1880, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Leiopeplus Broun, 1893.

Cibdelis Mannerheim, 282 [F]. Type Cibdelisblaschkii Mannerheim, 1843, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cilioncotus Koch, 41 [M]. Type Oncotusnamaquanus Koch, 1954, by original designation. Status: valid subgenus of Oncotus Blanchard, 1845 in Blaptinae: Platynotini: Eurynotina.

Cillibus Matthews, 1040, 1078 [M]. Type Saragusblackburni W.J. MacLeay, 1888, by original designation. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Cimicia Fairmaire, lxxxix [F]. Type Cimiciaspinipes Fairmaire, 1891, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Cimicichora Koch, 38 [F]. Type Cimicichoracrenulata Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Cimiciopsis Koch, 21, 87 [F]. Type Cimiciopsiscastleae Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Cimipsa Peyerimhoff, 346 [F]. Type Cimipsasergenti Peyerimhoff, 1911, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Circomus Fleischer, 236 [M]. Type Hypophlaeussubdepressus Wollaston, 1864, by monotypy. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Löbl et al. (2008b: 276).

Cirsa P.H. Lucas, lvi [F]. Type Cirsastriaticollis P.H. Lucas, 1857, by monotypy. Status: valid subgenus of Micipsa P.H. Lucas, 1855 in Pimeliinae: Tentyriini.

Cirta Gemminger in Gemminger and Harold, 1831 [F]. Type species [automatic]: Cirsastriaticollis P.H. Lucas, 1857, by monotypy. Status: junior synonym of Cirsa P.H. Lucas, 1857 in Pimeliinae: Tentyriini. Note: unjustified emendation of Cirsa P.H. Lucas, 1857, not in prevailing usage.

Cissides Chatanay, 475, 495 [M]. Type Heterophyluspunctatissimus Fairmaire, 1869, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cistela Fabricius, 116 [F]. Type Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: senior synonym of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: junior homonym of Cistela Geoffroy, 1762 [Coleoptera: Byrrhidae]. Note: the older name Cistela Geoffroy, 1762 was suppressed for purposes of the Principle of Priority but not for those of the Principle of Homonymy (ICZN 1994a, Opinion 1754).

Cistelampra Fairmaire, 386 [F]. Type Cistelamprapurpurina Fairmaire, 1897, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Cistelina Seidlitz, 195 [F]. Type Cisteladavidis Fairmaire, 1878, by subsequent designation (R. Lucas 1920: 187). Status: valid genus in Alleculinae: Cteniopodini.

Cistella Gistel, xi [F]. Type species [automatic]: Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: unjustified emendation of Cistela Fabricius, 1775, not in prevailing usage.

Cistelodema Borchmann, 380 [F]. Type Pseudocistelacyanea Pic, 1930, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Cisteloida Fairmaire, 256 [F]. Type Cisteloidacastanescens Fairmaire, 1882, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Cistelomorpha Redtenbacher, 134 [F]. Type Cistelomorphastraminea Redtenbacher, 1868, by subsequent designation (Borchmann 1932a: 358). Status: valid genus in Alleculinae: Cteniopodini.

Cistelopsis Fairmaire, 39 [F]. Type Cistelopsisrufina Fairmaire, 1896, by subsequent designation (R. Lucas 1920: 187). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Clamoris Gozis, 25 [F]. Type species [automatic]: Phtoracrenata Mulsant, 1854 (= Clamorisinsurgens Gozis, 1886), by monotypy. Status: valid genus in Phrenapatinae: Penetini. Note: replacement name for Phtora Mulsant, 1854.

Clastopus Fairmaire, 407 [M]. Type Clastopuseurynotoides Fairmaire, 1898, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Claudegirardius Iwan, 372 [M]. Type Claudegirardiusbertiae Iwan, 1999, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Clavatoodescelis Kaszab, 942, 974 [F]. Type Platyscelismelas Fischer, 1823, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini. Note: the First Reviser (Clavatoodescelis Kaszab, 1940 versus Oblongoodescelis Kaszab, 1940) is Egorov (2020: 380).

Cleognathus Gebien, 154 [M]. Type Cleognathusprosternalis Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Cleolaus Champion, 142 [M]. Type Penetasommeri Lacordaire, 1859, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Cleomis Fairmaire, 54 [M]. Type Cleomisviolaceipes Fairmaire, 1892, by subsequent designation (R. Lucas 1920: 189). Status: valid genus in Stenochiinae: Cnodalonini. Note: removed from synonymy with Psydus Pascoe, 1868 by Kaszab (1983b: 382).

Clinocranion Solier, 114 [N]. Type Clinocranionspinosum Solier, 1843, by subsequent designation (R. Lucas 1920: 190). Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina. Note: the alternative original spelling Clynocranion, used by Solier (1843: 4), was rejected by Koch (1955a: 70) who acted as the First Reviser.

Clitobius Mulsant & Rey, 141 [M]. Type Clitobiussabulicola Mulsant & Rey, 1859 (= Opatrumovatum Erichson, 1843), by monotypy. Status: valid genus and subgenus in Blaptinae: Opatrini: Ammobiina. Note: the original combination of the accepted name of the type species, Opatrumovatum Erichson, 1843, is a junior primary homonym of Opatrumovatum Fabricius, 1801.

Clypeophtora F. Soldati & L. Soldati, 4 [F]. Type Phtoratronqueti F. Soldati & L. Soldati, 2003, by original designation. Status: valid subgenus of Phtora Germar, 1836 in Diaperinae: Phaleriini.

Cnecosochara Reitter, 660 [F]. Type Cnecosocharapetriiformis Reitter, 1913, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Cnemandrosus Gebien, 42 [M]. Type Eucyrtussemipurpureus Fairmaire, 1896, by original designation. Status: junior synonym of Plamius Fairmaire, 1896 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1941: 1143).

Cnemeplatia Costa, 146 [F]. Type Cnemeplatiaatropos Costa, 1847, by monotypy. Status: valid genus in Pimeliinae: Cnemeplatiini: Cnemeplatiina.

Cnemodasus Gebien, 374 [M]. Type Cnemodasusrectangulus Gebien, 1914, by subsequent designation (Kaszab 1982b: 43). Status: junior synonym of Brachyidium Fairmaire, 1883 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1921a: 492).

Cnemodinus Cockerell, 242 [M]. Type species [automatic]: Cnemodustestaceus Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Cnemodinini. Note: replacement name for Cnemodus Horn, 1870.

Cnemodus Horn, 266 [M]. Type Cnemodustestaceus Horn, 1870, by monotypy. Status: senior synonym of Cnemodinus Cockerell, 1906 in Pimeliinae: Cnemodinini. Note: junior homonym of Cnemodus Herrich-Schaeffer, 1850 [Hemiptera].

Cnemophloeus Bremer, 10, 13 [M]. Type Hypophlaeusfilum Fairmaire, 1893, by original designation. Status: junior synonym of Stenophloeus Blair, 1921 in Diaperinae: Hypophlaeini. Synonymy: Löbl et al. (2008b: 312).

Cnemoplatia Wollaston, 411 [F]. Type species [automatic]: Cnemeplatiaatropos Costa, 1847, by monotypy. Status: junior synonym of Cnemeplatia Costa, 1847 in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Note: unjustified emendation of Cnemeplatia Costa, 1847, not in prevailing usage.

Cneocnemis Gebien, 32 [F]. Type Ulomahaemorrhoum Fairmaire, 1893, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Cnephalura Doyen, 313 [F]. Type Cnephaluraumbrata Doyen, 1988, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cnodalon Latreille, 23 [N]. Type Cnodalonviride Latreille, 1804, by subsequent monotypy (Latreille 1804: 321). Status: valid genus in Stenochiinae: Cnodalonini. Note: originally proposed without included nominal species; in order to promote nomenclatural stability Bousquet and Bouchard (2019: 114, 116) recommended to the ICZN to use its plenary powers to designate Cnodalonviride Latreille, 1804 as the type species for the nominal genus Cnodalon Latreille, 1797, as currently accepted in the literature, after the recent discovery that this species was in fact not an originally included nominal species in the genus Cnodalon, and so not available as the type species.

Cnodalum Agassiz, 91 [N]. Type species [automatic]: Cnodalonviride Latreille, 1804, by subsequent monotypy (Latreille 1804: 321). Status: junior synonym of Cnodalon Latreille, 1797 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Cnodalon Latreille, 1797, not in prevailing usage; in a recent application to the ICZN to fix type species problems with the genus Cnodalon Latreille, 1797, Bousquet and Bouchard (2019: 116) recommended that Cnodalum Agassiz, 1846 be placed on the Official Index of Rejected and Invalid Generic Names in Zoology in order to avoid homonymy problems with the name Cnodalum Emeljanov, 1978 [Hemiptera], which is currently used as valid.

Coccimarygmus Ardoin, 185, 187 [M]. Type Paramarygmusmorychoides Fairmaire, 1894, by original designation. Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Cochabambia Marcuzzi, 184 [F]. Type Cochabambiakulzeri Marcuzzi, 1985, by monotypy. Status: senior synonym of Allotriocochabambia Faúndez, Rider & Carvajal, 2014 in Tenebrionidae: incertae sedis. Note: junior homonym of Cochabambia Pirán, 1959 [Hemiptera].

Coeladesmia Reitter, 3 [F]. Type Metriopusplatynotus Gerstaecker, 1854, by subsequent designation (Gebien 1937a: 656). Status: valid subgenus of Metriopus Solier, 1835 in Pimeliinae: Adesmiini.

Coelocnemis Mannerheim, 280 [F]. Type Coelocnemisdilaticollis Mannerheim, 1843, by subsequent designation (R. Lucas 1920: 194). Status: valid genus in Stenochiinae: Cnodalonini.

Coelocnemodes Bates, 474 [M]. Type Coelocnemodesstoliczkanus Bates, 1879, by monotypy. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Coeloecetes Blair, 384 [M]. Type Coeloecetescavernicola Blair, 1929, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Coelolophus Mäklin, 502 [M]. Type Coelolophusbicarinatus Mäklin, 1867, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Coelometopus Solier, 154, 278 [M]. Type Coelometopusclypeatus Solier, 1848 (= Blaps clypeata Germar, 1813), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Coelomorpha Casey, 182 [F]. Type Coelomorphamaritima Casey, 1890, by monotypy. Status: junior synonym of Coelus Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 371).

Coelopalorus Blair, 135 [M]. Type Palorusfoveicollis Blair, 1930 (= Ulominacarinata Baudi di Selve, 1876), by monotypy. Status: junior synonym of Ulomina Baudi di Selve, 1876 in Tenebrioninae: Palorini. Synonymy: Scupola (2002: 186).

Coelophanes Iablokoff-Khnzorian, 309 [M]. Type Hedyphanesimpressicollis Faldermann, 1837 (= Hedyphaneslaticollis Fischer, 1832), by original designation. Status: junior synonym of Hedyphanes Fischer, 1820 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko (2005: 351).

Coelopleurum Gebien, 35 [N]. Type Coelopleurumglabratum Gebien, 1921, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Coelosattus Blaisdell, 166 [M]. Type Coelosattusfortineri Blaisdell, 1927 (= Eusattusdilatatus J.L. LeConte, 1851), by monotypy. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 373).

Coelotaxis Horn, 200 [F]. Type Coelotaxispunctulata Horn, 1876, by subsequent designation (Gebien 1938a: 289). Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 373).

Coelus Eschscholtz, 5 [M]. Type Coelusciliatus Eschscholtz, 1829, by monotypy. Status: valid genus in Pimeliinae: Coniontini.

Colasia Koch, 127, 131 [F]. Type Colasiaakisoides Koch, 1965, by monotypy. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Collariheliofugus Freude, 125, 126 [M]. Type Euschatiacollaris Germain, 1855, by original designation. Status: valid subgenus of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini.

Colophonesthes Koch, 48 [F]. Type Colophonesthesmontisatri Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Colparthrum Kirsch, 204 [N]. Type Colparthrumgerstaeckeri Kirsch, 1866, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Colpopatrum Reitter, 148 [N]. Type Opatrumasperipenne Reitter, 1897, by monotypy. Status: valid subgenus of Opatrum Fabricius, 1775 in Blaptinae: Opatrini: Opatrina.

Colpophorinus Escalera, 336 [M]. Type Opatrumgonocephaloides Escalera, 1914, by monotypy. Status: junior synonym of Colpophorus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Opatrina. Synonymy: Español (1954b: 317).

Colpophorus Mulsant & Rey, 44 [M]. Type Opatrumemarginatum P.H. Lucas, 1847, by subsequent designation (Iwan and Löbl 2008: 269). Status: valid subgenus of Opatrum Fabricius, 1775 in Blaptinae: Opatrini: Opatrina.

Colposcelis Dejean, 185 [F]. Type Tentyrialongicollis Zubkov, 1833, by subsequent designation (Gebien 1937a: 598). Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Colposceloides Schuster, 20 [M]. Type Colposcelislicenti Schuster, 1940, by monotypy. Status: valid subgenus of Colposcelis Dejean, 1834 in Pimeliinae: Tentyriini.

Colposcythis Reitter, 113 [F]. Type Colposcythiswalteri Reitter, 1889, by monotypy. Status: valid subgenus of Colposcelis Dejean, 1834 in Pimeliinae: Tentyriini.

Colposphena Semenov, 217 [F]. Type Sphenariabrevicollis Solsky, 1882, by subsequent designation (Gebien 1937a: 577). Status: valid genus in Pimeliinae: Tentyriini.

Colpotinoides Kaszab, 282, 354 [M]. Type Pseudoblapsgebieni Kaszab, 1942, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Colpotinus Fairmaire, xvii [M]. Type Colpotinussimulator Fairmaire, 1891, by monotypy. Status: valid genus in Blaptinae: Pedinini: Pedinina.

Colpotus Mulsant & Rey, 148, 208 [M]. Type Colpotusstrigicollis Mulsant & Rey, 1853, by subsequent designation (Gebien 1938a: 313). Status: valid subgenus of Pedinus Latreille, 1797 in Blaptinae: Pedinini: Pedinina.

Comphosida Macquart, 174 [F]. Type Mesostenapunctipennis Solier, 1835 (= Pimeliaangustata Fabricius, 1775), by monotypy. Status: junior synonym of Mesostena Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Lacordaire (1859a: 52).

Compsocula Fairmaire, 236 [F]. Type Stenogenaapicata Fairmaire, 1896, by subsequent designation (R. Lucas 1920: 199). Status: valid genus in Alleculinae: incertae sedis. Note: placed in Alleculinae by Chatanay (1915a: 526).

Compsomorphus Solier, 208 [M]. Type Compsomorphuselegans Solier, 1851, by original designation. Status: junior synonym of Apocrypha Eschscholtz, 1831 in Tenebrioninae: Apocryphini. Synonymy: Lacordaire (1859b: 433).

Conibiosoma Casey, 476 [N]. Type Conibiuselongatus Horn, 1870, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Conibius J.L. LeConte, 145 [M]. Type Conibiusseriatus J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 199). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Coniontellus Casey, 388 [M]. Type Coniontisobesa J.L. LeConte, 1851, by subsequent designation (Casey 1908: 57). Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 373).

Coniontides Casey, 57, 78 [M]. Type Coniontislata J.L. LeConte, 1866, by original designation. Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 373).

Coniontis Eschscholtz, 7 [F]. Type Coniontisviatica Eschscholtz, 1829, by subsequent designation (Casey 1908: 57). Status: valid genus in Pimeliinae: Coniontini.

Conipinus J.L. LeConte, 223 [M]. Type Eusattusdubius J.L. LeConte, 1851, by subsequent designation (Gebien 1938a: 284). Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: J.L. LeConte (1866a: 60).

Conisattus Casey, 614 [M]. Type Conisattusrectus Casey, 1895, by monotypy. Status: valid genus in Pimeliinae: Coniontini.

Conoecus Horn, 159 [M]. Type Conoecusovipennis Horn, 1885, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Conophthalmus Quedenfeldt, 13 [M]. Type Conophthalmussetulosus Quedenfeldt, 1885, by monotypy. Status: valid subgenus of Amatodes Dejean, 1834 in Blaptinae: Pedinini: Helopinina. Note: see Robiche (2013: 522) for placement of the genus.

Convexoodescelis Egorov, 593 [F]. Type Platyscelisbrevipennis Kaszab, 1938, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Cophodema Gebien, 402 [F]. Type species [automatic]: Cophosomahumeridens Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Cophosoma Gebien, 1928.

Cophosoma Gebien, 219, 222 [N]. Type Cophosomahumeridens Gebien, 1928, by monotypy. Status: senior synonym of Cophodema Gebien, 1943 in Stenochiinae: Cnodalonini. Note: junior homonym of Cophosoma Costa, 1849 [Hemiptera].

Copistethus Seidlitz, 524 [M]. Type Cistelaspadix Kiesenwetter, 1861, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Coracostira Fairmaire, 220 [F]. Type Coracostiraarmipes Fairmaire, 1899, by present designation. Status: junior synonym of Xenostethus Bates, 1868 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1936: 460).

Cordibates Kulzer, 909 [M]. Type Cordibateschilensis Kulzer, 1956, by original designation. Status: valid genus in Pimeliinae: Thinobatini.

Corinta Koch, 36 [F]. Type Corintalitoralis Koch, 1950, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Coripera Pascoe, 483 [F]. Type Adeliumdeplanatum Boisduval, 1835, by original designation. Status: valid genus in Lagriinae: Adeliini.

Cornopterus Koch, 40 [M]. Type Cornopteruswykehami Koch, 1950, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Cornucistela Campbell, 133 [F]. Type Cornucistelaserrata Campbell, 1980, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Cornugeton Bremer, 224 [M]. Type Platolenesmonilicornis Gebien, 1920, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Corticeus Piller & Mitterpacher, 87 [M]. Type Corticeusunicolor Piller & Mitterpacher, 1783, by monotypy. Status: valid genus and subgenus in Diaperinae: Hypophlaeini.

Coscinoptilix Allard, 15 [M]. Type Coscinoptilixgracilicornis Allard, 1876, by monotypy. Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Champion (1887: 312). Note: Allard’s original spelling Coscinopter is corrected to Coscinoptilix in the errata for volume 14 of ‘L’Abeille, Journal d’Entomologie’ (at the end of page 36 of the section “Table alphabétique”) and therefore Coscinoptilix is considered the correct original spelling (ICZN 1999, Article 32.5.1.1); see Bousquet et al. (2018: 183).

Cosmogaster Koch, 164 [F]. Type Anchophthalmusimpressicollis Fairmaire, 1897, by original designation. Status: senior synonym of Kochogaster Kamiński & Raś, 2011 in Blaptinae: Platynotini: Platynotina. Note: junior homonym of Cosmogaster Faust, 1904 [Coleoptera: Curculionidae].

Cosmonota Blanchard, pl. 14 [F]. Type Cosmonotaangustata Blanchard, 1842, by subsequent designation (Gebien 1928: 127). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cossyphodes Westwood, 168 [M]. Type Cossyphodeswollastonii Westwood, 1851, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: Cossyphodina.

Cossyphodinus Wasmann, 163 [M]. Type Cossyphodinusindicus Wasmann, 1899, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: Paramellonina.

Cossyphodites Brauns, 91 [M]. Type Cossyphodeswoodrooffei Péringuey, 1885, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: Cossyphoditina.

Cossyphus G.-A. Olivier, 121 [M]. Type Lampyrisdepressa Fabricius, 1781, by monotypy. Status: valid genus and subgenus in Lagriinae: Cossyphini.

Costallecula Pic, 235 [F]. Type Costalleculaluteocostata Pic, 1954, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Costatosora Pic, 32 [F]. Type Nemostiradistincticornis Pic, 1911, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Costiferolagria Pic, 5 [F]. Type Lagriasemialutacea Pic, 1915, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Coxelinus Fairmaire, 202 [M]. Type Coxelinusstricticollis Fairmaire, 1869, by subsequent designation (Gebien 1941: 826). Status: valid genus in Lagriinae: Lupropini.

Craniosphena Koch, 45, 54, 143 [F]. Type Himatismusjusti Fairmaire, 1880, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Craniotus J.L. LeConte, 142 [M]. Type Craniotuspubescens J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Cratidus J.L. LeConte, 239 [M]. Type Amphidoraosculans J.L. LeConte, 1851, by monotypy. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Cratopus Eschscholtz, 5, 8 [M]. Type Cratopuscastaneus Eschscholtz, 1831, by monotypy. Status: senior synonym of Scelosodis Solier, 1835 in Pimeliinae: Tentyriini. Note: junior homonym of Cratopus Schönherr, 1826 [Coleoptera: Curculionidae].

†Cretaceites Wang, 203 [M]. Type Cretaceitesjingxiensis Wang, 1997, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Lower Cretaceous deposits (China).

Cribrasida Reitter, 11, 38 [F]. Type Asidagrandipalpis Allard, 1869, by subsequent designation (F. Soldati 2008: 32). Status: junior synonym of Elongasida Escalera, 1906 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 290).

Crististibes Koch, 62 [M]. Type Planostibesbinodosus Gebien, 1920, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Crossoscelis Gebien, 52 [F]. Type Crossoscelisclauda Gebien, 1914, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Masumoto (1999a: 123).

Cruracurvamtenebrio Robiche, 97 [M]. Type Tenebriokamgangi Robiche, 2019, by monotypy. Status: valid subgenus of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini.

Cryphaeus Klug, 89 [M]. Type Cryphaeusaries Klug, 1833, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Toxicina.

Crypsinous Fairmaire, 262 [M]. Type Crypsinousacutispina Fairmaire, 1891, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Crypsis C.O. Waterhouse, 73 [F]. Type Crypsisviolaceipennis C.O. Waterhouse, 1877, by monotypy. Status: valid genus in Diaperinae: Leiochrinini.

Cryptadius Fairmaire, 395 [M]. Type species [automatic]: Crypsinousacutispina Fairmaire, 1891, by monotypy. Status: junior synonym of Crypsinous Fairmaire, 1891 in Tenebrioninae: Amarygmini. Note: unnecessary replacement name for Crypsinous Fairmaire, 1891 (as “Crypsinon”); junior homonym of Cryptadius J.L. LeConte, 1851 [Coleoptera: Tenebrionidae: Pimeliinae: Edrotini].

Cryptadius J.L. LeConte, 140 [M]. Type Cryptadiusinflatus J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Cryptasida Koch, 129 [F]. Type Asidanamaqua Péringuey, 1899, by original designation. Status: valid genus in Pimeliinae: Asidini.

Crypticanus Fairmaire, 119 [M]. Type Crypticanuscuneatus Fairmaire, 1897 (= Melanopterusedwarsii Mulsant & Rey, 1854), by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Crypticocatops Kaszab, 102 [M]. Type Platydemacatopoides Fairmaire, 1896, by original designation. Status: valid subgenus of Microcrypticus Gebien, 1921 in Diaperinae: Crypticini.

Crypticoides Fairmaire, 389 [M]. Type Crypticoidesmellyi Fairmaire, 1898, by monotypy. Status: junior synonym of Oxycara Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Koch (1960: 382).

Crypticomorpha Casey, 81, 140 [F]. Type Coniontistenuis Casey, 1908, by monotypy. Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Aalbu et al. (2002: 487).

Crypticopsis Antoine, 270 [F]. Type Crypticuscorticeus Fairmaire, 1871, by original designation. Status: valid subgenus of Crypticus Latreille, 1816 in Diaperinae: Crypticini.

Crypticus Latreille, 298 [M]. Type Helopsglaber Fabricius, 1775 (= Tenebrioquisquilius Linnaeus, 1761), by monotypy. Status: valid genus and subgenus in Diaperinae: Crypticini.

Cryptobates Fairmaire, 231 [M]. Type Cryptobatesrubigineus Fairmaire, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cryptobatoides Kaszab, 2, 15 [F]. Type Cryptobatoidesopaca Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cryptobrachys Kaszab, 4, 14 [M]. Type Cryptobatescrassecostatus Fairmaire, 1898, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cryptocarpes Koch, 191 [M]. Type Cryptocarpeselongatus Koch, 1952, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini. Note: the First Reviser (Cryptocarpes Koch, 1952 versus Lornamus Koch, 1952) is Endrödy-Younga (1996: 21).

Cryptochile Latreille, 576 [F]. Type Pimeliamaculata Fabricius, 1781, by monotypy. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina. Note: discovery of the older unused name Phymatium Billberg, 1820 threatens the validity of Cryptochile Latreille, 1828 and its associated family-group names currently used as valid; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to conserve usage of Cryptochile Latreille, 1828.

Cryptogenius Solier, 37, 122 [M]. Type Cryptogeniusdentatus Solier, 1843, by original designation. Status: senior synonym of Phrynocolus Lacordaire, 1859 in Pimeliinae: Sepidiini: Molurina. Note: junior homonym of Cryptogenius Westwood, 1842 [Coleoptera: Hybosoridae].

Cryptoglossa Solier, 638, 680 [F]. Type Cryptoglossabicostata Solier, 1837, by monotypy. Status: valid genus in Pimeliinae: Cryptoglossini.

†Cryptohelops Nabozhenko & Kirejtshuk, 68 [M]. Type Cryptohelopsmenaticus Nabozhenko & Kirejtshuk, 2014, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina. Note: described from Middle-Upper Paleocene deposits (France).

Cryptomysia Pic, 260 [F]. Type Cryptomysiaminor Pic, 1954, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Cryptops Solier, 235 [M]. Type Cryptopsulomoides Solier, 1851 (= Tenebriodiaperinus Panzer, 1797), by original designation. Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Gemminger in Gemminger and Harold (1870: 1962). Note: junior homonym of Cryptops Leach, 1814 [Chilopoda].

Cryptostenophanes Kaszab, 5, 12 [M]. Type Cryptostenophanesborneensis Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cryptotrophus Gistel, xi [M]. Type species [automatic]: Pimeliamaculata Fabricius, 1781, by monotypy. Status: junior synonym of Cryptochile Latreille, 1828 in Pimeliinae: Cryptochilini: Cryptochilina. Note: unnecessary replacement name for Cryptochile Latreille, 1828.

Cryptozoon Schaufuss, 47 [N]. Type Cryptozooncivile Schaufuss, 1882, by subsequent designation (Bousquet et al. 2018: 284). Status: valid genus in Diaperinae: Gnathidiini: Anopidiina. Note: transferred from Colydiidae (see Doyen and Lawrence 1979: 366).

Csikiola Kaszab, 493 [F]. Type Csikiolathesileiformis Kaszab, 1955, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Csiro G.S. Medvedev & Lawrence, 563 [F]. Type Hyocissubparallelus Champion, 1894, by original designation. Status: valid genus and subgenus in Diaperinae: Hyociini: Hyociina.

Cteisa Solier, 242 [F]. Type Cteisahirta Solier, 1835, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Cteisodella Novák, 50 [F]. Type Cteisodellaassamica Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Cteisodes Borchmann, 307 [F]. Type Cteisodessericea Borchmann, 1932, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Cteniopinus Seidlitz, 200 [M]. Type Cistelaaltaica Gebler, 1830, by subsequent designation (R. Lucas 1920: 212). Status: valid genus and subgenus in Alleculinae: Cteniopodini.

Ctenioposomus Reitter, 131 [M]. Type Cteniopusfrater Reitter, 1903, by subsequent designation (Novák and Pettersson 2008: 330). Status: valid subgenus of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: the alternative original spelling Ctenoposomus, used by Reitter (1906b: 135), was rejected by Bousquet et al. (2015: 140) who acted as the First Revisers.

Cteniopus Solier, 245, 246 [M]. Type Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Westwood 1838: 32). Status: valid genus and subgenus in Alleculinae: Cteniopodini.

Ctenogria Borchmann, 48, 101 [F]. Type Ctenogriavermiculata Borchmann, 1916, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Ctesicles Champion, 7 [M]. Type Ctesiclesinsularis Champion, 1896, by subsequent designation (R. Lucas 1920: 214). Status: junior synonym of Diastolinus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Blapstinina. Synonymy: Ivie and Hart (2016: 468).

Ctimene Bates, 359 [F]. Type Ctimenebreweri Bates, 1873, by monotypy. Status: senior synonym of Mitrothorax Carter, 1914 in Tenebrioninae: Heleini: Cyphaleina. Note: junior homonym of Ctimene Boisduval, 1832 [Lepidoptera].

Cuemus Bouchard, 96 [M]. Type Cuemusmonteithi Bouchard, 2000, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cuphotes Champion, 332 [F]. Type species [automatic]: Spheniscuserotyloides W. Kirby, 1819, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: replacement name for Spheniscus W. Kirby, 1819.

Curimosphena Gebien, 42, 43 [F]. Type Himatismusvillosus Haag-Rutenberg, 1870, by original designation. Status: junior synonym of Imatismus Dejean, 1834 in Pimeliinae: Tentyriini. Synonymy: Bouchard et al. (2007: 386).

Curtolyprops Pic, 11 [M]. Type Curtolypropslatipennis Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Curtopeltoides Pic, 14 [M]. Type Curtopeltoidesrufescens Pic, 1916, by subsequent designation (Gebien 1940: 1062). Status: valid genus in Tenebrioninae: Ulomini.

Cybopiestes Reitter, 148, 150 [M]. Type Cybopiestescsikii Reitter, 1917, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cybostira Borchmann, 429, 430 [F]. Type Cybostiracaligata Borchmann, 1936, by original designation. Status: valid subgenus of Epicydes Champion, 1889 in Lagriinae: Lagriini: Statirina.

Cybotus Casey, 482 [M]. Type Blapstinusestriatus J.L. LeConte, 1878, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Cychrachna Koch, 341, 361 [F]. Type Cychrachnacarcharoides Koch, 1950, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Cychrochile Koch, 160 [F]. Type Cychrochileerodioides Koch, 1953, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Cyclobiomorphus Pic, 1 [M]. Type Cyclobiomorphusundulatus Pic, 1916, by subsequent designation (Gebien 1940: 420). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cyclobium Pic, 11 [N]. Type Cyclobiumvesiculiferum Pic, 1916, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cyclocnera Leo, 50 [F]. Type Cyclocneraazarovi Leo, 2018, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Cyclonesus Fairmaire, 104 [M]. Type Cyclonesusparvicollis Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cyclophanes Carter, 92 [M]. Type Cyclophanesvariegatus Carter, 1913, by subsequent designation (R. Lucas 1920: 217). Status: junior synonym of Hemicyclus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 497).

Cyclosattus Casey, 710 [M]. Type Eusattuswebsteri Casey, 1891 (= Celibecostata Solier, 1848), by monotypy. Status: junior synonym of Saragus Erichson, 1842 in Tenebrioninae: Heleini: Heleina. Synonymy: Doyen et al. (1990: 244). Note: the type species was originally described from Colorado [United States of America] in error, Doyen et al. (1990: 244) pointed out that Eusattuswebsteri Casey, 1891 is a synonym of the Australian species Celibecostata Solier, 1848.

Cylindrinotus Faldermann, 73 [M]. Type Cylindrinotuslugubris Faldermann, 1837 (= Helopsfemoratus Faldermann, 1837), by subsequent designation (Gebien 1943: 425). Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina. Note: Gebien (1943: 425) selected Helopsfemoratus Faldermann, 1837 as type species, which is not an originally included species in Cylindrinotus; however, since Gebien listed Cylindrinotuslugubris Faldermann, 1837 (a species originally included in Cylindrinotus) as a synonym of Helopsfemoratus Faldermann, 1837, he is deemed to have designated the latter as type species (ICZN 1999, Article 69.2.2); the original combination of the accepted name of the type species, Helopsfemoratus Faldermann, 1837, is a junior primary homonym of Helopsfemoratus Fabricius, 1798.

Cylindronotus Agassiz, 111 [M]. Type species [automatic]: Cylindrinotuslugubris Faldermann, 1837 (= Helopsfemoratus Faldermann, 1837), by subsequent designation (Gebien 1943: 425). Status: junior synonym of Cylindrinotus Faldermann, 1837 in Tenebrioninae: Helopini: Cylindrinotina. Note: unjustified emendation of Cylindrinotus Faldermann, 1837, not in prevailing usage.

Cylindrosia Gebien, 289 [F]. Type Cylindrosiafoveifrons Gebien, 1922, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Cylindrosora Borchmann, 237, 375 [F]. Type Soramacer Borchmann, 1930, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Cylindrostira Borchmann, 237, 374 [F]. Type Casnonideacorporaali Borchmann, 1929, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Cylindrothorus Solier, 4, 50 [M]. Type Cylindrothoruspilosus Solier, 1843, by original designation. Status: valid genus and subgenus in Alleculinae: Alleculini: Mycetocharina.

Cymatodes Agassiz, 112 [M]. Type species [automatic]: Helopsundatus Fabricius, 1792 (= Erotylusnebulosus Fabricius, 1781), by monotypy. Status: junior synonym of Cymatothes Dejean, 1834 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Cymatothes Dejean, 1834, not in prevailing usage; the older name Cymatodes W. Kirby & Spence, 1828 [Coleoptera: Curculionidae] is not nomenclaturally available.

Cymatothes Dejean, 208 [M]. Type Helopsundatus Fabricius, 1792 (= Erotylusnebulosus Fabricius, 1781), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Cymbeba Pascoe, 483 [F]. Type Cymbebadissimilis Pascoe, 1866, by monotypy. Status: valid genus in Lagriinae: Adeliini. Note: as mentioned by Matthews and Bouchard (2008: 350) the type species was erroneously described from Australia; the genus Cymbeba Pascoe, 1866 is endemic to New Caledonia.

Cynaeus J.L. LeConte, 233 [M]. Type Platydemaangusta J.L. LeConte, 1851, by original designation. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Cyphaleus Westwood, 43 [M]. Type Helopsrugosus Gray, 1831, by subsequent designation (Duponchel 1844a: 547). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: nomen protectum (see Appendix 2); Cyphaleus is sometimes attributed to Hope (1841: 126) in the literature (e.g., Matthews 631); however, Hope did not describe the genus and there is no indication in his publication that “Cyphaleusrugosus Hope”; the only species he included in the genus is the same as Helopsrugosus Gray, 1831 (“Cyphaleus Hope” was treated as an unavailable name by Neave (1939: 939)); the First Reviser (Cyphaleus Westwood, 1841 versus Chartopteryx Westwood, 1841) is Matthews (1992: 490).

Cyphelops Fairmaire, 73 [M]. Type Cyphelopsinflatus Fairmaire, 1901, by monotypy. Status: junior synonym of Miotodera Fairmaire, 1901 in Stenochiinae: Stenochiini. Synonymy: Chatanay (1915a: 526).

Cyphogenia Solier, 638, 677 [F]. Type Tenebrioauritus Pallas, 1781, by monotypy. Status: valid genus and subgenus in Pimeliinae: Akidini.

Cyphonotus Guérin-Méneville, pl. 5 [M]. Type Cyphonotusdromedarius Guérin-Méneville, 1831, by monotypy. Status: senior synonym of Homocyrtus Dejean, 1834 in Tenebrionidae: incertae sedis. Note: junior homonym of Cyphonotus Fischer, 1823 [Coleoptera: Scarabaeidae].

Cyphostethe Marseul, xxxix [F]. Type Himatismusferrugineus Marseul, 1866, by subsequent designation (Gebien 1937a: 573). Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Cyphostethoides Löbl & Merkl in Löbl et al., 2020: 3 [M]. Type Cyphostethebrunnea Kaszab, 1962, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini. Note: name first proposed by Kaszab (1979c: 273) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 244) designated Cyphostethebrunnea Kaszab, 1962 as the type species of Kaszab’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Cyptus Gerstaecker, 61 [M]. Type Cyptusscabrosus Gerstaecker, 1871, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Cyriogeton Pascoe, 356 [M]. Type Cyriogetoninsignis Pascoe, 1871, by monotypy. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Cyrta Lacordaire, 724 [F]. Type species [automatic]: Cirsastriaticollis P.H. Lucas, 1857, by monotypy. Status: junior synonym of Cirsa P.H. Lucas, 1857 in Pimeliinae: Tentyriini. Note: unjustified emendation of Cirsa P.H. Lucas, 1857, not in prevailing usage.

Cyrtoderes Dejean, 181 [M]. Type Sepidium lacunosum Thunberg, 1784 (= Tenebriocristatus DeGeer, 1778), by subsequent designation (Bousquet and Bouchard 2013a: 45). Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Cyrtomius Casey, 379 [M]. Type Cyrtomiuscavicauda Casey, 1907 (= Epitragusplicatus Champion, 1884), by original designation. Status: valid genus and subgenus in Pimeliinae: Epitragini.

Cyrtosoma Perty, 59 [N]. Type Cyrtosomaunicolor Perty, 1830, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cyrtostrongylium Blair, 246 [N]. Type Cyrtostrongyliumrhysopaussoides Blair, 1929 (= Macrosynopticuscostatus Pic, 1922), by monotypy. Status: junior synonym of Macrosynopticus Pic, 1922 in Tenebrioninae: Amarygmini. Synonymy: Pic (1930d: 12).

Cyrtotyche Pascoe, 469 [F]. Type Cyrtotychesatanas Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cyrtotyctus Kolbe, 241 [M]. Type Cyrtotyctusosdaroides Kolbe, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Dactylocalcar Gebien, 46 [N]. Type Dactylocalcarcaecus Gebien, 1938, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Daedrosis Bates, 266 [F]. Type Daedrosiscrenatostriata Bates, 1868, by subsequent designation (R. Lucas 1920: 224). Status: valid genus in Lagriinae: Adeliini.

Dailognatha Steven, 88 [F]. Type Tentyriaaequalis Tauscher, 1812, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Dalmanius Bremer, 90, 95 [M]. Type Dalmaniusperegrinus Bremer, 2001, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Damatris Laporte, 224 [F]. Type Tetraphyllusformosus Laporte & Brullé, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Danielomira Weise, 119 [F]. Type Isomiracantabrica Weise, 1974, by original designation. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Danodema Gebien, 570 [F]. Type Danodemasubcalva Gebien, 1925, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Daochus Champion, 139 [M]. Type Daochusmandibularis Champion, 1886, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Dasus Motschulsky, 78 [M]. Type Opatrumfuscum Herbst, 1793 (= Opatrumrusticum G.-A. Olivier, 1812), by original designation. Status: junior synonym of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Bedel (1894: 154).

Dasyplonyx Bremer, 37 [M]. Type Cyriogetonmaculosus Pic, 1922, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Dasytoxystropus Pic, 12 [M]. Type Dasytoxystropussubparallelus Pic, 1921, by original designation. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Dauresia Ferrer, 187 [F]. Type Dauresiamontisusti Ferrer, 2001, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Davaona Borchmann, 442, 524 [F]. Type Casnonideaperforata Borchmann, 1913, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Debeauxiella Bouchard & Bousquet, new subgenus [F]. Type Pachyceraangulophthalma Koch, 1943, by present designation. Status: valid subgenus of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: Koch (1943a: 524, 546) introduced the new subgenus name Debeauxiella for three nominal species, but unfortunately did not designate a type species; the subgenusDebeauxiella, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Pachyceraangulophthalma Koch, 1943 as type species and referring to Koch (1943a: 524) for the character states that characterise and differentiate Debeauxiella.

Dechius Pascoe, 455 [M]. Type Dechiusaphodioides Pascoe, 1866 (= Tenebriocancellatus Montrouzier, 1860), by monotypy. Status: junior synonym of Scotoderus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Bates (1873f: 473).

Dechiustes Blair, 137 [M]. Type Dechiustescarolinensis Blair, 1940, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Decialma Pascoe, 288, 291 [F]. Type Decialmatenuitarsis Pascoe, 1869, by monotypy. Status: junior synonym of Olisthaena Erichson, 1842 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1913a: 62).

Decoriplus Louw, 117, 120 [M]. Type Psammodespictus Haag-Rutenberg, 1871, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Delognatha Agassiz, 116, 118 [F]. Type species [automatic]: Tentyriaaequalis Tauscher, 1812, by monotypy. Status: junior synonym of Dailognatha Steven, 1828 in Pimeliinae: Tentyriini. Note: unjustified emendation of Dailognatha Steven, 1828, not in prevailing usage; suppressed for the purposes of both the Principle of Priority and the Principle of Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology by the ICZN (2010b, Opinion 2250).

Delognatha Lacordaire, 315 [F]. Type Delognathalacordairei Lacordaire, 1859, by subsequent designation (Gebien 1940: 756). Status: valid genus in Phrenapatinae: Phrenapatini. Note: the senior homonym Delognatha Agassiz, 1846, an unjustified emendation of Dailognatha Steven, 1828 [Coleoptera: Tenebrionidae: Pimeliinae: Tentyriini], was placed on the Official Index of Rejected and Invalid Generic Names in Zoology and suppressed for the purposes of both the Principle of Priority and the Principle of Homonymy by the ICZN (2010b, Opinion 2250).

Delonurops Reitter, 24, 25 [M]. Type Entomogonusclavimanus Reitter, 1903, by subsequent designation (Iablokoff-Khnzorian 1964: 304). Status: valid subgenus of Entomogonus Solier, 1848 in Tenebrioninae: Helopini: Helopina.

Delopygus J.L. LeConte, 129 [M]. Type Delopyguscrenatus J.L. LeConte, 1866, by monotypy. Status: junior synonym of Eutochia J.L. LeConte, 1862 in Tenebrioninae: Ulomini. Synonymy: Horn (1870: 372).

Dema Gistel, xi [F]. Type species [automatic]: Opatrumclathratum Fabricius, 1787, by monotypy. Status: junior synonym of Opatrinus Dejean, 1821 in Blaptinae: Platynotini: Platynotina. Note: unnecessary replacement name for OpatrinusDejean 1821.

Demtrius Broun, 243 [M]. Type Demtriuscarinulatus Broun, 1895, by monotypy. Status: valid genus in Tenebrioninae: Titaenini. Note: placed in Titaenini by Matthews (2003b: 13).

Dendarophylan Español, 312, 313 [M]. Type Phylan pardoi Español, 1945, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Dendaroscelis Reitter, 79 [F]. Type Dendarusserripes Reitter, 1904, by monotypy. Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Dendarus Dejean, 65 [M]. Type Helopstristis Rossi, 1790, by subsequent designation (Blanchard 1844: pl. 48). Status: valid genus and subgenus in Blaptinae: Dendarini: Dendarina. Note: as pointed out by Silfverberg (1984: 59) and Löbl et al. (2008a: 41) the Helopstristis Herbst of Laporte (1840) (= Pandaruscoarcticollis Mulsant, 1854) was used as the type species of Dendarus (e.g., Gebien 1938a: 299) because Helopstristis Rossi, 1790 was treated as a nomen dubium; however, the Helopstristis Herbst of Laporte (1840), in addition to being nomenclaturally unavailable, was not originally included in Dendarus Dejean, 1821 and therefore cannot be used as the type species; the earliest, and valid, type species designation is that of Blanchard (1844: pl. 48, as “Pedinus (Dendarus) tristis Rossi”).

Dendroscopius Gistel, 125 [M]. Type species [automatic]: Trogossitathoracica Fabricius, 1792, by monotypy. Status: junior synonym of Bius Dejean, 1834 in Tenebrioninae: Tenebrionini. Note: unnecessary replacement name for Bius Dejean, 1834.

Dengitha Reitter, 516 [F]. Type Dengithalutea Reitter, 1887, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Dentatoploedipus Kaszab, 355, 383 [M]. Type Dentatoploedipussembilanicus Kaszab, 1984, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Dentibirus G.S. Medvedev, 170, 204 [M]. Type Heliopatespusillus Ménétriés, 1849, by original designation. Status: junior synonym of Cabirutus Strand, 1929 in Blaptinae: Pedinini: Pedinina. Synonymy: Kamiński and Iwan (2017: 595).

Deplanchesia Fauvel, 310 [F]. Type Deplanchesiametallescens Fauvel, 1860, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Deretus Gahan, 10 [M]. Type Deretusdenticollis Gahan, 1900, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Deriles Motschulsky, 24 [M]. Type Upisexcavata Herbst, 1797, by original designation. Status: valid subgenus of Amenophis J. Thomson, 1858 in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Derilis, used by Motschulsky (1872: 27).

Derispia Lewis, 389 [F]. Type Diaperismaculipennis Marseul, 1876, by original designation. Status: valid genus in Diaperinae: Leiochrinini.

Derispiella Kaszab, 357, 364 [F]. Type Derispiellahingstoni Kaszab, 1961, by original designation. Status: valid genus in Diaperinae: Leiochrinini.

Derispiola Kaszab, 30, 115 [F]. Type Derispiolafruhstorferi Kaszab, 1946, by original designation. Status: valid genus in Diaperinae: Leiochrinini. Note: Derispiola was used earlier by Gebien (1940: 434) but is unavailable from that date because it was published without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1).

Derispiolina Kaszab, 279 [F]. Type Derispiolinapterolomoides Kaszab, 1979, by original designation. Status: junior synonym of Falsotithassa Pic, 1934 in Lagriinae: Lupropini. Synonymy: Schawaller (2000: 2).

Derolagria Borchmann, 61 [F]. Type Lagriaplicatula Borchmann, 1909, by subsequent designation (Borchmann 1936: 151). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Deroplatus Solier, 133 [M]. Type Geoboruscostatus Blanchard, 1842, by subsequent designation (Gebien 1937a: 563). Status: junior synonym of Geoborus Blanchard, 1842 in Pimeliinae: Epitragini. Synonymy: Lacordaire (1859a: 77).

Derosalax Gebien, 85 [M]. Type Derosalaxbruchi Gebien, 1926, by monotypy. Status: valid genus in Pimeliinae: Trilobocarini.

Derosimus Fairmaire, 62 [M]. Type Derosimusquadricollis Fairmaire, 1904, by monotypy. Status: junior synonym of Rhypasma Pascoe, 1862 in Lagriinae: Belopini. Synonymy: Blair (1935a: 104).

Derosphaerius Westwood, 362 [M]. Type Derosphaeriusanthracinus Westwood, 1881, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Derosphaerus J. Thomson, 99 [M]. Type Derosphaerusglobicollis J. Thomson, 1858, by subsequent designation (R. Lucas 1920: 233). Status: valid genus in Stenochiinae: Cnodalonini. Note: nomen protectum (see Bouchard and Bousquet 2020b: 6).

Derostethe Koch, 285 [F]. Type Himatismuspunctatissimus Haag-Rutenberg, 1870, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini.

Derostira Fairmaire, 388 [F]. Type Derostiracrenulipennis Fairmaire, 1897, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Derostirostaius Borchmann, 477 [M]. Type Derostiralaticollis Pic, 1913, by original designation. Status: valid subgenus of Staius Fairmaire, 1896 in Lagriinae: Lagriini: Statirina. Note: first proposed by Pic (1934: 32) without original type species designation; it is therefore unavailable from that date (ICZN 1999, Article 13.3).

Derostrophus Fairmaire, 186 [M]. Type species [automatic]: Derosphaeriusanthracinus Westwood, 1881, by monotypy. Status: junior synonym of Derosphaerius Westwood, 1881 in Pimeliinae: Tentyriini. Note: unnecessary replacement name for Derosphaerius Westwood, 1881.

Desertosochrus Koch, 149 [M]. Type Drosochruspiligaster Koch, 1958, by original designation. Status: valid subgenus of Drosochrus Erichson, 1843 in Blaptinae: Pedinini: Helopinina.

Diabolicobates Pic, 30 [M]. Type Diabolicobatescornutus Pic, 1930 (= Mechanetesplatitubera Kaszab, 1941), by monotypy. Status: junior synonym of Mechanetes C.O. Waterhouse, 1887 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1941a: 43).

Diachoriops Ando, 8 [M]. Type species [automatic]: Schizommacucumericola Gebien, 1921, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Schizomma Gebien, 1921.

Diacis Koch, 105 [M]. Type Trachynotusregalis Haag-Rutenberg, 1876, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Diaclina Jacquelin du Val, 296 [F]. Type Tenebriochrysomelinus Herbst, 1799 (= Tenebriotestudineus Piller & Mitterpacher, 1783), by original designation. Status: valid genus in Tenebrioninae: Alphitobiini.

Diaderma Koch, 399, 405 [N]. Type Diadermapunticum Koch, 1960, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Diaperis Geoffroy, 337 [F]. Type Chrysomelaboleti Linnaeus, 1758, by subsequent monotypy (Müller 74; see ICZN 1994a, Opinion 1754). Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: originally proposed without included nominal species; placed on the Official List of Generic names in Zoology by the ICZN (1994a, Opinion 1754).

Diaphanidus Reitter, 299, 301 [M]. Type Arthrodeisantennatus Reitter, 1894, by subsequent designation (R. Lucas 1920: 235). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Diaspirus Matthews, 704, 759 [M]. Type Diaspirusbellendenus Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Diastanus Fairmaire, 338 [M]. Type Diastanusnitidiventris Fairmaire, 1902, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Diastixus Allard, 5 [M]. Type Helopspunctipennis P.H. Lucas, 1846 (= Helopsheteromorphus P.H. Lucas, 1846), by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Nesotes Allard, 1876 in Tenebrioninae: Helopini: Helopina. Synonymy: Antoine (1949: 134).

Diastoleus Solier, 8, 67 [M]. Type Scotobiuscollaris Guérin-Méneville, 1834, by original designation. Status: valid genus in Tenebrioninae: Scotobiini.

Diastolinus Mulsant & Rey, 70, 74 [M]. Type Blapsclathrata Fabricius, 1792, by subsequent designation (R. Lucas 1920: 236). Status: valid genus in Blaptinae: Opatrini: Blapstinina. Note: the First Revisers (Diastolinus Mulsant & Rey, 1859 versus Sellio Mulsant & Rey, 1859) are Ivie and Hart (2016: 468).

Diceroderes Solier, 30, 46 [M]. Type Diceroderesmexicanus Solier, 1841, by original designation. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Dichastops Gerstaecker, 63 [M]. Type Dichastopssubaeneus Gerstaecker, 1871, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Dichillesthes Reitter, 156 [F]. Type Dichilluscordicollis Reitter, 1886, by monotypy. Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillinus Reitter, 156, 161 [M]. Type Tageniapusilla Ménétriés, 1849, by subsequent designation (G.S. Medvedev 1975: 599). Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillocerus Reitter, 155, 167 [M]. Type Hyperopspertusus Kiesenwetter, 1861, by subsequent designation (G.S. Medvedev 1975: 603). Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillodontus Reitter, 156, 166 [M]. Type Dichillusadriani Reitter, 1916, by monotypy. Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillomessor Reitter, 155 [M]. Type Dichillushimalayanus Fairmaire, 1896, by subsequent designation (G.S. Medvedev 1975: 594). Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillus Jacquelin du Val, 253 [M]. Type Tageniaminuta Solier, 1838, by original designation. Status: valid genus and subgenus in Pimeliinae: Stenosini: Dichillina.

Dichomma Solier, 253, 271 [N]. Type Dichommamaillei Solier, 1835 (= Tentyriadardana Steven, 1828), by subsequent designation (Jacquelin du Val 249). Status: valid genus in Pimeliinae: Tentyriini.

Dichotymus Fairmaire, ccxcv [M]. Type Dichotymusstriatipennis Fairmaire, 1891, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Dichromma Mulsant & Rey, 94 [N]. Type Pandarinusforaminosus Mulsant & Rey, 1855, by monotypy. Status: junior synonym of Paroderus Mulsant & Rey, 1854 in Blaptinae: Dendarini: Dendarina. Synonymy: Reitter (1904: 89, through placement of the type species in Paroderus Mulsant & Rey, 1854).

Dichtha Haag-Rutenberg, 39 [F]. Type Cryptogeniusinflatus Gerstaecker, 1854, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Dicraeosis Gebien, 355 [F]. Type species [automatic]: Dicraeusbacillus Marseul, 1876, by monotypy. Status: junior synonym of Stenochinus Motschulsky, 1860 in Stenochiinae: Cnodalonini. Note: replacement name for Dicraeus Marseul, 1876.

Dicraeus Marseul, 103 [M]. Type Dicraeusbacillus Marseul, 1876, by monotypy. Status: junior synonym of Stenochinus Motschulsky, 1860 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1925a: 79). Note: junior homonym of Dicraeus Loew, 1873 [Diptera].

Dictysomorphus Pic, 24 [M]. Type Dictysomorphusdentaticornis Pic, 1921, by monotypy. Status: junior synonym of Ceropria Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Synonymy: Kaszab (1983a: 132).

Dictysus Rye, 281 [M]. Type species [automatic]: Dietysusconfusus Pascoe, 1866 (= Cnodalonmetallicum Perty, 1831), by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Note: unjustified emendation for Dietysus Pascoe, 1866, not in prevailing usage.

Dicyrtodes Matthews, 706, 740 [M]. Type Dicyrtodesarneius Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Dicyrtus Duponchel, 5 [M]. Type Dicyrtusgibbosus Duponchel, 1844, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Diemenoma Matthews, 707, 710 [F]. Type Adeliumcommodum Pascoe, 1869, by original designation. Status: valid genus in Lagriinae: Adeliini.

Diesia Fischer, pl. 14 [F]. Type Diesiaquadridentata Fischer, 1820, by subsequent designation (Hope 1841: 118). Status: valid genus in Pimeliinae: Pimeliini.

Diesiola Skopin, 397, 398 [F]. Type Diesiaquadridentata Fischer, 1820, by original designation. Status: junior synonym of Diesia Fischer, 1820 in Pimeliinae: Pimeliini. Synonymy: Skopin (1971: 328).

Diestecopus Solier, 152, 194 [M]. Type Diestecopuserodioides Solier, 1848, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Diestesoma Péringuey, 271 [N]. Type Diestesomapulchrum Péringuey, 1904, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Diestica Pascoe, xii [F]. Type Diesticaviridipennis Pascoe, 1868, by monotypy. Status: junior synonym of Poecilesthus Dejean, 1834 in Stenochiinae: Stenochiini. Synonymy: Gebien (1911b: 589).

Dietomorpha Reymond, 143 [F]. Type Dietomorphapardalis Reymond, 1938, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Dietopsis Solier, 236 [F]. Type Alleculasolieri Laporte, 1840, by subsequent monotypy (Laporte 1840: 243). Status: valid subgenus of Allecula Fabricius, 1801 in Alleculinae: Alleculini: Alleculina. Note: originally proposed without included nominal species; Laporte (1840: 243), by describing one new species Alleculasolieri Laporte, 1840 in association with the subgenus “Dietopsis Sol.”, was the first author to subsequently and expressly include nominal species in Dietopsis (ICZN 1999, Article 67.2.2).

Dietysus Pascoe, 486 [M]. Type Dietysusconfusus Pascoe, 1866 (= Cnodalonmetallicum Perty, 1831), by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1921a: 409).

Dignamptus J.L. LeConte, 421 [M]. Type Dignamptusstenochinus J.L. LeConte, 1878, by subsequent designation (Bousquet et al. 2018: 325). Status: junior synonym of Talanus Jacquelin du Val, 1857 in Stenochiinae: Talanini. Synonymy: Champion (1887: 321).

Dignathosis Koch, 75 [F]. Type Dignathosisarcana Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Dila Fischer von Waldheim, 111 [F]. Type Blapslaevicollis Gebler, 1841, by subsequent designation (Motschulsky 1860c: 530). Status: valid genus in Blaptinae: Blaptini: Blaptina.

Dilablaps Bogatchev, 98 [F]. Type Dilablapsparadoxa Bogatchev, 1976, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Dilamus Jacquelin du Val, 279 [M]. Type Borosrufipes P.H. Lucas, 1846, by original designation. Status: valid genus and subgenus in Blaptinae: Opatrini: Ammobiina.

Dillacerus Solier, 497 [M]. Type Dillacerusduponti Solier, 1835 (= Dolichoderusacuminatus Klug, 1833), by monotypy. Status: senior synonym of Macellocerus Solier, 1848 in Tenebrioninae: Toxicini: Nycteropina. Note: discovery of this forgotten name threatens the stability of the junior synonym Macellocerus Solier, 1848; although Dillacerus Solier, 1835 has not been used as valid in the literature after 1899, we could not find usage of Macellocerus Solier, 1848 in at least 25 works, published by at least ten authors in the immediately preceding 50 years, and therefore reversal of precedence cannot be used to treat Dillacerus Solier, 1835 as a nomen oblitum; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to conserve usage of Macellocerus Solier, 1848, a genus that includes approximately 50 valid species.

Dilopersina Reitter, 117 [F]. Type Prosodesjakowlewi Semenov, 1894, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Dimeriseis Solier, 530 [F]. Type Erodiuslaevigatus G.-A. Olivier, 1792, by subsequent designation (Löbl et al. 2008a: 41). Status: valid subgenus of Erodius Fabricius, 1775 in Pimeliinae: Erodiini.

Dimoniacis Koch, 44 [F]. Type Dimoniacisjacksoni Koch, 1958, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Dimorphochilus Borchmann, 352 [M]. Type Dimorphochilusapicalis Borchmann, 1908, by subsequent designation (R. Lucas 1920: 240). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Dinax Gistel, xi [M]. Type species [automatic]: Erotylusfasciatus Fabricius, 1781, by subsequent designation (Hope 1841: 133). Status: junior synonym of Poecilesthus Dejean, 1834 in Stenochiinae: Stenochiini. Note: unnecessary replacement name for Poecilesthus Dejean, 1834.

Dineria Motschulsky, 530 [F]. Type Blapsconfusa Ménétriés, 1832 (= Blaps halophila Fischer, 1820), by subsequent designation (G.S. Medvedev and Iwan 2006: 617). Status: valid subgenus of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina.

Dinomus Brême, 113 [M]. Type Dinomusperforatus Brême, 1842, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Dinoria Pascoe, 141 [F]. Type Dinoriapicta Pascoe, 1869, by monotypy. Status: junior synonym of Brycopia Pascoe, 1869 in Lagriinae: Adeliini. Synonymy: Carter (1920a: 238).

Dinoscelis Gerstaecker, 533 [F]. Type Dinoscelispasserinii Gerstaecker, 1854, by monotypy. Status: valid subgenus of Pycnocerus Westwood, 1841 in Lagriinae: Pycnocerini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Diodontes Solier, 508, 518 [M]. Type Diodontesporcatus Solier, 1834, by subsequent designation (Hope 1841: 114). Status: valid genus in Pimeliinae: Erodiini.

Dioedus J.L. LeConte, 238 [M]. Type Dioeduspunctatus J.L. LeConte, 1862, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Diopethes Pascoe, 32 [M]. Type Diopethesarachnoides Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Diopoenus Champion, 445 [M]. Type Diopoenuscompressicornis Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Diorhychina Borchmann, 239, 472 [F]. Type Diorhychinaperforata Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Dioscoridemus Koch, 115 [M]. Type Dioscoridemusvonhayekae Koch, 1970, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Dioxycula Fairmaire, 115 [F]. Type Dioxyculaaranea Fairmaire, 1896, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Diphyrrhynchus Fairmaire, 445 [M]. Type Diphyrrhynchuschalceus Fairmaire, 1849, by monotypy. Status: valid genus in Blaptinae: Opatrini: Heterotarsina. Note: Diphyrrhynchus is an incorrect subsequent spelling of the original spelling Diphyrhynchus, first used by Gemminger in Gemminger and Harold (1870: 1955), and in prevailing usage; Diphyrrhynchus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Diplocyrtus Quedenfeldt, 257 [M]. Type Diplocyrtusfloccosus Quedenfeldt, 1887, by monotypy. Status: valid genus in Tenebrioninae: Apocryphini.

Diprosodes Reitter, 115 [M]. Type Prosodesbiformis Semenov, 1894, by subsequent designation (G.S. Medvedev 1997: 570). Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Dirosis Miller, 115 [F]. Type Dirosisnervosa Miller, 1858 (= Erodiusservillei Solier, 1834), by monotypy. Status: valid subgenus of Erodius Fabricius, 1775 in Pimeliinae: Erodiini.

Dischidus Kolbe, 297 [M]. Type Helopssinuatus Fabricius, 1801 (= Tenebriolaevigatus Fabricius, 1781), by present designation. Status: junior synonym of Taraxides C.O. Waterhouse, 1876 in Stenochiinae: Cnodalonini. Synonymy: Champion (1891: 637). Note: the original combination of the accepted name of the type species, Tenebriolaevigatus Fabricius, 1781, is a junior primary homonym of Tenebriolaevigatus Linnaeus, 1767.

Dischizillus Wasmann, 244 [M]. Type species [automatic]: Schizillusrogersi Wasmann, 1899, by monotypy. Status: junior synonym of Pseudethas Fairmaire, 1896 in Pimeliinae: Stenosini: Dichillina. Note: replacement name for Schizillus Wasmann, 1899.

Discodemus J.L. LeConte, 223 [M]. Type Zophosisreticulata Say, 1824, by monotypy. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: J.L. LeConte (1866a: 60).

Discogenia J.L. LeConte, 117 [F]. Type Eleodesscabricula J.L. LeConte, 1858, by subsequent designation (Bousquet et al. 2018: 149). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Discopleurus Lacordaire, 105 [M]. Type species [automatic]: Pleurophorusquadricollis Solier, 1851, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina. Note: replacement name for Pleurophorus Solier, 1851.

Discotus Reitter, 161 [M]. Type Opatrumdilectans Faldermann, 1836, by subsequent designation (G.S. Medvedev 1990: 185). Status: valid subgenus of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina.

Disema Mäklin, 646 [F]. Type Disemabimaculata Mäklin, 1875, by subsequent designation (R. Lucas 1920: 244). Status: valid genus in Lagriinae: Lagriini: Statirina.

Disemorpha Pic, 16 [F]. Type Disemorphaciliata Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Dissonomus Jacquelin du Val, 280 [M]. Type species [automatic]: Heliopatespicipes Faldermann, 1837, by subsequent designation (Gebien 1938a: 397). Status: valid genus and subgenus in Tenebrioninae: Dissonomini. Note: replacement name for Heterophylus Mulsant & Rey, 1859.

Distretus Haag-Rutenberg, 42 [M]. Type Molurisamplipennis Fåhraeus, 1870, by subsequent designation (Rye 1873: 287). Status: valid genus and subgenus in Pimeliinae: Sepidiini: Molurina.

Ditaphronotus Casey, 289, 341 [M]. Type Emmenastusfoveicollis Champion, 1884, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Diversogria Pic, 229 [F]. Type Heterogrialepersonnei Pic, 1954, by original designation. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Dividiopsa Koch, 158 [F]. Type Gnophotaangolensis Harold, 1879, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Divieta Reitter, 390 [F]. Type Vietacostata Allard, 1874, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Vieta Laporte, 1840 in Pimeliinae: Sepidiini: Sepidiina. Synonymy: Lesne (1922: 696).

Doderoella Schuster, 133 [F]. Type Doderoellacyrenaica Schuster, 1926, by monotypy. Status: junior synonym of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Synonymy: Kwieton (1982: 35).

Dolamara Reichardt, 251 [F]. Type Melanimoncupreomicans Reitter, 1915, by original designation. Status: valid genus in Tenebrioninae: Melanimonini.

Dolichasida Reitter, 40, 58 [F]. Type Asidamoraguezi Schaufuss, 1879, by subsequent designation (Wilke 1922: 258). Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: Gebien (1937a: 733; with Insulasida Escalera, 1922, a junior synonym of Asida Latreille, 1802).

Dolichoderus Klug, 87 [M]. Type Dolichoderusacuminatus Klug, 1833, by monotypy. Status: senior synonym of Macellocerus Solier, 1848 in Tenebrioninae: Toxicini: Nycteropina. Note: junior homonym of Dolichoderus Lund, 1831 [Hymenoptera].

Doliema Pascoe, 50 [N]. Type Doliemaplatisoides Pascoe, 1860, by monotypy. Status: junior synonym of Adelina Dejean, 1835 in Diaperinae: Diaperini: Adelinina. Synonymy: Fleutiaux and Sallé (1890: 428), Spilman (1973: 40).

Doliodesmus Spilman, 149 [M]. Type Doliodesmuscharlesi Spilman, 1967, by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Doliopines Horn, 427 [M]. Type Doliopinescucujinus Horn, 1894, by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Dolphus Blanchard, pl. 11 [M]. Type Dolphusglobipennis Blanchard, 1847, by monotypy. Status: valid genus in Tenebrioninae: Helopini: incertae sedis.

Donaciolagria Pic, 14 [F]. Type Donaciolagriaimpressipennis Pic, 1914, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Donisiellus Bremer, 111, 113 [M]. Type Donisiellusdecellei Bremer, 1992, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Doranalia Novák, 481 [F]. Type Cistelarufipennis Marseul, 1876, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Dordanea Reitter, 357 [F]. Type Dordaneaelegans Reitter, 1887, by monotypy. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Dorelogena Péringuey, 280 [F]. Type Dorelogenacastanea Péringuey, 1904, by subsequent designation (R. Lucas 1920: 248). Status: valid genus in Stenochiinae: Stenochiini.

Dorota Novák, 452 [F]. Type Allecularufoposticalis Pic, 1944, by original designation. Status: valid genus in Alleculina: Alleculini: Alleculina.

Dorrigonum Matthews, 708, 752 [N]. Type Licinomaumbilicata Carter, 1924, by original designation. Status: valid genus in Lagriinae: Adeliini.

Doryagus Pascoe, 12 [M]. Type Doryagustalpa Pascoe, 1887, by monotypy. Status: junior synonym of Stizopus Erichson, 1843 in Blaptinae: Opatrini: Stizopodina. Synonymy: Gebien (1920: 120).

Doyenia Matthews & Lawrence, 532 [F]. Type Doyeniacrematogastri Matthews & Lawrence, 2005, by original designation. Status: valid genus in Lagriinae: Belopini.

Doyenus Iwan, 385, 386 [M]. Type Doyenusuncus Iwan, 1996, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Drepanomela Borchmann, 119 [F]. Type Lagriacribratula Schaufuss, 1887, by original designation. Status: valid subgenus of Cerogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina.

Drocleana Bates, 291 [F]. Type Camariachalcoptera Klug, 1833, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Drosochrus Erichson, 243 [M]. Type Drosochrusbrunnipes Erichson, 1843, by subsequent designation (Gebien 1943: 910). Status: valid genus and subgenus in Blaptinae: Pedinini: Helopinina.

Dryadigmus Bremer, 26 [M]. Type Amarygmuscechovskyi Bremer, 2007, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Duocula Novák, 56 [F]. Type Oraculaclara Novák, 2019, by original designation. Status: valid subgenus of Oracula Novák, 2019 in Alleculinae: Alleculini: Alleculina.

Durandius Kaszab, 115 [M]. Type Durandiusardoini Kaszab, 1970, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Rondoniellina.

Durasida Reitter, 9, 14 [F]. Type Tenebriosilphoides Linnaeus, 1767, by subsequent designation (F. Soldati 2008: 32). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Dymonus Solier, 7, 121 [M]. Type Sepidium vestitum Guérin-Méneville, 1831, by original designation. Status: junior synonym of Vieta Laporte, 1840 in Pimeliinae: Sepidiini: Sepidiina. Synonymy: Lacordaire (1859a: 205).

Dysantes Pascoe, 31 [M]. Type Dicerodereselongatus Redtenbacher, 1868, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina. Note: this name is not a junior homonym of Dysantes Forster, 1869 [Hymenoptera], as previously reported in the literature, since the name in Coleoptera was published 1 January 1869 whereas the name in Hymenoptera was issued in May 1869 (see Bouchard and Bousquet 2020a: 101 for additional comments).

Dysarchus Pascoe, 449 [M]. Type Dysarchusodewahnii Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Dysceladus C.O. Waterhouse, 411 [M]. Type Dysceladustuberculatus C.O. Waterhouse, 1875 (= Polposipusherculeanus Solier, 1848), by monotypy. Status: junior synonym of Polposipus Solier, 1848 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1922b: 268, 311).

Dysgena Mäklin, 558 [F]. Type Dysgenalugubris Mäklin, 1863, by subsequent designation (R. Lucas 1920: 252). Status: valid genus in Tenebrioninae: Praeugenini.

Dysodera Borchmann, 238, 470 [F]. Type Dysoderamethneri Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Dysopinus Borchmann, 236, 379 [M]. Type Nemostiraserra Borchmann, 1913, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Dystalica Pascoe, 142 [F]. Type Dystalicahomogenea Pascoe, 1869, by monotypy. Status: junior synonym of Adelium W. Kirby, 1819 in Lagriinae: Adeliini. Synonymy: Matthews (1998: 777).

Dzhungaropterocoma Skopin, 150 [F]. Type Pterocomasubnuda Reitter, 1897, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Earophanta Semenov, 172 [F]. Type species [automatic]: Platyopeserrata Semenov, 1893, by original designation. Status: valid genus in Pimeliinae: Pimeliini. Note: replacement name for Earophila Semenov, 1903.

Earophila Semenov, 9 [F]. Type Platyopeserrata Semenov, 1893, by original designation. Status: senior synonym of Earophanta Semenov, 1903 in Pimeliinae: Pimeliini. Note: junior homonym of Earophila Gumppenberg, 1887 [Lepidoptera].

Earophilina Strand, 99 [F]. Type species [automatic]: Platyopeserrata Semenov, 1893, by original designation. Status: junior synonym of Earophanta Semenov, 1903 in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Earophila Semenov, 1903.

Eba Pascoe, 122, 129 [F]. Type Ebacerylonoides Pascoe, 1863, by monotypy. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Carter and Zeck (1937: 194). Note: transferred from “Tenebrionoidea: Colydiidae” by Carter and Zeck (1937: 194).

Ebenolus Fairmaire, 227 [M]. Type Ebenolusvernicatus Fairmaire, 1897, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Kaszab (1977b: 29).

Ebertius Jedlička, 98 [M]. Type Ebertiusnepalensis Jedlička, 1965 (= Laenalongipilis Schuster, 1926), by original designation. Status: junior synonym of Laena Dejean, 1821 in Lagriinae: Laenini. Synonymy: Kaszab (1970d: 429, through placement of the type species in Laena Dejean, 1821). Note: originally described in the family Carabidae.

Eccoptostira Borchmann, 236, 377 [F]. Type Nemostirarohdei Pic, 1912, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: we act as First Revisers and reject the alternative original spelling Ecoptostira, used by Borchmann (1936: 236).

Eccoptostoma Gebien, 70 [N]. Type Taraxidesruficrus Fairmaire, 1894, by present designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: redescribed as new by Gebien (1921b: 62, 82).

Echinotrigon Skopin, 170 [M]. Type Trigonoscelisgranulata Reitter, 1915, by original designation. Status: valid subgenus of Trigonoscelis Dejean, 1834 in Pimeliinae: Pimeliini.

Echinotus Solier, 30 [M]. Type Sepidium spinicolle Laporte, 1840, by original designation [p. 122]. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Echocerus Horn, 364, 366 [M]. Type Trogossitamaxillosa Fabricius, 1801, by monotypy. Status: valid subgenus of Gnatocerus Thunberg, 1814 in Diaperinae: Diaperini: Adelinina.

Ecnocera Borchmann, 22, 220 [F]. Type Porrolagriagracilis Borchmann, 1909, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Ecnolagria Borchmann, 49, 139 [F]. Type Lagriagrandis Gyllenhal, 1817, by original designation. Status: valid genus and subgenus in Lagriinae: Lagriini: Lagriina.

Ecnomoderes Gebien, 109, 110 [M]. Type Ecnomoderesbarbatus Gebien, 1928, by subsequent designation (Gebien 1937a: 690). Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Ecnomosternum Gebien, 104 [N]. Type Ecnomosternumvermiculatum Gebien, 1928, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Ecphoroma Solier, 195 [N]. Type Pimeliahemisphaerica Solier, 1836, by subsequent designation (Löbl et al. 2008b: 157). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Ecripsis Pascoe, 456 [F]. Type Ecripsispubescens Pascoe, 1866, by monotypy. Status: junior synonym of Ammidium Erichson, 1843 in Blaptinae: Opatrini: Ammobiina. Synonymy: Blair in Carter (1914c: 406). Note: this African genus was originally described from Australia in error (see Matthews and Bouchard 2008: 351).

Ectateus Koch, 230 [M]. Type Anchophthalmusmodestus Fairmaire, 1887, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Ectatocera Fåhraeus, 325 [F]. Type Ectatoceralongicornis Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Ectatocnemis Horn, 400 [F]. Type Ectatocnemismultilineata Horn, 1867, by monotypy. Status: junior synonym of Anomalipus Guérin-Méneville, 1831 in Blaptinae: Platynotini: Platynotina. Synonymy: Endrödy-Younga (1988: 19). Note: this taxon was originally described from Chile in error, it is native to South Africa (see Endrödy-Younga 1988: 19).

Ectenostoma Fåhraeus, 317 [N]. Type Ectenostomanigriventris Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Ectomopsis Fairmaire, 299 [F]. Type Ectomopsisbruchi Fairmaire, 1905, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ectromopsis Antoine, 125, 145 [F]. Type Catomuspoliticollis Allard, 1876, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Ectyche Pascoe, 143 [F]. Type Ectycheerebea Pascoe, 1869, by monotypy. Status: valid genus in Diaperinae: Ectychini. Note: “Ectyche? nana Pascoe, 1869” was doubtfully included in this genus by Pascoe (1869: 143) and is therefore not an originally included species (ICZN 1999: Article 67.2.5).

Edalus Broun, 1159 [M]. Type Edalusopacus Broun, 1893, by subsequent designation (R. Lucas 1920: 255). Status: senior synonym of Wattadelium Emberson, 2000 in Lagriinae: Adeliini. Note: junior homonym of Edalus Broun, 1886 [Coleoptera: Zopheridae].

Edrotes J.L. LeConte, 140 [M]. Type Edrotesventricosus J.L. LeConte, 1851, by monotypy. Status: valid genus and subgenus in Pimeliinae: Edrotini.

Edrotinus Fairmaire, 461 [M]. Type Edrotinustucumanus Fairmaire, 1904, by original designation. Status: junior synonym of Trilobocara Solier, 1851 in Pimeliinae: Trilobocarini. Synonymy: Gebien (1937a: 592).

Edrotopus Haag-Rutenberg, 129 [M]. Type Edrotopusstrigicollis Haag-Rutenberg, 1877, by monotypy. Status: junior synonym of Platyholmus Dejean, 1834 genus in Pimeliinae: Praociini. Synonymy: Fairmaire (1884a: 508).

Edylius Champion, 353 [M]. Type Edyliuscanescens Champion, 1894, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Eichleria Kamiński, 132 [F]. Type Eichleriaostrowskii Kamiński, 2015, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Elaeodes Gemminger in Gemminger and Harold, 1868 [F]. Type species [automatic]: Eleodesdentipes Eschscholtz, 1829, by subsequent designation (Hope 1841: 124). Status: junior synonym of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini. Note: unjustified emendation of Eleodes Eschscholtz, 1829, not in prevailing usage.

Elaeus Gemminger in Gemminger and Harold, 1969 [M] Type species [automatic]: Heleaperforata Latreille, 1816, by subsequent monotypy (Latreille 1817: 261). Status: junior synonym of Helea Latreille, 1804 in Tenebrioninae: Heleini: Heleina. Note: unjustified emendation of Helea Latreille, 1804 (as “Helaeus Latreille”), not in prevailing usage.

Elasmocera Mäklin, 504 [F]. Type Elasmoceradentipes Mäklin, 1867, by monotypy. Status: senior synonym of Elasmocerella Strand, 1935 in Stenochiinae: Stenochiini. Note: junior homonym of Elasmocera Rondani, 1846 [Diptera].

Elasmocerella Strand, 302 [F]. Type species [automatic]: Elasmoceradentipes Mäklin, 1867, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: replacement name for Elasmocera Mäklin, 1867.

Eleates Casey, 253 [M]. Type Eleatesoccidentalis Casey, 1886, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Eledona Latreille, 19 [F]. Type Opatrumagricola Herbst, 1783, by subsequent monotypy (Latreille 1802: 162). Status: valid genus in Tenebrioninae: Bolitophagini. Note: originally proposed without included nominal species; Latreille (1802: 162)) , by including the species “Bolitophagusagricola” in association with this name, was the first author to subsequently and expressly include nominal species in Eledona (ICZN 1999, Article 67.2.2).

Eledonoprius Reitter, 329, 338 [M]. Type Opatrumarmatum Panzer, 1799, by subsequent designation (R. Lucas 1920: 258). Status: valid genus in Tenebrioninae: Bolitophagini.

Elenophorus Dejean, 64 [M]. Type Tenebriocollaris Linnaeus, 1767, by monotypy. Status: junior synonym of Leptoderis Billberg, 1820 in Pimeliinae: Elenophorini: Elenophorina. Synonymy: Silfverberg (1984: 59).

Eleodes Eschscholtz, 8 [F]. Type Eleodesdentipes Eschscholtz, 1829, by subsequent designation (Hope 1841: 124). Status: valid genus and subgenus in Blaptinae: Amphidorini.

Eleodimorpha Blaisdell, 477 [F]. Type Eleodimorphabolcan Blaisdell, 1909, by original designation. Status: valid genus in Blaptinae: Amphidorini.

Eleodopsis Blaisdell, 52 [F]. Type Eleodopsissubvestita Blaisdell, 1939, by original designation. Status: junior synonym of Blapylis Horn, 1870 in Blaptinae: Amphidorini. Synonymy: Spilman (1962a: 59).

Eleoselinus Kamiński, 101 [M]. Type Ectateusvilliersi Ardoin, 1965, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Elixota Pascoe, 475 [F]. Type Elixotacuprea Pascoe, 1866 (= Amarygmushopei Bremer, 2001), by original designation. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer (2001a: 65).

Ellaemus Pascoe, 495 [M]. Type Emcephalussubmaculatus Brême, 1842, by present designation. Status: junior synonym of Emcephalus W. Kirby, 1828 in Tenebrioninae: Heleini: Heleina. Synonymy: Matthews and Bouchard (2008: 304).

Ellidoneus Wilke, 277 [M]. Type Cardigeniusgranulatus Fairmaire, 1873, by original designation. Status: valid subgenus of Cardigenius Solier, 1836 in Pimeliinae: Asidini.

Ellipsodes Wollaston, 485 [M]. Type Sphaeridiumglabratum Fabricius, 1781, by monotypy. Status: valid genus and subgenus in Diaperinae: Crypticini.

Elomosda Bates, 273 [F]. Type Elomosdabeltii Bates, 1870, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Elongasida Escalera, 306 [F]. Type Asidagrandipalpis Allard, 1869, by subsequent designation (Viñolas and Cartagena 2005: 164). Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini.

Embaphion Say, 254 [N]. Type Akismuricata Say, 1824, by monotypy. Status: valid genus in Blaptinae: Amphidorini.

Emcephalus W. Kirby, 524 [M]. Type Emcephalusgibbosus W. Kirby, 1828, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Emeacoides Gebien, 698 [M]. Type species [automatic]: Enneacoidesvinculiger Fairmaire, 1881 (= Nyctoporiscarinata J.L. LeConte, 1851), by monotypy. Status: junior synonym of Nyctoporis Eschscholtz, 1831 in Pimeliinae: Nyctoporini. Note: unjustified emendation of Enneacoides Fairmaire, 1881, not in prevailing usage.

Emeax Pascoe, 450 [M]. Type Emeaxsculpturatus Pascoe, 1866 (= Nyctoporiscristata Eschscholtz, 1831), by monotypy. Status: junior synonym of Nyctoporis Eschscholtz, 1831 in Pimeliinae: Nyctoporini. Synonymy: J.L. LeConte (1873: 334). Note: as mentioned by Matthews and Bouchard (2008: 351) the type species was originally described from Australia in error; the genus Nyctoporis Eschscholtz, 1831 is endemic to the Nearctic realm.

Emmallodera Blanchard, pl. 13 [F]. Type Emmalloderacrenaticostata Blanchard, 1842, by monotypy. Status: valid genus in Tenebrioninae: Scotobiini. Note: unjustified emendation of the original spelling Emalodera, introduced by Agassiz (1846b: 137), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Emmallus Agassiz, 137 [M]. Type species [automatic]: Emmaluspilosus Erichson, 1843, by monotypy. Status: junior synonym of Emmalus Erichson, 1843 in Blaptinae: Opatrini: Ammobiina. Note: unjustified emendation of Emmalus Erichson, 1843, not in prevailing usage.

Emmalus Erichson, 251 [M]. Type Emmaluspilosus Erichson, 1843, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Emmenastrichus Horn, 413 [M]. Type Emmenastrichuscribratus Horn, 1894, by subsequent designation (Casey 1907: 289). Status: valid genus in Pimeliinae: Edrotini.

Emmenastus Motschulsky, 75 [M]. Type Emmenastusrugosus Motschulsky, 1845, by subsequent designation (J.L. LeConte 1866b: 106). Status: junior synonym of Oxycara Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Aalbu et al. (1995: 484).

Emmenides Casey, 329 [M]. Type Emmenastuspunctatus J.L. LeConte, 1866, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Emydodes Pascoe, 56 [M]. Type Emydodescollaris Pascoe, 1860, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Emyon Gerstaecker, 532 [M]. Type Emyoncaelatus Gerstaecker, 1854, by monotypy. Status: junior synonym of Helopinus Solier, 1848 in Blaptinae: Pedinini: Helopinina. Synonymy: Koch (1958: 149).

Emypsara Pascoe, 460 [F]. Type Emypsaraadamsii Pascoe, 1866 (= Diaperisriederii Faldermann, 1833), by subsequent designation (Löbl et al. 2008a: 41). Status: valid genus in Diaperinae: Phaleriini.

Enanea Lewis, 467 [F]. Type Enaneatestacea Lewis, 1894, by monotypy. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Encara Gemminger, 124 [N]. Type species [automatic]: Emcephalusgibbosus W. Kirby, 1828, by monotypy. Status: junior synonym of Emcephalus W. Kirby, 1828 in Tenebrioninae: Heleini: Heleina. Note: replacement name for Emcephalus W. Kirby, 1828 (as “Encephalus Brême”).

Encephalus Agassiz, 137 [M]. Type species [automatic]: Emcephalusgibbosus W. Kirby, 1828, by monotypy. Status: junior synonym of Emcephalus W. Kirby, 1828 in Tenebrioninae: Heleini: Heleina. Note: unjustified emendation of Emcephalus W. Kirby, 1828, not in prevailing usage; junior homonym of Encephalus Stephens, 1832 [Coleoptera: Staphylinidae].

Encyalesthus Motschulsky, 139 [M]. Type Encyalesthussubviolaceus Motschulsky, 1860, by monotypy. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1987: 43).

Endostomus Gemminger in Gemminger and Harold, 1973 [M]. Type species [automatic]: Cossyphussenegalensis Laporte, 1833, by subsequent monotypy (Duponchel 1844b: 315). Status: junior synonym of Endustomus Brême, 1842 in Lagriinae: Cossyphini. Note: unjustified emendation of Endustomus Brême, 1842, not in prevailing usage.

Endostostomus Jakobson, 528 [M]. Type species [automatic]: Cossyphussenegalensis Laporte, 1833, by subsequent monotypy (Duponchel 1844b: 315). Status: junior synonym of Endustomus Brême, 1842 in Lagriinae: Cossyphini. Note: unjustified emendation of Endustomus Brême, 1842, not in prevailing usage.

Endothina Carter, 536 [F]. Type Endothinasquamosa Carter, 1924 (= Opatrumcanaliculatum Fabricius, 1798), by monotypy. Status: junior synonym of Leichenum Dejean, 1834 in Blaptinae: Pedinini: Leichenina. Synonymy: Carter (1928: 284).

Endroeditagalus Schawaller & Bouchard, 192 [M]. Type Endroeditagalusntsubanus Schawaller & Bouchard, 2019, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Endustomus Brême, 17 [M]. Type Cossyphussenegalensis Laporte, 1833, by subsequent monotypy (Duponchel 1844b: 315). Status: valid genus in Lagriinae: Cossyphini. Note: originally proposed without included nominal species; Duponchel (1844b: 315), by including the species “Cossyphussenegalensis” in association with this name, was the first author to subsequently and expressly include nominal species in Endustomus (ICZN 1999, Article 67.2.2).

Enganodia Fairmaire, 398 [F]. Type Enganodiasanguinicrus Fairmaire, 1898, by monotypy. Status: junior synonym of Lophocnemis Mäklin, 1867 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 543).

Enicmonota Ardoin, 61 [F]. Type Enicmosomacrassicorne Ardoin, 1959, by monotypy. Status: junior synonym of Enicmosoma Gebien, 1922 in Lagriinae: Lupropini. Synonymy: Schawaller (2013a: 139).

Enicmosoma Gebien, 312 [N]. Type Enicmosomapunctum Gebien, 1922, by subsequent designation (Gebien 1941: 823). Status: valid genus in Lagriinae: Lupropini.

Enigmatica Ferrer, 202 [F]. Type Enigmaticaendroedyi Ferrer, 2005, by monotypy. Status: valid genus in Lagriinae: Laenini.

Enneacoides Fairmaire, 277 [M]. Type Enneacoidesvinculiger Fairmaire, 1881 (= Nyctoporiscarinata J.L. LeConte, 1851), by monotypy. Status: junior synonym of Nyctoporis Eschscholtz, 1831 in Pimeliinae: Nyctoporini. Synonymy: Gebien (1908b: 287).

Ennychiatus Koch, 28 [M]. Type Stizopuscaraboides Fairmaire, 1897, by original designation. Status: junior synonym of Parastizopus Gebien, 1938 in Blaptinae: Opatrini: Stizopodina. Synonymy: Iwan and Schimrosczyk (2017: 384).

Ennychius Fåhraeus, 299 [M]. Type Ennychiusmorio Fåhraeus, 1870, by monotypy. Status: junior synonym of Helibatus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Stizopodina. Synonymy: Gebien (1938a: 74).

Enoplopus Solier, 151, 158 [M]. Type species [automatic]: fixed herein (ICZN 1999, Article 70.3) as Tenebriovelikensis Piller & Mitterpacher, 1783, misidentified as Tenebriocaraboides Linnaeus, 1758 in the original designation by monotypy in Dejean (1821). Status: junior synonym of Accanthopus Dejean, 1821 in Tenebrioninae: Helopini: Enoplopodina. Note: unnecessary replacement name for Accanthopus Dejean, 1821.

Entinopoda Gebien, 64 [F]. Type Eustolopusoctoseriatus Gebien, 1938, by monotypy. Status: junior synonym of Eustolopus Gebien, 1938 in Pimeliinae: Adesmiini. Synonymy: Penrith (1979: 40).

Entomobalia Flores & Triplehorn, 607 [F]. Type Asidaplatynotos Perty, 1830, by original designation. Status: valid genus in Pimeliinae: Nycteliini.

Entomochilus Gay & Solier in Solier, 48 [M]. Type Entomochiluspilosus Gay & Solier, 1843, by original designation. Status: valid genus in Pimeliinae: Physogasterini.

Entomoderes Solier, 308, 346 [M]. Type Entomodereserebi Solier, 1836, by monotypy. Status: valid genus in Pimeliinae: Nycteliini.

Entomogonus Solier, 151, 155 [M]. Type Entomogonusbarthelemyi Solier, 1848, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina.

Entypodera Gerstaecker, 66 [F]. Type Entypoderaanthicoides Gerstaecker, 1871, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: new placement [OM], previously included in Lagriinae: Lagriini: Lagriina.

†Eoallognosis Haupt, 115, 137 [F]. Type Eoallognosisundulata Haupt, 1950, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

†Eocallidium Haupt, 143 [N]. Type Eocallidiumrugulosum Haupt, 1950, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: originally included in the family Cerambycidae, transferred to Tenebrionidae by Vitali (2008: 8); described from Middle Eocene deposits (Germany).

Eocyphogenia G.S. Medvedev, 897, 898 [F]. Type Akisrugipennis Faldermann, 1835, by original designation. Status: junior synonym of Cyphogenia Solier, 1837 in Pimeliinae: Akidini. Synonymy: Ren and Yu (2000: 54).

Eodirosis Kwieton, 25 [F]. Type Erodiusquadrilineatus Kraatz, 1865, by original designation. Status: valid subgenus of Erodius Fabricius, 1775 in Pimeliinae: Erodiini. Note: the original combination of the accepted name of the type species, Erodiusquadrilineatus Kraatz, 1865, is a junior primary homonym of Erodiusquadrilineatus G.-A. Olivier, 1792.

†Eodromus Haupt, 113, 120 [M] Type †Ancylochiraagilis Meunier, 1915, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: originally proposed in the family Carabidae by Pongrácz (1935: 538) without type species (ICZN 1999, Article 13.3); we act as First Revisers and reject the alternative original spelling Edromus, used by Haupt (1950: 120); described from Middle Eocene deposits (Germany).

†Eohelaeus Haupt, 115, 135 [M]. Type Eohelaeussublaevis Haupt, 1950, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Epairops Fåhraeus, 282 [M]. Type Epairopsfragilis Fåhraeus, 1870, by monotypy. Status: junior synonym of Ossiporis Pascoe, 1866 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Gebien (1937b: 37).

Epairopsis Koch, 47 [F]. Type Trachynotusfrontalis Haag-Rutenberg, 1873, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Epantius J.L. LeConte, 144 [M]. Type Epantiusobscurus J.L. LeConte, 1851, by monotypy. Status: valid genus in Tenebrioninae: Eulabini.

Epeurycaulus Kolbe, 579 [M]. Type Epeurycaulusaldabricus Kolbe, 1902, by present designation. Status: junior synonym of Plesioderes Mulsant, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1922b: 268).

Ephalus J.L. LeConte, 228 [M]. Type Heliopateslatimanus J.L. LeConte, 1847, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: transferred from Ammobiina by Lumen et al. (2020: 344).

Ephidonius Pascoe, 151 [M]. Type Ephidoniusacuticornis Pascoe, 1869, by original designation. Status: junior synonym of Brises Pascoe, 1869 in Tenebrioninae: Heleini: Heleina. Synonymy: Carter (1914b: 46).

Epicalla Lacordaire, 309 [F]. Type Epicallavaripes Champion, 1886, by subsequent designation (R. Lucas 1920: 268). Status: valid genus in Stenochiinae: Cnodalonini. Note: the authorship of this genus name has been credited to Champion (1886: 249) in the literature; however, Lacordaire (1859a: 309) mentioned that the genus Epicalla has the “mandibules entières au bout”, which fulfils the requirements of availability (ICZN 1999, Article 12.1); Champion (1886: 249–251), by describing three new species in association with the genus Epicalla, was the first author to subsequently and expressly include nominal species in this genus (ICZN 1999, Article 67.2.2).

Epicydes Champion, 60 [M]. Type Epicydesoculatus Champion, 1889, by subsequent designation (Borchmann 1936: 429). Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Epilamprus Gistel, xiv [M]. Type species [automatic]: Helopsindutus Wiedemann, 1819, by subsequent designation (Duponchel 1844b: 359). Status: junior synonym of Ceropria Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Note: unjustified emendation of Epilampus Dejean, 1834, not in prevailing usage.

Epilampus Dejean, 198 [M]. Type species [automatic]: Helopsindutus Wiedemann, 1819, by subsequent designation (Duponchel 1844b: 359). Status: junior synonym of Ceropria Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Note: unnecessary replacement name for Ceropria Laporte & Brullé, 1831 (see Bousquet and Bouchard 2013a: 52).

Epilasium Erichson, 237 [N]. Type Trichotoncayennense Hope, 1841, by monotypy. Status: junior synonym of Trichoton Hope, 1841 in Blaptinae: Opatrini: Blapstinina. Synonymy: Erichson (1842a: 237). Note: the name Epilasium was listed as synonym of Trichoton Hope, 1841 by Erichson (1842a: 237), it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Curtis 1844: 222), Epilasium was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Epipagus Haag-Rutenberg, 274, 311 [M]. Type Epipagusbenguelensis Haag-Rutenberg, 1872, by monotypy. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Epipedodema Gebien, 54 [F]. Type Epipedodemadepressa Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Alphitobiini.

Epipedonota Solier, 307, 342 [F]. Type Epipedonotaebenina Solier, 1836, by subsequent designation (Solier 1851: 159). Status: valid genus in Pimeliinae: Nycteliini. Note: the alternative original spelling Epipedonata, used by Solier (1836: 342), was rejected by Solier (1838a: 487) who acted as the First Reviser.

Epiphaleria Lewis, 382 [F]. Type Epiphaleriaatriceps Lewis, 1894, by original designation. Status: valid subgenus of Phaleria Latreille, 1802 in Diaperinae: Phaleriini.

Epiphysa Dejean, 178 [F]. Type Pimeliaflavicollis Fabricius, 1794, by monotypy. Status: valid genus in Pimeliinae: Adesmiini.

Epiplecta Mäklin, 498 [F]. Type Epiplectamaculata Mäklin, 1867, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Episopus Bates, 372 [M]. Type Episopuspolitus Bates, 1873, by subsequent designation (R. Lucas 1920: 270). Status: valid genus in Stenochiinae: Cnodalonini.

Epitoxicum Bates, 46 [N]. Type Epitoxicumhaplandroides Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Toxicina.

Epitragella Kulzer, 184 [F]. Type Epitragelladimorpha Kulzer, 1958, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Epitragodes Casey, 365 [M]. Type Epitragustomentosus J.L. LeConte, 1866, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Epitragoma Casey, 386 [N]. Type Epitragusvestitus Casey, 1891, by monotypy. Status: junior synonym of Pechalius Casey, 1907 in Pimeliinae: Epitragini. Synonymy: Freude (1968: 61).

Epitragopsis Casey, 386 [F]. Type Epitragusgodmani Champion, 1884, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Epitragosoma K.W. Brown & Triplehorn, 515 [N]. Type Epitragosomaarenaria K.W. Brown & Triplehorn, 2002, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Epitragus Latreille, 165 [M]. Type Epitragusfuscus Latreille, 1804, by subsequent monotypy (Latreille 1804: 322). Status: valid genus and subgenus in Pimeliinae: Epitragini. Note: originally proposed without included nominal species since the species listed (Helopsvariegatus ? Fab.) is conditionally included; Latreille (1804: 322), by including the species Epitragusfuscus Latreille, 1804 in association with this name, was the first author to subsequently and expressly include a nominal species in Epitragus (ICZN 1999, Article 67.2.2).

Epitrichia Gebler, 475 [F]. Type Helopstomentosus Gebler, 1842, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Epityria Koch, 298 [F]. Type Himatismusstriatopunctatus Haag-Rutenberg, 1877, by original designation. Status: valid subgenus of Derosphaerius Westwood, 1881 in Pimeliinae: Tentyriini.

Epomidus Matthews, 705, 763 [M]. Type Epomidusprionodes Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Erelus Mulsant & Rey, 185 [M]. Type Erelussulcipennis Mulsant & Rey, 1853, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: included in a list of “Tenebrionidaenomina dubia” by Iwan et al. (2020: 475).

Eremobates Gebien, 120 [M]. Type Eremobatescrux Gebien, 1921, by monotypy. Status: senior synonym of Eremobatodes Gebien, 1943 in Stenochiinae: Cnodalonini. Note: junior homonym of Eremobates Banks, 1900 [Arachnida].

Eremobatodes Gebien, 404 [M]. Type species [automatic]: Eremobatescrux Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Eremobates Gebien, 1921.

Eremocantor Smith & Wirth, 582 [M]. Type Eremocantormarioni Smith & Wirth, 2016, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Eremoecus Lacordaire, 69 [M]. Type Hyperopseschscholtzii Solier, 1851, by monotypy. Status: valid genus in Pimeliinae: Trilobocarini.

Eremonomus Wollaston, 199 [M]. Type Eremonomushuttoni Wollaston, 1861 (= Ammidiumciliatum Erichson, 1843), by monotypy. Status: junior synonym of Ammidium Erichson, 1843 in Blaptinae: Opatrini: Ammobiina. Synonymy: Ardoin (1971: 358).

Eremophaleria Español, 32, 34 [F]. Type Phaleriabedeli Chobaut, 1900, by original designation. Status: valid subgenus of Phaleria Latreille, 1802 in Diaperinae: Phaleriini.

Eremostibes Koch, 60 [M]. Type Eremostibesopacus Koch, 1963, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Ergenna Fairmaire, 139 [F]. Type Ergennacaerulescens Fairmaire, 1897, by monotypy. Status: junior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Synonymy: De Moor (1970: 42).

Erionura Reitter, 18 [F]. Type Helopsgiganteus Kraatz, 1862, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Ernocharis C.G. Thomson, 118 [F]. Type Cistelabrevis Illiger, 1794 (= Cistelamaura Fabricius, 1792), by original designation. Status: valid subgenus of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina.

Erodibius Löbl, Bouchard, Merkl & Bousquet, 2 [M]. Type Arthrodeiscyphonotus Fairmaire, 1887, by original designation. Status: valid subgenus of Arthrodibius Lesne, 1915 in Pimeliinae: Erodiini. Note: name first proposed by Koch (1960: 354) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl et al. (2008a: 40) designated Arthrodeiscyphonotus Fairmaire, 1887 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Erodinus Reitter, 299 [M]. Type none designated. Status: undetermined taxon in Pimeliinae: Erodiini. Note: this genus was included in a key, which fulfils the criteria of availability for new genus-group names proposed before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Erodinus and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Erodiontes Reitter, 48, 79 [M]. Type Erodiontesvermiculatus Reitter, 1914, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Erodius Fabricius, 258 [M]. Type Erodiusgibbus Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Erulipothydemus Pic, 19 [M]. Type Erulipothydemuslatior Pic, 1918, by subsequent designation (Gebien 1943: 902). Status: valid genus in Tenebrioninae: Helopini: incertae sedis.

Erulipus Fairmaire, 14 [M]. Type Erulipusfruhstorferi Fairmaire, 1903, by monotypy. Status: valid subgenus of Ainu Lewis, 1894 in Stenochiinae: Cnodalonini.

Erxias Champion, 460 [M]. Type Erxiasbicolor Champion, 1888, by subsequent designation (Bousquet et al. 2015: 138). Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Erycastus Fairmaire, 133 [M]. Type Erycastusnavicularis Fairmaire, 1897, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Eryx Stephens, 27 [M]. Type Pyrochroanigra DeGeer, 1775 (= Helopsater Fabricius, 1775), by monotypy. Status: senior synonym of Prionychus Solier, 1835 in Alleculinae: Alleculini: Alleculina. Synonymy: Mulsant (1856a: 12, 61). Note: junior homonym of Eryx Daudin, 1803 [Reptilia].

Erzika Novák, 53 [F]. Type Erzikatamdaoica Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Eschatomoxys Blaisdell, 125 [M]. Type Eschatomoxyswagneri Blaisdell, 1935, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Eschatoporis Blaisdell, 76 [F]. Type Eschatoporisnunenmacheri Blaisdell, 1906, by monotypy. Status: valid genus in Lagriinae: Eschatoporiini.

Eschatostena Keleinikova, 654 [F]. Type Eschatostenakuznetzovi Keleinikova, 1977, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Esemephe Steiner, 385 [F]. Type Esemephetumi Steiner, 1980, by original designation. Status: valid genus in Pimeliinae: Cossyphodini: Esemephina.

Espagnolina Kaszab, 117 [F]. Type Espagnolinaassamica Kaszab, 1965, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Espidium Rafinesque, 113 [N]. Type species [automatic]: Sepidium tricuspidatum Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Sepidium Fabricius, 1775 in Pimeliinae: Sepidiini: Sepidiina. Note: unnecessary replacement name for Sepidium Fabricius, 1775.

Espites Pascoe, 32 [M]. Type Espitesbasalis Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Espitomorphus Pic, 24 [M]. Type Espitomorphusmulticolor Pic, 1921, by monotypy. Status: junior synonym of Camarimena Motschulsky, 1863 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 136).

Etazeta Fairmaire, 358 [F]. Type Etazetaaeneicolor Fairmaire, 1889, by monotypy. Status: junior synonym of Luprops Hope, 1833 in Lagriinae: Lupropini. Synonymy: Gebien (1914a: 35).

Ethas Pascoe, 324 [M]. Type Ethascarbonarius Pascoe, 1862, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Ethmomerus Koch, 243 [M]. Type Ethmussubcylindricus Koch, 1953, by original designation. Status: valid subgenus of Ethmus Haag-Rutenberg, 1873 in Pimeliinae: Sepidiini: Trachynotina.

Ethmophobes Koch, 244 [M]. Type Ethmuslatus Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Ethmus Haag-Rutenberg, 1873 in Pimeliinae: Sepidiini: Trachynotina.

Ethmus Haag-Rutenberg, 44 [M]. Type Ethmusmaculatus Haag-Rutenberg, 1873, by subsequent designation (Gebien 1937a: 778). Status: valid genus and subgenus in Pimeliinae: Sepidiini: Trachynotina. Note: nomenclatural stability is threatened by the discovery of an older type species designation (Ethmuslatus Haag-Rutenberg, 1873, by subsequent designation by R. Lucas (1920: 276), which is the type species of the valid subgenusEthmophobes Koch, 1953); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Gebien (1937a: 778).

Eubalia Laporte, 257 [F]. Type Statiraovalis Laporte, 1840, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Euboeus Boieldieu, 10 [M]. Type Euboeusmimonti Boieldieu, 1865, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Eucaliga Fairmaire & Germain, 5 [F]. Type Eucaligasanguinicollis Fairmaire & Germain, 1861, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Eucamaria Gebien, 28, 149 [F]. Type Camariaspectabilis Pascoe, 1860, by original designation. Status: junior synonym of Falsocamaria Pic, 1917 in Stenochiinae: Cnodalonini. Synonymy: Masumoto (1993a: 142).

Eucamptus Germar, 444 [M]. Type Eucamptusiridis Germar, 1842 (= Hegemonaresplendens Laporte, 1840), by monotypy. Status: junior synonym of Hegemona Laporte, 1840 in Stenochiinae: Cnodalonini. Synonymy: Duponchel (1845b: 498).

Euclarkia Lea, 180 [F]. Type Euclarkiacostata Lea, 1919, by monotypy. Status: valid genus in Lagriinae: Belopini.

Eucolus Mulsant & Rey, 39, 67 [M]. Type Eucoluspolinierii Mulsant & Rey, 1853, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Euconibius Casey, 618 [M]. Type Notibiusgagates Horn, 1870, by monotypy. Status: junior synonym of Conibius J.L. LeConte, 1851 in Blaptinae: Opatrini: Blapstinina. Synonymy: Aalbu in Bousquet et al. (2018: 202).

Eucosmus Gistel, x [M]. Type species [automatic]: Spheniscuserotyloides W. Kirby, 1819, by monotypy. Status: junior synonym of Cuphotes Champion, 1887 in Stenochiinae: Stenochiini. Note: replacement name for Spheniscus W. Kirby, 1819; junior homonym of Eucosmus Agassiz, 1846 [Echinoidea].

Eucrossoscelis Nakane, 29 [F]. Type Eucrossoscelisbroscosomoides Nakane, 1963, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Eucyrtus Lacordaire, 417 [M]. Type Eucyrtuspretiosus Lacordaire, 1859, by subsequent designation (Gebien 1921a: 335). Status: valid genus in Stenochiinae: Cnodalonini. Note: the name Eucyrtus was listed as synonym of Scotaeus Hope, 1834 by Lacordaire (1859b: 417); as it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Pascoe 1866a: 473), Eucyrtus was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Eudissonomus G.S. Medvedev, 218 [M]. Type Heterophylussubstriatus Reitter, 1898, by original designation. Status: valid subgenus of Dissonomus Jacquelin du Val, 1861 in Tenebrioninae: Dissonomini.

Eudysantes Bouchard, Lawrence, Davies & Newton, 508 [M]. Type species [automatic]: Dicerodereselongatus Redtenbacher, 1868, by original designation. Status: junior synonym of Dysantes Pascoe, 1871 in Tenebrioninae: Toxicini: Dysantina. Note: unnecessary replacement name for Dysantes Pascoe, 1871 (see Bouchard and Bousquet 2020a: 101 for additional comments).

Euglyptonotus Gestro, 744 [M]. Type Euglyptonotusmagrettii Gestro, 1901, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Euhelaeus Gebien, 281 [M]. Type Euhelaeusspeculiferus Gebien, 1921, by monotypy. Status: junior synonym of Emcephalus W. Kirby, 1828 in Tenebrioninae: Heleini: Heleina. Synonymy: Matthews (1993: 1074).

Euhemicera Ando, 189, 197 [F]. Type Epilampuspulcher Hope, 1843, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Eulabis Eschscholtz, 14 [F]. Type Eulabisbicarinata Eschscholtz, 1829, by subsequent designation (Blaisdell 1932: 44). Status: valid genus in Tenebrioninae: Eulabini.

Eulea Carter, 131 [F]. Type Euleacaeca Carter, 1937, by monotypy. Status: valid genus in Lagriinae: Belopini.

Euleantus Haag-Rutenberg, 85 [M]. Type Euleantushumeralis Haag-Rutenberg, 1876, by monotypy. Status: junior synonym of Rhammatodes Haag-Rutenberg, 1876 in Pimeliinae: Tentyriini. Synonymy: Koch (1952a: 133).

Eulipus Wollaston, 448 [M]. Type Tentyriaelongata Brullé, 1839 (= Tentyriabrullaei Wollaston, 1865), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Eulytus C.O. Waterhouse, 175 [M]. Type Eulytusnodipennis C.O. Waterhouse, 1882, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Eumicrositus Viñolas, 57, 62 [M]. Type Pedinusulissiponensis Germar, 1823, by original designation. Status: valid subgenus of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina. Note: this name was first proposed by Español (1947: 11, 15) without type species designation and is therefore unavailable from that date.

Eumolpamarygmus Pic, 11 [M]. Type Eumolpamarygmusbigibbosus Pic, 1923, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Eumolparamarygmus Bremer, 8 [M]. Type Eumolpamarygmusnitidus Pic, 1935, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: the earlier usage of Eumolparamarygmus by Pic (1935: 24) is interpreted as an incorrect subsequent spelling of Eumolpamarygmus Pic, 1923 (see Bremer 2005: 210).

Eumolpocyriogeton Pic, 305 [M]. Type Eumolpocyriogetonconvexus Pic, 1923 (= Plesiophthalmusconcameratus Bremer & Lillig, 2014), by monotypy. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Eumylada Reitter, 170 [F]. Type Myladinapunctifera Reitter, 1889, by subsequent designation (Gebien 1939: 461). Status: valid genus in Blaptinae: Opatrini: Opatrina.

Eunotiodes Casey, 519 [M]. Type Eunotiodesbrevicollis Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Euomma Boheman, 101 [N]. Type Euommalaterale Boheman, 1858, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Euomophlus Iablokoff-Khnzorian, 134 [M]. Type Cistelarugosicollis Brullé, 1832, by original designation. Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini.

†Eupachypterus Kirejtshuk, Nabozhenko & Nel, 192 [M]. Type Eupachypteruseocenicus Kirejtshuk, Nabozhensko & Nel, 2010, by original designation. Status: valid genus in Blaptinae: Opatrini: Neopachypterina. Note: described from Eocene deposits (France).

Eupezoplonyx Pic, 12 [M]. Type Eupezoplonyxater Pic, 1922, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Eupezus Dejean, 211 [M]. Type Helopslongipes Fabricius, 1781, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Euphloeus Pascoe, 15 [M]. Type Euphloeusverrucosus Pascoe, 1887, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Euphron Dejean, 206 [M]. Type Helopscoerulescens Guérin-Méneville, 1831, by monotypy. Status: senior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Bousquet and Bouchard (2013a: 55). Note: nomen oblitum (see Bouchard and Bousquet 2020b: 6).

Euphrynus Fairmaire, 114 [M]. Type Euphrynusspinithorax Fairmaire, 1897, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Eupomeca Solier, 289 [F]. Type Blapscylindrica Herbst, 1799 (= Blaps obtusa Fabricius, 1798), by subsequent designation (Nabozhenko 2008: 35). Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Gemminger in Gemminger and Harold (1870: 1866).

Eupsophulus Cockerell, 242 [M]. Type species [automatic]: Eupsophuscastaneus Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Vacronini. Note: replacement name for Eupsophus Horn, 1870.

Eupsophus Horn, 344, 347 [M]. Type Eupsophuscastaneus Horn, 1870, by monotypy. Status: senior synonym of Eupsophulus Cockerell, 1906 in Pimeliinae: Vacronini. Note: junior homonym of Eupsophus Fitzinger, 1843 [Amphibia].

Eupterocoma Skopin, 152 [F]. Type Pterocomaganglbaueri Reitter, 1890, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Eurepipleura Bogdanov-Katjkov, 2 [F]. Type Anatolicaminima Bogdanov-Katjkov, 1915, by monotypy. Status: valid subgenus of Anatolica Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Eurhysodina Wasmann, 18 [F]. Type Rhyzodinamarshalli Blair, 1913, by original designation. Status: valid subgenus of Rhyzodina Chevrolat, 1873 in Tenebrioninae: Rhyssopaussini.

Euryasida Reitter, 40, 58 [F]. Type Asidabarceloi Pérez Arcas, 1868, by monotypy. Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: F. Soldati (2008: 34).

Eurycaulinus Koch, 468 [M]. Type Eurycaulusquedenfeldti Heyden, 1890, by monotypy. Status: valid subgenus of Eurycaulus Fairmaire, 1868 in Blaptinae: Opatrini: Sclerina.

Eurycaulus Fairmaire, 492 [M]. Type Eurycaulusmarmottani Fairmaire, 1868, by monotypy. Status: valid genus and subgenus in Blaptinae: Opatrini: Sclerina.

Eurychora Thunberg, 9 [F]. Type Pimeliaciliata Fabricius, 1781, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: Eurychora is an incorrect subsequent spelling of the original spelling Evrychora, first used by Thunberg (1791: 116), in prevailing usage; Eurychora is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Eurychorula Koch, 7 [F]. Type Eurychoraacuminata Fairmaire, 1891, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Eurygona Laporte, 187 [F]. Type Aulacuschilensis Gray, 1832, by monotypy. Status: senior synonym of Orthogonoderes Solier, 1841 in Pimeliinae: Praociini. Note: replacement name for Aulacus Gray, 1832; junior homonym of Eurygona Boisduval, 1836 [Lepidoptera].

Euryhelops Reitter, 209 [M]. Type Helopschampioni Reitter, 1891, by monotypy. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Euryhelops Reitter, 214 [M]. Type Euryhelopstiro Reitter, 1902, by subsequent designation (Gebien 1943: 424). Status: senior synonym of Zophohelops Reitter, 1902 in Tenebrioninae: Helopini: Cylindrinotina. Note: junior homonym of Euryhelops Reitter, 1902 [Tenebrionidae: Blaptinae: Platyscelidini].

Eurymetopon Eschscholtz, 5, 8 [N]. Type Eurymetoponrufipes Eschscholtz, 1831, by subsequent designation (Casey 1907: 288). Status: valid genus in Pimeliinae: Edrotini.

Eurymetopum Agassiz, 151 [N]. Type species [automatic]: Eurymetoponrufipes Eschscholtz, 1831, by subsequent designation (Casey 1907: 288). Status: junior synonym of Eurymetopon Eschscholtz, 1831 in Pimeliinae: Edrotini. Note: unjustified emendation of Eurymetopon Eschscholtz, 1831, not in prevailing usage.

Eurynotus W. Kirby, 418 [M]. Type Eurynotusmuricatus W. Kirby, 1819 (= Helopscapensis Fabricius, 1794), by monotypy. Status: valid genus and subgenus in Blaptinae: Platynotini: Eurynotina.

Euryostola Reitter, 202, 207 [F]. Type Pachyscelisminor Baudi di Selve, 1875, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Eurypera Pascoe, 106 [F]. Type Euryperacuprea Pascoe, 1870 (= Amarygmuspascoei Gebien, 1911), by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Blackburn (1893a: 92).

Eurypimelia Reitter, 9, 49 [F]. Type Tenebriosubglobosus Pallas, 1781, by subsequent designation (Chernei 104). Status: junior synonym of Camphonota Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Gebien (1937a: 837).

Euryprosodes Reitter, 122 [M]. Type Prosodesareolata Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Euryprosternum Chatanay, 3 [N]. Type Andremiusparallelus Chatanay, 1913, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Eusarca Chevrolat, 526 [F]. Type Eusarcairidipennis Chevrolat, 1845 (= Hegemonaresplendens Laporte, 1840), by monotypy. Status: junior synonym of Hegemona Laporte, 1840 in Stenochiinae: Cnodalonini. Synonymy: Duponchel (1845a: 498). Note: junior homonym of Eusarca Hübner, 1813 [Lepidoptera].

Eusattodes Casey, 56, 64 [M]. Type Eusattuslaevis J.L. LeConte, 1866, by original designation. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: Triplehorn (1968: 379).

Eusattus J.L. LeConte, 131 [M]. Type Eusattusdifficilis J.L. LeConte, 1851, by subsequent designation (Casey 1908: 56). Status: valid genus in Pimeliinae: Coniontini.

Euschatia Solier, 227 [F]. Type Euschatiaproxima Solier, 1851, by present designation. Status: junior synonym of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini. Synonymy: Lacordaire (1859b: 443).

Euschides J.L. LeConte, 127 [F]. Type species [automatic]: Stenomorphablapsoides Solier, 1836, by subsequent designation (Desmarest 150). Status: junior synonym of Stenomorpha Solier, 1836 in Pimeliinae: Asidini. Note: unnecessary replacement name for Stenomorpha Solier, 1836.

Euspinamarygmus Masumoto, 295 [M]. Type Euspinamarygmuskaszabi Masumoto, 1989, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Eustenia Fairmaire, 303 [F]. Type Eusteniatenuimembris Fairmaire, 1905, by monotypy. Status: senior synonym of Tucumana Gebien, 1911 in Alleculinae: Alleculini: Xystropodina. Note: junior homonym of Eustenia Snellen, 1899 [Lepidoptera].

Eustenomacidius Nabozhenko, 807 [M]. Type Helopsluridus Ménétriés, 1849 (= Stenomaxlaevicollis Kraatz, 1882), by original designation. Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina.

Eustolopus Gebien, 61 [M]. Type Eustolopuscalcaratus Gebien, 1938, by original designation. Status: valid genus in Pimeliinae: Adesmiini. Note: the First Reviser (Eustolopus Gebien, 1938 versus Entinopoda Gebien, 1938) is Penrith (1979: 40).

Eustrongylium Kolbe, 366 [N]. Type Strongyliumepiscopale Kolbe, 1895, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1911b: 590).

Eusyntelia C.O. Waterhouse, 473 [F]. Type Eusynteliabalfouri C.O. Waterhouse, 1881, by subsequent designation (R. Lucas 1920: 290). Status: valid genus in Pimeliinae: Tentyriini.

Eutagenia Reitter, 100, 125 [F]. Type Stenosissmyrnensis Solier, 1848, by subsequent designation (Iwan et al. 2020: 208). Status: valid genus in Pimeliinae: Stenosini: Stenosina. Note: Reitter (1916: 152) first noted that Stenosissmyrnensis Solier of Reitter (1886) was misidentified and proposed the new species Eutageniacribricollis for it; the earlier selection of the taxonomic species Stenosissmyrnensis Solier sensu Reitter, 1886 (= Eutageniacribricollis Reitter, 1916) by Gebien (1937a: 685) as type species is invalid since such an action would have required approval from the International Commission on Zoological Nomenclature at the time; Iwan et al. (2020: 208) fixed the nominal species Stenosissmyrnensis Solier, 1848 as the type species following Article 70.3.1 (ICZN 1999).

Euteleocera Agassiz, 152 [F]. Type species [automatic]: Euteloceraviatica Solier, 1841, by original designation. Status: junior synonym of Eutelocera Solier, 1841 in Pimeliinae: Praociini. Note: unjustified emendation of Eutelocera Solier, 1841, not in prevailing usage.

Eutelocera Solier, 209, 237 [F]. Type Euteloceraviatica Solier, 1841, by original designation. Status: valid genus in Pimeliinae: Praociini.

Eutelogonus Reitter, 24 [M]. Type Helopsdavidis Fairmaire, 1884, by monotypy. Status: valid subgenus of Entomogonus Solier, 1848 in Tenebrioninae: Helopini: Helopina.

Eutelonodolinus Robiche, 450 [M]. Type Eutelonodolinusjolyi Robiche, 2007, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Eutelonotus Fairmaire in Alluaud, 456 [M]. Type species [automatic]: Eutelusrequieni Solier, 1843, by subsequent designation (R. Lucas 1920: 291). Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Eutelus Solier, 1843 (see Bousquet 2016a: 44).

Eutelus Solier, 56 [M]. Type Eutelusrequieni Solier, 1843, by subsequent designation (R. Lucas 1920: 291). Status: senior synonym of Eutelonotus Fairmaire in Alluaud, 1902 in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Lutelus, used by Solier (1843: 4); junior homonym of Eutelus Walker, 1834 [Hymenoptera].

Eutermicola Lea, 273 [M]. Type Eutermicolasculpticollis Lea, 1916, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Eutherama Carter, 405 [N]. Type Eutheramacyaneum Carter, 1914, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Euthriptera Reitter, 203, 229 [F]. Type Thripteragrisescens Fairmaire, 1875, by monotypy. Status: valid genus in Pimeliinae: Pimeliini. Note: we act as First Revisers and reject the alternative original spelling Enthriptera, used by Reitter (1893: 203).

Euthysternum Chatanay, 508 [N]. Type Chemolanusepiscopalis Fairmaire, 1886, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Eutichus Haag-Rutenberg, 4, 58 [M]. Type Eutichuswahlbergi Haag-Rutenberg, 1875, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Eutochia J.L. LeConte, 238 [F]. Type species [automatic]: Ulomapiceum Melsheimer, 1846, by monotypy. Status: valid genus in Tenebrioninae: Ulomini. Note: replacement name for Aniara Melsheimer, 1853.

Eutomus Lacordaire, 369 [M]. Type Eutomusmicrographus Lacordaire, 1865, by subsequent designation (Barber 1914: 191). Status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: J.L. LeConte and Horn (1883: 232). Note: originally described as a member of Curculionoidea: Curculionidae: Scolytinae.

Eutoreuma Carter, 78 [N]. Type species [automatic]: Toreumacupreum Carter, 1913, by monotypy. Status: senior synonym of Atoreuma Gebien, 1941 in Tenebrioninae: Heleini: Cyphaleina. Note: replacement name for Toreuma Carter, 1913; junior homonym of Eutoreuma Grote, 1872 [Lepidoptera].

Eutrapela Dejean, 215 [F]. Type Crioceriselongata Fabricius, 1781 (= Chrysomelaunifasciata DeGeer, 1778), by subsequent designation (Duponchel 1845a: 533). Status: senior synonym of Neoeutrapela Bousquet & Bouchard, 2013 in Lagriinae: Lagriini: Statirina. Synonymy: Bousquet and Bouchard (2013a: 61). Note: junior homonym of Eutrapela Hübner, 1809 [Lepidoptera].

Eutrapelodes Borchmann, 132 [F]. Type Cteniopusgracillimus Fairmaire, 1888, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Eutriorophus Casey, 296 [M]. Type Eutriorophustuckeri Casey, 1924, by original designation. Status: junior synonym of Stibia Horn, 1870 in Pimeliinae: Edrotini. Synonymy: Blaisdell (1933: 210).

Euzadenos Koch, 286 [M]. Type Eurynotusdelalandii Mulsant & Rey, 1854, by original designation. Status: junior synonym of Selenepistoma Dejean, 1834 in Blaptinae: Dendarini: Melambiina. Synonymy: new synonym [PB]. Note: Euzadenos Koch, 1956 was recently treated as a valid subgenus of Zadenos Laporte, 1840 (Kamiński 549); however, the type species of the older available genus name Selenepistoma Dejean, 1834 is currently placed in Euzadenos and therefore Selenepistoma has priority.

Evaniosomus Guérin-Méneville, 14 [M]. Type Evaniosomusorbignianus Guérin-Méneville, 1834, by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Evaostetha Novák, 208 [F]. Type Evaostethapetri Novák, 2008, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Evelina J. Thomson, 22 [F]. Type Evelinalacordairei J. Thomson, 1860, by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Eviropodus Koch, 84 [M]. Type Trigonopusalternans Fåhraeus, 1870, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Evoplus J.L. LeConte, 128 [M]. Type Evoplusferrugineus J.L. LeConte, 1866, by monotypy. Status: junior synonym of Neomida Latreille, 1829 in Diaperinae: Diaperini: Diaperina. Synonymy: Champion (1886: 175, with Arrhenoplita W. Kirby, 1837, a junior synonym of Neomida Latreille, 1829).

Exadelium Watt, 29 [N]. Type Adeliumrufilabrum Broun, 1886, by original designation. Status: valid genus in Lagriinae: Adeliini.

Exangeltus Blackburn, 93 [M]. Type Exangeltusangustus Blackburn, 1897, by monotypy. Status: valid genus in Pimeliinae: Vacronini.

Exapinaeus Pascoe, 34 [M]. Type Exapinaeuspolitus Pascoe, 1882, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Exechophthalmus Ardoin, 168 [M]. Type Exechophthalmusguillaumeti Ardoin, 1974, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Exerestus Bates, 268 [M]. Type Exerestusjansonii Bates, 1870 (= Rhinandruselongatus Horn, 1866), by monotypy. Status: junior synonym of Rhinandrus J.L. LeConte, 1866 in Tenebrioninae: Tenebrionini. Synonymy: Bates (1872a: 98).

Exeniotis Pascoe, 353 [F]. Type Exeniotiscollaris Pascoe, 1871, by monotypy. Status: valid genus in Lagriinae: Belopini.

Exocolena Gebien, 43 [F]. Type Exocolenalongipes Gebien, 1914, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Exostira Borchmann, 353 [F]. Type Exostirasellata Borchmann, 1925, by subsequent designation (Borchmann 1930a: 512). Status: valid genus in Lagriinae: Lagriini: Statirina.

Extetranosis Koch, 741 [F]. Type Tetranosisethasicornis Koch, 1940, by monotypy. Status: valid subgenus of Microtelopsis Koch, 1940 in Pimeliinae: Stenosini: Stenosina. Note: we act as First Revisers and select Microtelopsis Koch, 1940 as the valid name for this genus instead of Extetranosis Koch, 1940 and Hypermicrotelopsis Koch, 1940.

Fahraeus Ardoin, 309, 349 [M]. Type Oplocheiruspunctatissimus Fåhraeus, 1870, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Falacer Laporte, 233 [M]. Type Accanthopuscupreus Laporte, 1840 (= Helopscontractus Palisot de Beauvois, 1811), by subsequent designation (Bousquet et al. 2018: 141). Status: junior synonym of Meracantha W. Kirby, 1837 in Tenebrioninae: Amarygmini. Synonymy: Lacordaire (1859b: 466).

Falsammidium Koch, 395, 405 [N]. Type Clitobiuslaevipennis Fairmaire, 1892, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Falsandrosus Kaszab, 307 [M]. Type Falsandrosustetrops Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: Faslandrosus was introduced earlier by Kaszab (1979a: 104) but the name is unavailable from that date since it was published after 1930 without a description, a definition or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1).

Falsaspila Koch, 28 [F]. Type Adelostomabatesi Haag-Rutenberg, 1875, by original designation. Status: junior synonym of Zarudnionymus Semenov-Tjan-Shansky & Bogatchev, 1947 in Pimeliinae: Adelostomini. Synonymy: Ardoin (1976: 149).

Falsastenochirus Pic, 13 [M]. Type Asthenochiruscontractus Fairmaire, 1894, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Falsasthenochirus Ardoin, 637 [M]. Type species [automatic]: Asthenochiruscontractus Fairmaire, 1894, by monotypy. Status: junior synonym of Falsastenochirus Pic, 1938 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Falsastenochirus Pic, 1938, not in prevailing usage.

Falsoarthroconus Kaszab, 52, 57 [M]. Type Falsoarthroconusnocturnus Kaszab, 1978, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Falsoaugolesthus Masumoto, 38 [M]. Type Eucyrtuspulcher Pic, 1927, by original designation. Status: junior synonym of Tetragonomenes Chevrolat, 1878 in Stenochiinae: Cnodalonini. Synonymy: Ando et al. (2016: 65).

Falsobates Kaszab, 5, 27 [M]. Type Falsobatesxantusi Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Falsobrachys Kulzer, 65 [M]. Type Falsobrachyslongipes Kulzer, 1954, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Falsocaedius Español, 140, 142 [M]. Type Clitobiusfossulatus Escalera, 1914, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Falsocalcar Pic, 9 [N]. Type Falsocalcarbicolor Pic, 1925, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Falsocamaria Pic, 19 [F]. Type Falsocamariaobscura Pic, 1917, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Falsocamariodes Ardoin, 159 [M]. Type Falsocamariodesviettei Ardoin, 1956, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Falsocasnonidea Pic, 31 [F]. Type Nemostiradiversipennis Pic, 1923, by monotypy. Status: junior synonym of Casnonidea Fairmaire, 1882 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1936: 393).

Falsocatomulus Pic, 10 [M]. Type Falsocatomuluseuphraticus Pic, 1914, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Falsocosmonota Kaszab, 75 [F]. Type Falsocosmonotacheni Kaszab, 1962, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Falsocossyphus Pic, 4 [M]. Type Falsocossyphuspilosus Pic, 1916, by monotypy. Status: valid genus in Tenebrioninae: Falsocossyphini.

Falsocuphotes Pic, 22 [F]. Type Falsocuphotescurticornis Pic, 1918, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Falsodiopethes Pic, 12 [M]. Type Falsodiopethesgounellei Pic, 1924, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Falsoencyalesthus Pic, 29 [M]. Type Falsoencyalesthuslatipennis Pic, 1923, by monotypy. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1969b: 125, with Encyalesthus Motschulsky, 1860, a junior synonym of Derosphaerus J. Thomson, 1858).

Falsogauromaia Pic, 22 [F]. Type Falsogauromaiaannulipes Pic, 1921, by monotypy. Status: valid subgenus of Gauromaia Pascoe, 1866 in Stenochiinae: Cnodalonini.

Falsolagria Pic, 44 [F]. Type Falsolagriamarmorata Pic, 1927, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Falsolobodera Kaszab, 24 [F]. Type Falsoloboderaskopini Kaszab, 1967, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Falsolophocnemis Pic, 13 [F]. Type Falsolophocnemissinuatipes Pic, 1917, by subsequent designation (Gebien 1948: 542). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Kaszab (1977b: 29).

Falsomicrodera Kaszab, 291, 294 [F]. Type Microderaturkestanica Schuster, 1915, by original designation. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Falsomophlus Pic, 11 [M]. Type Falsomophlusniger Pic, 1925, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Falsomycterus Pic, 127 [M]. Type Falsomycterusdiversipes Pic, 1907, by monotypy. Status: valid genus in Pimeliinae: Falsomycterini.

Falsonannocerus Pic, 150 [M]. Type Falsonannocerusdentaticeps Pic, 1947, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Falsonemostira Pic, 14 [F]. Type Nemostiraannulipes Pic, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Falsonotostrongylium Kaszab, 538, 552 [N]. Type Falsonotostrongyliumbradymeroides Kaszab, 1955, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Falsoperichilus Pic, 16 [M]. Type Falsoperichilussemirugosus Pic, 1920, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Falsophthora Kaszab, 306, 309 [F]. Type Tagalusfordi Kulzer, 1957, by monotypy. Status: valid subgenus of Pseudophthora Kaszab, 1970 in Phrenapatinae: Penetini.

Falsoplonyx Ardoin, 308, 337 [M]. Type Gonocnemisrubripes Fairmaire, 1899, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Falsopraocis Kulzer, 5, 102 [M]. Type Amphidoraricardae (as “richardae”) Solier, 1851, by original designation. Status: valid genus in Pimeliinae: Praociini.

Falsopsilonycha Pic, 303 [F]. Type Psilonychausambarana Pic, 1917, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Falsostrongylium Pic, 11 [N]. Type Falsostrongyliumsemirufum Pic, 1915, by subsequent designation (Gebien 1948: 539). Status: valid genus in Stenochiinae: Stenochiini.

Falsosynopticus Pic, 17 [M]. Type Falsosynopticusater Pic, 1936, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Falsotagalus Kaszab, 301, 310 [M]. Type Falsotagalussubcoecus Kaszab, 1977, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Falsotithassa Pic, 18 [F]. Type Falsotithassasumatrana Pic, 1934, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Falsozialeus Pic, 11 [M]. Type Falsozialeusater Pic, 1951 (= Asyleptusfumosus Péringuey, 1896), by monotypy. Status: junior synonym of Asyleptus Péringuey, 1896 in Tenebrioninae: Amarygmini. Synonymy: Ardoin (1962b: 957); Schawaller and Bremer (2013: 81).

Falsozotypus Kaszab, 334 [M]. Type Falsozotypusopacipennis Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: this name was first published by Kaszab (1979a: 108) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article (13.1) and is therefore unavailable from that date.

Farsarthrosis Kaszab, 86 [F]. Type Farsarthrosisbenardi Kaszab, 1979, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Faustia Kraatz, 92 [F]. Type Faustiamodesta Kraatz, 1882, by monotypy. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Ferganoprosodes G.S. Medvedev, 595 [M]. Type Prosodesangulicollis Kraatz, 1883, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Ferveoventer Smith, 604 [M]. Type Ferveoventerbrowni Smith, 2013, by original designation. Status: valid genus in Pimeliinae: Asidini.

Fifina Novák, 470 [F]. Type Fifinaromani Novák, 2018, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Fifinoides Novák, 78 [M]. Type Fifinoideschinensis Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Filotarsus Solier, 209, 239 [M]. Type Filotarsustenuicornis Solier, 1841, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini.

Fitzsimonsia Koch, 415 [F]. Type Fitzsimonsiacymbium Koch, 1955, by original designation. Status: senior synonym of Fitzsimonsium Koch, 1962 in Pimeliinae: Stenosini: Platamodina. Note: junior homonym of Fitzsimonsia Witte, 1943 [Reptilia].

Fitzsimonsium Koch, 152 [N]. Type species [automatic]: Fitzsimonsiacymbium Koch, 1955, by original designation. Status: valid genus in Pimeliinae: Stenosini: Platamodina. Note: replacement name for Fitzsimonsia Koch, 1955.

Flabellalogista Pic, 256 [F]. Type Flabellalogistaminuta Pic, 1954, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Flabellolagria Pic, 27 [F]. Type Flabellolagrialuteovittata Pic, 1927, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Flabellostrongylium Pic, 18 [N]. Type Flabellostrongyliumatronitidum Pic, 1938, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Flavipoda Campbell, 21 [F]. Type Helopsflavipes Fabricius, 1792 [as “Alleculaflavipes Jacquelin du Val, 1857”], by original designation. Status: valid subgenus of Lobopoda Solier, 1835 in Alleculinae: Alleculini: Alleculina.

Foleya Peyerimhoff, 71 [F]. Type Foleyabrevicornis Peyerimhoff, 1916, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Foochounus Pic, 22 [M]. Type Foochounusconvexipennis Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Foranotum Nabozhenko & Sadeghi, 165 [N]. Type Foranotumperforatum Nabozhenko & Sadeghi, 2017, by original designation. Status: valid genus in Kuhitangiinae: Foranotini.

Fossilochile Koch, 63 [F]. Type Fossilochilerufa Koch, 1952, by original designation. Status: junior synonym of Pachynotelus Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Synonymy: Penrith and Endrödy-Younga (1994: 54).

Fourtaus Pic in Alfieri, 47 [M]. Type Fourtausbrevicornis Pic, 1921 (= Hegeterocaraarabicum Reitter, 1900), by monotypy. Status: junior synonym of Hegeterocara Reitter, 1900 in Pimeliinae: Tentyriini. Synonymy: Koch (1941: 294).

Foveostatira Pic, 24 [F]. Type Statirafoveicollis Pic, 1918, by subsequent designation (Borchmann 1936: 244). Status: valid subgenus of Statira Lepeletier & Audinet-Serville, 1828 in Lagriinae: Lagriini: Statirina.

Freudeia Kaszab, 216 [F]. Type Freudeianepalica Kaszab, 1961, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Freudella Ardoin, 30 [F]. Type Freudellaauripunctata Ardoin, 1961, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Freyitia Koch, 522, 525 [F]. Type Freyitiaornatipes Koch, 1943, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Freyula Koch, 591, 593 [F]. Type Freyulapsammarina Koch, 1959, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Fundulus Gistel, 125 [M]. Type species [automatic]: Opatrumtibiale Fabricius, 1781, by monotypy. Status: junior synonym of Melanimon Steven, 1828 in Tenebrioninae: Melanimonini. Note: unnecessary replacement name for Microzoum Dejean, 1834; junior homonym of Fundulus Lacepède, 1803 [Pisces].

Gabonia Fairmaire, 326 [F]. Type Gaboniadenticulata Fairmaire, 1894, by monotypy. Status: senior synonym of Gabonisca Fairmaire, 1894 in Lagriinae: Pycnocerini. Note: junior homonym of Gabonia Jacoby, 1893 [Coleoptera: Chrysomelidae].

Gabonisca Fairmaire, 395 [F]. Type species [automatic]: Gaboniadenticulata Fairmaire, 1894, by monotypy. Status: valid genus in Lagriinae: Pycnocerini. Note: replacement name for Gabonia Fairmaire, 1894.

Gahanosis Penrith, 371, 379 [F]. Type Zophosisundulata Gahan, 1900, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Gamaxus Bates, 315 [M]. Type Gamaxushauxwellii Bates, 1868, by monotypy. Status: valid genus in Lagriinae: Goniaderini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Garambanus Ardoin, 847 [M]. Type Garambanusverschureni Ardoin, 1964, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Gargilius Fairmaire, 251 [M]. Type Gargiliustrispinosus Fairmaire, 1891, by subsequent designation (Gebien 1940: 421). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Garridoa Marcuzzi, 180 [F]. Type Garridoakaszabi Marcuzzi, 1985, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Gastrhaema Jacquelin du Val, 353 [N]. Type Cistelarufiventris Waltl, 1835, by subsequent designation (Novák and Pettersson 2008: 331). Status: valid genus in Alleculinae: Cteniopodini.

Gaurobates Gebien, 170, 184 [M]. Type Gaurobatespictus Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Gauromaia Pascoe, 473 [F]. Type Gauromaiadives Pascoe, 1866, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Gebienella Kaszab, 4, 21 [F]. Type Gebienellainterrumpens Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Gebienia Borchmann, 219, 352 [F]. Type Gebieniarimulosa Borchmann, 1921, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Gebieniella Koch, 736 [F]. Type Ethasstenosides Pascoe, 1862, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Gebienocamaria Masumoto, 223, 232 [F]. Type Camariaangulicollis Fairmaire, 1896, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Gebleria Motschulsky, 410 [F]. Type Dilaphilacoides Fischer von Waldheim, 1844, by monotypy. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Gedeon Reiche & Saulcy, 219 [M]. Type Gedeonhierichonticus Reiche & Saulcy, 1857, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Gedrosia Bogatchev, 111 [F]. Type Gedrosiamonstrosa Bogatchev, 1961 (= Pseudopodhomalagabrieli Schuster, 1938), by original designation. Status: junior synonym of Pseudopodhomala Schuster, 1938 in Pimeliinae: Pimeliini. Synonymy: Löbl et al. (2008b: 166). Note: junior homonym of Gedrosia Stål, 1862 [Hemiptera].

Genateropa Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Apterogenacanonnei Ardoin, 1962, by original designation. Status: valid genus in Stenochiinae: Stenochiini. Note: replacement name for Apterogena Ardoin, 1962.

Genoblaps Bauer, 230 [F]. Type Blaps tentyroides Seidlitz, 1893 (= Blapssocia Seidlitz, 1893), by monotypy. Status: valid subgenus of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina.

Gentinadis Laporte, 240 [M]. Type Stenochiacaerulea Laporte, 1840 (= Strongyliumazureum Germar, 1823), by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Lacordaire (1859b: 484).

Geoborus Blanchard, pl. 13 [M]. Type Geoboruscostatus Blanchard, 1842 (= Gyriosomuslineatus Guérin-Méneville, 1834), by subsequent designation (Pizarro-Araya and Flores 2006: 95). Status: valid genus in Pimeliinae: Epitragini.

Geophanus Haag-Rutenberg, 4, 46 [M]. Type Psaryphisconfusa Fåhraeus, 1870, by subsequent designation (Gebien 1937a: 675). Status: valid genus in Pimeliinae: Adelostomini.

Gerandryus Rottenberg, 217 [M]. Type species [automatic]: Parablopsaetnensis Rottenberg, 1871, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: replacement name for Parablops Rottenberg, 1871.

Gerardia Pic, 258 [F]. Type Gerardiasublineata Pic, 1954, by monotypy. Status: senior synonym of Piccula Bousquet & Bouchard, 2015 in Alleculinae: Alleculini: Gonoderina. Note: junior homonym of Gerardia Lacaze-Duthiers, 1864 [Cnidaria].

Gerdacula Novák, 145 [F]. Type Gerdaculafujianica Novák, 2015, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Ghaleca Péringuey, 316 [F]. Type Ghalecalaeta Péringuey, 1899 (= Alymonprolatus Pascoe, 1866), by monotypy. Status: junior synonym of Alymon Pascoe, 1866 in Tenebrioninae: Amarygmini. Synonymy: Péringuey (1904: 297, through synonymy of its type species with Alymonprolatus Pascoe, 1866).

Gibbostrongylium Pic, 18 [N]. Type Strongyliummedanense Pic, 1917, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Gigantopigeus Kaszab, 355, 374 [M]. Type Gigantopigeusmirabilis Kaszab, 1984, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Girardius L. Soldati, 73 [M]. Type Micipsapersica Baudi, 1874, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Girardocamaria Masumoto, 227, 232 [F]. Type Girardocamariaardoini Masumoto, 1993, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Glabrasida Escalera, 408 [F]. Type Glabrasidaconspuata Escalera, 1910, by subsequent designation (F. Soldati 2008: 32). Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini.

Glabrilobopoda Campbell, 46 [F]. Type Lobopodaglabrata Champion, 1888, by original designation. Status: valid subgenus of Lobopoda Solier, 1835 in Alleculinae: Alleculini: Alleculina.

Globasida Escalera, 430 [F]. Type Asidaoblonga Rambur, 1838, by subsequent designation (Viñolas and Cartagena 2005: 209). Status: valid subgenus of Asida Latreille, 1802 in Pimeliinae: Asidini.

Globularthrodosis Kaszab, 95 [F]. Type Diaphanidussemenowi Reitter, 1900, by original designation. Status: junior synonym of Diaphanidus Reitter, 1900 in Pimeliinae: Erodiini. Synonymy: G.S. Medvedev and Nepesova (1985: 52).

Glyptariobius Koch, 423 [M]. Type Hoplariobiusexacavatus Koch, 1948, by original designation. Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Glyptasida Casey, 75, 95 [F]. Type Ophryastessordidus J.L. LeConte, 1853, by original designation. Status: valid subgenus of Philolithus Lacordaire, 1858 in Pimeliinae: Asidini.

Glyptophrynus Fairmaire, 532 [M]. Type Glyptophrynustenuesculptus Fairmaire, 1899, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Glyptopteryx Gebien, 376 [F]. Type Glyptopteryxforticostis Gebien, 1910 (= Selinusquadricollis Fairmaire, 1887), by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Glyptothorax Borchmann, 223 [M]. Type Glyptothoraxpilosus Borchmann, 1937, by original designation. Status: senior synonym of Borchmannius Bousquet & Bouchard, 2015 in Alleculinae: incertae sedis. Note: junior homonym of Glyptothorax Blyth, 1860 [Pisces]; taxon also described as new by Borchmann (1938: 122).

Glyptotus J.L. LeConte, 75 [M]. Type Glyptotuscribratus J.L. LeConte, 1858, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Gnaptor Brullé, 254 [M]. Type Tenebriospinimanus Pallas, 1781, by monotypy. Status: valid genus and subgenus in Blaptinae: Blaptini: Gnaptorina. Note: this name was previously attributed to Brullé (1832: 202 as Petrobius, corrected to Gnaptor in the “Errata” of the same work) in the literature.

Gnaptorina Reitter, 364 [F]. Type Gnaptorinafelicitana Reitter, 1887, by monotypy. Status: valid genus and subgenus in Blaptinae: Blaptini: Gnaptorinina.

Gnathelops Gebien, 320 [M]. Type Gnathelopschatanayi Gebien, 1922, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: removed from the tribe Helopini and placed as Tenebrionidae incertae sedis by Nabozhenko (2018: 183).

Gnathidium Gebien, 41 [N]. Type Gnathidiumcephalotes Gebien, 1921, by monotypy. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Gnathocerus Agassiz, 164 [M]. Type species [automatic]: Gnatocerusruber Thunberg, 1814 (= Trogossitacornuta Fabricius, 1798), by monotypy (see ICZN 1975, Opinion 1039). Status: junior synonym of Gnatocerus Thunberg, 1814 in Diaperinae: Diaperini: Adelinina. Note: unjustified emendation of Gnatocerus Thunberg, 1814, not in prevailing usage.

Gnathosia Fischer, 13 [F]. Type Gnathosiaglabra Fischer, 1821, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Gnatocerus Thunberg, 47 [M]. Type Gnatocerusruber Thunberg, 1814 (= Trogossitacornuta Fabricius, 1798), by monotypy (see ICZN 1975, Opinion 1039). Status: valid genus and subgenus in Diaperinae: Diaperini: Adelinina. Note: placed on the Official List of Generic Names in Zoology by the ICZN (1975, Opinion 1039).

Gnesis Pascoe, 477 [M]. Type Gnesishelopioides Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Gnophota Erichson, 237 [F]. Type Gnophotaantracina Erichson, 1843, by subsequent designation (R. Lucas 1920: 306). Status: valid genus in Pimeliinae: Tentyriini.

Goajiria Ivie & Hart, 474 [F]. Type Diastolinuscurtus Mulsant & Rey, 1859, by original designation. Status: valid subgenus of Diastolinus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Blapstinina. Note: name first introduced by Marcuzzi (1986: 180) without a type species designation.

Gobretus Freude, 148, 152 [M]. Type Epitraguscephalotes Freude, 1967, by monotypy. Status: valid subgenus of Epitragus Latreille, 1802 in Pimeliinae: Epitragini.

Gonasida Casey, 75, 117 [F]. Type Pelecyphoruselatus J.L. LeConte, 1853, by original designation. Status: valid subgenus of Philolithus Lacordaire, 1858 in Pimeliinae: Asidini.

Gondvanadelium Kaszab, 81 [N]. Type Gondvanadeliumseirotranoides Kaszab, 1981, by original designation. Status: valid genus in Lagriinae: Adeliini.

Gondwanocrypticus Español, 10 [M]. Type Crypticusplatensis Fairmaire, 1884, by original designation. Status: valid genus in Diaperinae: Crypticini. Note: this name was introduced earlier by Koch (1950c: 64) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Gondwanodilamus Kaszab, 320 [M]. Type Conibiusfranzi Kaszab, 1969, by original designation. Status: junior synonym of Conibius J.L. LeConte, 1851 in Blaptinae: Opatrini: Blapstinina. Synonymy: new synonym [RLA]. Note: the new synonymy is based on taxonomic studies of all known species in the genus Conibius J.L. LeConte, 1851 and follows the previous synonymization of the two other subgenera previously treated as valid, Ooconibius Casey, 1895 and Euconibius Casey, 1895, by Aalbu in Bousquet et al. (2018: 202).

Gonespites Gebien, 325, 396 [M]. Type Gonespitessubcrenatus Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Goniadera Perty, 62 [F]. Type Goniaderacrenata Perty, 1832, by monotypy. Status: valid genus and subgenus in Lagriinae: Goniaderini. Note: the alternative original spelling Gonyodera, used by Perty (1832: 63), was rejected by Perty (1833: 14) who acted as the First Reviser.

†Gonialaena Nabozhenko, Bukejs & Telnov, 254 [F]. Type Gonialaenagroehni Nabozhenko, Bukejs & Telnov, 2019, by original designation. Status: valid genus in Lagriinae: Gonialaenini. Note: described from Eocene Baltic amber.

Goniodera Agassiz, 165 [F]. Type species [automatic]: Goniaderacrenata Perty, 1832, by monotypy. Status: junior synonym of Goniadera Perty, 1832 in Lagriinae: Goniaderini. Note: unjustified emendation of Goniadera Perty, 1832, not in prevailing usage.

Gonocephalum Solier, 498 [N]. Type Opatrumfuscum Herbst, 1793 (= Opatrumrusticum G.-A. Olivier, 1812), by subsequent designation (Gebien 1939: 443). Status: valid genus and subgenus in Blaptinae: Opatrini: Opatrina.

Gonocnemis J. Thomson, 101 [F]. Type Gonocnemisstrigipennis J. Thomson, 1858, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Gonocnemocistela Pic, 22 [F]. Type Gonocnemocistelalutea Pic, 1935, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Gonodera Mulsant, 41 [F]. Type Cistelafulvipes Fabricius, 1792 (= Cistelaluperus Herbst, 1783), by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Gonogenius Solier, 8, 48 [M]. Type Scotobiusvulgaris Guérin-Méneville, 1834, by original designation. Status: junior synonym of Scotobius Germar, 1823 in Tenebrioninae: Scotobiini. Synonymy: Lacordaire (1859a: 129).

Gonopterus Solier, 101 [M]. Type Sepidiumrugosum Fabricius, 1781, by monotypy. Status: junior synonym of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Gemminger in Gemminger and Harold (1870: 1900).

Gonopus Latreille, 580 [M]. Type Blapstibialis Fabricius, 1798, by monotypy. Status: valid genus and subgenus in Blaptinae: Platynotini: Platynotina.

Gonospa Champion, 216 [F]. Type Gonospaphaedonoides Champion, 1886, by subsequent designation (Gebien 1940: 426). Status: valid genus in Stenochiinae: Cnodalonini.

Grabulax Kanda, 558 [M]. Type Grabulaxdarlingtoni Kanda, 2016, by original designation. Status: valid genus in Lagriinae: Laenini.

Gracilasida Escalera, 445 [F]. Type Asidaariasi Escalera, 1909, by subsequent designation (F. Soldati 2008: 32). Status: valid subgenus of Asida Latreille, 1802 in Pimeliinae: Asidini. Note: this genus was established without included nominal species; Asidaariasi Escalera, 1909 and A.pusillima Kraatz, 1874 were first subsequently and expressly included in Gracilasida by Escalera (1909: 135–136).

Graecopachys Skopin, 99 [M]. Type Tentyriaquadricollis Brullé, 1832, by original designation. Status: junior synonym of Phymatiotris Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Löbl et al. (2008a: 43).

Grammicus G.R. Waterhouse, 323 [M]. Type Grammicuschilensis G.R. Waterhouse, 1845, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Granasida Reitter, 9, 14 [F]. Type Asidagranulifera Chevrolat, 1840, by monotypy. Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini.

Grandelagria Pic, 3 [F]. Type Grandelagriabicoloripes Pic, 1940, by monotypy. Status: valid subgenus of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina.

Grandicyrtomius Freude, 225 [M]. Type Epitragusgrandis Champion, 1884, by original designation. Status: valid subgenus of Cyrtomius Casey, 1907 in Pimeliinae: Epitragini.

Granulasida Escalera, 427 [F]. Type Asidasetipennis Allard, 1869, by subsequent designation (Viñolas and Cartagena 2005: 192). Status: junior synonym of Gracilasida Escalera, 1905 in Pimeliinae: Asidini. Synonymy: F. Soldati (2008: 34, with Planasida Escalera, 1907, a junior synonym of Gracilasida Escalera, 1905).

Granulophanes Nabozhenko, 2 [M]. Type Hedyphaneslutosus Allard, 1877, by original designation. Status: valid subgenus of Hedyphanes Fischer, 1820 in Tenebrioninae: Helopini: Helopina.

Graptopezus Gebien, 296 [M]. Type Seteniscostipennis Blair, 1915 (= Nyctozoiluscrenaticollis W.J. MacLeay, 1886), by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Gressittiola Kaszab, 462, 464 [F]. Type Gressittiolaplatydemoides Kaszab, 1955, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Gridellia Kammerer, 270 [F]. Type species [automatic]: Tenebrioclypealis Gebien, 1920, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini. Note: replacement name for Villiersia Gridelli, 1951.

Gridelliopus Koch, 358 [M]. Type Gridelliopussubsquamosus Koch, 1956, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Gronophora Borchmann, 48, 103 [F]. Type Gronophoragravida Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Guanobius Grimm, 375 [M]. Type Guanobiusborneensis Grimm, 2008, by original designation. Status: valid genus in Tenebrioninae: Alphitobiini.

Guildia Antoine, 346, 359 [F]. Type Microsituspunctistriatus Escalera, 1925, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Gunarellus Reitter, 22 [M]. Type Helopsgratus Frivaldszky, 1894, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Stenohelops Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko et al. (2020b: 297). Note: Gunarellus was first described by Reitter (1922a: 22, in key; issued 30 March 1922) without originally included species; Reitter (1922b: 114–115; issued 25 October 1922) was the first author to subsequently and expressly include nominal species in Gunarellus (ICZN 1999, Article 67.2.2).

Gunarus Gozis, 25 [M]. Type Helopshirtulus Reiche, 1862, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Gymnetasida Reitter, 10, 22 [F]. Type Asidatricostata Allard, 1869, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Gymnognathus Solier, 136 [M]. Type Gymnognathusfuscus Solier, 1851, by monotypy. Status: junior synonym of Arthroconus Solier, 1851 in Pimeliinae: Edrotini. Synonymy: Lacordaire (1859a: 67). Note: junior homonym of Gymnognathus Schönherr, 1823 [Coleoptera: Anthribidae].

Gynandrocera Gebien, 46 [F]. Type Gynandroceracephalotes Gebien, 1920 (= Afrinusminor Péringuey, 1908), by subsequent designation (Koch 1950b: 328). Status: valid subgenus of Afrinus Fairmaire, 1888 in Pimeliinae: Tentyriini.

Gyrasida Koch, 127, 143 [F]. Type Afrasidapropensa Wilke, 1922, by original designation. Status: valid genus in Pimeliinae: Praociini. Note: elevated to the rank of genus and transferred from Pimeliinae: Asidini by Flores and Vidal (2009: 52); as mentioned by Flores and Vidal (2009: 48, 56) the type species was originally described from South Africa in the genus Gyrasida Koch, 1962 is endemic to Chile.

Gyrinodes Fauvel, 61 [M]. Type Helopsgagatinus Küster, 1850, by monotypy. Status: junior synonym of Nesotes Allard, 1876 in Tenebrioninae: Helopini: Helopina. Synonymy: new synonym [YB]. Note: this genus-group taxon has been forgotten in the literature; its type species is currently placed in the nominotypical subgenus of Nesotes Allard, 1876 and for that reason Fauvel’s name is regarded as a new junior synonym of Nesotes.

Gyriosomus Guérin-Méneville, 6 [M]. Type Nyctelialuczotii Guérin-Méneville, 1831, by subsequent designation (Duponchel 1845b: 449). Status: valid genus in Pimeliinae: Nycteliini.

Gyrosis Gebien, 33, 37 [F]. Type Zophosisorbicularis Deyrolle, 1867, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Habrobates Semenov, 11 [M]. Type Habrobatesvernalis Semenov, 1903, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Habrochiton Semenov-Tjan-Shansky, 177, 179 [M]. Type Habrochitonvernus Semenov-Tjan-Shansky, 1907, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Hades J. Thomson, 13 [M]. Type Hadestenebrosus J. Thomson, 1860 (= Platydemahemisphaerica Laporte & Brullé, 1831), by monotypy. Status: senior synonym of Pimplema Pascoe, 1887 in Diaperinae: Leiochrinini. Synonymy: Gebien (1940: 755). Note: junior homonym of Hades Westwood, 1851 [Lepidoptera].

Hadroderus Koch, 347 [M]. Type Hadroderustuberculiferus Koch, 1956, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Hadrodes Wollaston, 226 [M]. Type Hadrodeshelenensis Wollaston, 1877, by monotypy. Status: valid genus in Blaptinae: Opatrini: AMMOBIINA.

Hadromelambius Koch, 432 [M]. Type Melambiustelueticus Escalera, 1914, by original designation. Status: valid subgenus of Melambius Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Hadrophasis Ferrer, 87 [F]. Type Hadrophasisangolensis Ferrer, 1992, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Hadrus Wollaston, 502 [M]. Type Hadrusalpinus Wollaston, 1854, by subsequent designation (R. Lucas 1920: 313). Status: senior synonym of Wolladrus Iwan & Kamiński, 2016 in Blaptinae: Opatrini: Opatrina. Note: junior homonym of Hadrus Perty, 1833 [Diptera].

Haemerophygus Baudi di Selve, 266 [M]. Type Helopsmucoreus Waltl, 1838, by monotypy. Status: junior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Synonymy: Nabozhenko et al. (2016b: 608).

Haemodus Gebien, 904 [M]. Type species [automatic]: Haemuscarinatipennis Péringuey, 1904, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina. Note: replacement name for Haemus Péringuey, 1904.

Haemus Péringuey, 228 [M]. Type Haemuscarinatipennis Péringuey, 1904, by monotypy. Status: senior synonym of Haemodus Gebien, 1943 in Blaptinae: Dendarini: Melambiina. Note: junior homonym of Haemus Stål, 1862 [Hemiptera].

Halammobia Semenov, 92 [F]. Type Tenebriopellucidus Herbst, 1799, by monotypy. Status: valid genus in Diaperinae: Phaleriini.

Halonomus Wollaston, 201 [M]. Type Halonomusgrayii Wollaston, 1861 (= Opatrumovatum Erichson, 1843), by subsequent designation (Iwan and Löbl 2007: 734). Status: junior synonym of Clitobius Mulsant & Rey, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Fairmaire (1886b: ccv). Note: the original combination of the accepted name of the type species, Opatrumovatum Erichson, 1843, is a junior primary homonym of Opatrumovatum Fabricius, 1801.

Halophalerus Crotch, 107 [M]. Type Phaleriarotundata J.L. LeConte, 1851, by subsequent designation (Bousquet et al. 2018: 289). Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Synonymy: Austin (1880: 38).

Hangaya Matthews & Merkl, 479 [F]. Type Hangayaenigmatica Matthews & Merkl, 2015, by original designation. Status: valid genus in Tenebrioninae: incertae sedis. Note: according to Matthews and Lawrence (2019: 628) the closest relative of this genus within the subfamily Tenebrioninae remains to be determined.

Hanstroemium Koch, 19 [N]. Type Hanstroemiumadelostomoide Koch, 1953, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Haplandrus J.L. LeConte, 230 [M]. Type Helopsfemoratus Fabricius, 1798 (= Upisfulvipes Herbst, 1797), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Haporema Fairmaire, 109 [F]. Type Haporemadecipiens Fairmaire, 1892, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hapsida Gemminger in Gemminger and Harold, 1955 [F]. Type species [automatic]: Apsidachrysomelina Lacordaire, 1859, by original designation. Status: junior synonym of Apsida Lacordaire, 1859 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Apsida Lacordaire, 1859, not in prevailing usage.

Harvengia Ferrer, 367 [F]. Type Harvengiavietnamita Ferrer, 2004, by original designation. Status: valid genus in Pimeliinae: Stenosini: Harvengiina.

Hasticollinum Kaszab, 96 [N]. Type Hasticollinumpodagrarium Kaszab, 1939, by original designation. Status: junior synonym of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Ferrer (1995a: 72).

Havanalia Novák, 489 [F]. Type Havanaliaqazvinica Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Hectus Pascoe, 288, 289 [M]. Type Hectusanthracinus Pascoe, 1869, by monotypy. Status: junior synonym of Olisthaena Erichson, 1842 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1913a: 62).

Hedrotes Gemminger in Gemminger and Harold, 1816 [M]. Type species [automatic]: Edrotesventricosus J.L. LeConte, 1851, by monotypy. Status: junior synonym of Edrotes J.L. LeConte, 1851 in Pimeliinae: Edrotini. Note: unjustified emendation of Edrotes J.L. LeConte, 1851, not in prevailing usage.

Hedyphanes Fischer, pl.15 [M]. Type Hedyphanescoerulescens Fischer, 1820, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina.

Hegemona Laporte, 230 [F]. Type Hegemonaresplendens Laporte, 1840, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hegeter Latreille, 172 [M]. Type Hegeterstriatus Latreille, 1804 (= Blapstristis Fabricius, 1792), by subsequent monotypy (Latreille 1804: 276). Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: originally proposed without included nominal species; Latreille (1804: 276), by including the species Hegeterstriatus Latreille, 1804 in association with this name, was the first author to subsequently and expressly include nominal species in Hegeter (ICZN 1999, Article 67.2.2); the previously accepted type species for this genus was Blapselongata G.-A. Olivier, 1795 (= Blapstristis Fabricius, 1792), by subsequent designation (Latreille 1810: 429); Latreille (1804: 276) mentioned “Le type de ce genre paroît être le blaps alongé d’Olivier” however he did not use a Latin name for the species.

Hegeterocara Reitter, 94, 190 [N]. Type Hegeterocaraarabicum Reitter, 1900, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Hegeteromorpha Escalera, 38 [F]. Type Pachychilaexternecostata Haag-Rutenberg, 1875, by subsequent monotypy (Escalera 1914: 281). Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: originally proposed without included nominal species; Escalera (1914: 281), by including the species Pachychilaexternecostata Haag-Rutenberg, 1875 in association with this name, was the first author to subsequently and expressly include nominal species in Hegeteromorpha (ICZN 1999, Article 67.2.2).

Heinrichesia Carl, 258 [F]. Type Heinrichesiaschaeferi Carl, 2000, by monotypy. Status: junior synonym of Waterhousia Skopin, 1973 in Pimeliinae: Pimeliini. Synonymy: G.S. Medvedev (2005a: 309). Note: combined description of new genus-group taxon and new species (ICZN 1999, Article 13.4).

Helea Latreille, 326 [F]. Type Heleaperforata Latreille, 1817, by subsequent monotypy (Latreille 1817: 261). Status: valid genus in Tenebrioninae: Heleini: Heleina. Note: the name Helea Meigen, 1800 (Diptera) was suppressed for the purposes of zoological nomenclature by the ICZN (1963, Opinion 678); originally proposed without included nominal species; Latreille (1817: 261), by including the new species Heleaperforata Latreille, 1817 [as Heleusperforatus] in association with this name, was the first author to subsequently and expressly include nominal species in Helea (ICZN 1999, Article 67.2.2).

Heledona Agassiz, 135, 174 [F]. Type species [automatic]: Opatrumagricola Herbst, 1783, by subsequent monotypy (Latreille 1802: 162). Status: junior synonym of Eledona Latreille, 1797 in Tenebrioninae: Bolitophagini. Note: unjustified emendation of Eledona Latreille, 1797, not in prevailing usage.

Helenomelas Ardoin, 190 [M]. Type Helenomelasbasilewskyi Ardoin, 1972, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Helenophorus Gemminger in Gemminger and Harold, 1851 [M]. Type species [automatic]: Tenebriocollaris Linnaeus, 1767, by monotypy. Status: junior synonym of Leptoderis Billberg, 1820 in Pimeliinae: Elenophorini: Elenophorina. Note: unjustified emendation of Elenophorus Dejean, 1821, not in prevailing usage.

Helibatus Mulsant & Rey, 94, 100 [M]. Type Helibatusmorio Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Helioarthrodibius Koch, 353 [M]. Type Arthrodeisruguliventris Fairmaire, 1884, by monotypy. Status: valid subgenus of Arthrodibius Lesne, 1915 in Pimeliinae: Erodiini.

Heliocaes Bedel, 92 [M]. Type Blaps emarginata Fabricius, 1792, by present designation. Status: junior synonym of Heliopates Dejean, 1834 in Blaptinae: Dendarini: Dendarina. Synonymy: Gebien (1938a: 304). Note: this genus was proposed as a replacement name for “Heliopathes” Dejean sensu Mulsant, 1854, an incorrect subsequent spelling of Heliopates Dejean, 1834, which is unavailable (ICZN 1999, Article 33.3); Heliocaes Bedel, 1906 was made available by indication (ICZN 1999, Article 12.2.1); Escalera (1914: 326) was the first author to subsequently and expressly include nominal species in Heliocaes (ICZN 1999, Article 67.2.2).

Heliocrates Reitter, 98 [M]. Type Heliophilushumerangulus Reitter, 1904, by subsequent designation (Iwan and Löbl 2007: 734). Status: valid subgenus of Heliopates Dejean, 1834 in Blaptinae: Dendarini: Dendarina.

Heliodromus Brullé, 196 [M]. Type Heliodromusrotundatus Brullé, 1832, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Tentyria Latreille, 1802 in Pimeliinae: Tentyriini. Synonymy: Solier (1835b: 314).

Heliofugus Guérin-Méneville, pl. 4 [M]. Type Heliofugusarenosus Guérin-Méneville, 1831, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Heliomelasma Koch, 408 [N]. Type Melasmanaappenhageni Koch, 1948, by original designation. Status: valid subgenus of Melasmana Strand, 1935 in Blaptinae: Dendarini: Melambiina.

Heliomophlus Reitter, 147 [M]. Type Heliotaurusscabriusculus Fairmaire, 1866, by subsequent designation (R. Lucas 1920: 319). Status: valid genus in Alleculinae: Cteniopodini.

Heliopates Dejean, 191 [M]. Type Tenebriolusitanicus Herbst, 1797, by subsequent designation (Gebien 1938a: 304). Status: valid genus and subgenus in Blaptinae: Dendarini: Dendarina. Note: this genus was originally proposed as a replacement name for the junior homonym Heliophilus Dejean, 1821 (see Bousquet and Bouchard 2013a: 46) and should have the same type species as Heliophilus Dejean, 1821 (i.e., Pedinushybridus Latreille, 1804); however, recent authors (e.g., Silfverberg 1984: 59, Iwan and Löbl 2008: 279, Iwan and Löbl 2020: 302) have followed the type species designation proposed by Gebien (1938a: 304) and treated Heliopates as a separate genus with Tenebriolusitanicus Herbst, 1797 as its type species; in order to maintain nomenclatural stability we recommend that an application be submitted to the International Commission on Zoological Nomenclature to request that the type species designation proposed by Gebien (1938a: 304) be used for Heliopates Dejean, 1834.

Heliopathes Gebien, 304 [M]. Type Tenebriolusitanicus Herbst, 1797, by subsequent designation (Gebien 1938a: 304). Status: junior synonym of Heliopates Dejean, 1834 in Blaptinae: Dendarini: Dendarina. Note: unjustified emendation of Heliopates Dejean, 1834, not in prevailing usage; see comments regarding the type species of Heliopates Dejean, 1834 under that name.

Heliophilus Dejean, 65 [M]. Type Pedinushybridus Latreille, 1804, by subsequent designation (Duponchel 1845b: 517). Status: senior synonym of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina. Synonymy: Iwan and Löbl (2008: 282). Note: junior homonym of Heliophilus Meigen, 1803 [Diptera]; the currently accepted valid name of the type species, “Phylan abbreviatus (G.-A. Olivier, 1795)”, should not be considered as an available name since G.-A. Olivier (1795: [57] 17) clearly refers to Tenebrioabbreviatus Fabricius, 1775.

Heliophosis Koch, 95 [F]. Type Heliophosiskalaharica Koch, 1952, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Heliophygus Agassiz, 175 [M]. Type species [automatic]: Heliofugusarenosus Guérin-Méneville, 1831, by monotypy. Status: junior synonym of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Heliofugus Guérin-Méneville, 1831, not in prevailing usage.

Heliosteres Hope, 124 [M]. Type species [automatic]: Heliofugusarenosus Guérin-Méneville, 1831, by monotypy. Status: junior synonym of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Heliofugus Guérin-Méneville, 1831.

Heliostrhaema Reitter, 34 [N]. Type Heliotaurusrolphii Fairmaire, 1867, by subsequent designation (R. Lucas 1920: 319). Status: valid genus in Alleculinae: Cteniopodini. Note: Heliostrhaema is an unjustified emendation of the original spelling Heliosthraema, introduced by Seidlitz (1896: 223), in prevailing usage and attributed to the original author and date; it is considered here to be a justified emendation (ICZN 1999, Article 33.2.3.1).

Heliotaurus Mulsant, 73 [M]. Type Cisteladistincta Laporte, 1840, by subsequent designation (Novák and Pettersson 2008: 332). Status: valid genus and subgenus in Alleculinae: Cteniopodini.

Helogria Borchmann, 48, 110 [F]. Type Lagriapruinosa Chevrolat, 1841, by subsequent designation (Borchmann 1936: 143). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Helopelius Reitter, 123, 152 [M]. Type Stenomaxaeneipennis Allard, 1876, by subsequent designation (Nabozhenko 2008: 38). Status: valid subgenus of Stenohelops Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Note: downgraded to a subgenus of Stenohelops Reitter, 1922 by Nabozhenko et al. (2020b: 293).

Helopidesthes Fairmaire, 446 [F]. Type Helopidesthescoriaria Fairmaire, 1895, by monotypy. Status: valid genus in Tenebrioninae: Helopini: incertae sedis.

Helopidoxus Reitter, 32, 44 [M]. Type Helopssuperbus Mulsant & Godart, 1855, by monotypy. Status: valid subgenus of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina.

Helopimorphus Desbrochers des Loges, 140 [M]. Type Helopimorphusangulipennis Desbrochers des Loges, 1881, by monotypy. Status: junior synonym of Heterotarsus Latreille, 1829 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Seidlitz (1896: 24).

Helopinus Solier, 152, 197 [M]. Type Helopinuscostatus Solier, 1848, by monotypy. Status: valid subgenus of Drosochrus Erichson, 1843 in Blaptinae: Pedinini: Helopinina. Note: the First Reviser (Helopinus Solier, 1848 versus Pteraulus Solier, 1848) is Koch (1958: 149)

Helopocerodes Reitter, 122, 144 [M]. Type Helopsfaldermanni Faldermann, 1837, by subsequent designation (Nabozhenko 2001a: 633). Status: junior synonym of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: Keskin et al. (2017: 726).

Helopogonus Reitter, 122, 150 [M]. Type Helopsviridicollis Schaufuss, 1869, by monotypy. Status: valid subgenus of Nesotes Allard, 1876 in Tenebrioninae: Helopini: Helopina.

Helopondrus Reitter, 123, 153 [M]. Type Stenomaxsareptanus Allard, 1876, by subsequent designation (Nabozhenko 2001a: 652). Status: junior synonym of Horistelops Gozis, 1910 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: new synonym [YB]. Note: Horistelops Gozis, 1910 has been forgotten in the literature; its type species is currently included in the subgenusHelopondrus Reitter, 1922 and for that reason Reitter’s name is considered a new junior synonym of Horistelops.

Helopostygnus Antoine, 133, 153 [M]. Type Helopsatlantis Antoine, 1926, by original designation. Status: valid subgenus of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina.

Helopotrichus Reitter, 32, 44 [M]. Type Helopsvillosipennis P.H. Lucas, 1846, by subsequent designation (Viñolas and Cartagena 2005: 38). Status: valid subgenus of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina.

Helops Fabricius, 257 [M]. Type Tenebriocaeruleus Linnaeus, 1758, by subsequent designation (Hope 1841: 133; see ICZN 2009, Opinion 2237). Status: valid genus in Tenebrioninae: Helopini: Helopina. Note: as mentioned in Bousquet et al. (2018: 183) and Vela et al. (2020: 63) the type of the nominal species Tenebriocaeruleus Linnaeus, 1758 is actually a leaf beetle that belongs to the genus Timarcha Samouelle, 1819 (Coleoptera: Chrysomelidae) based on examination of a syntype; since Tenebriocaeruleus Linnaeus, 1758 was placed on the Official List of Specific Names in Zoology and fixed as the type species of Helops Fabricius, 1775 in Opinion 2237 (ICZN 2009), an application to the International Commission on Zoological Nomenclature is necessary to request that the type species of Helops be changed to Tenebriocaeruleus Linnaeus sensu Fabricius, 1775 (= Helopschalibaeus Rossi, 1790).

Helopsallecula Pic, 33 [F]. Type Helopsalleculaminutissima Pic, 1936, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Helopsisomira Pic, 63 [F]. Type Helopsisomirakochi Pic, 1952, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Hemasodes Casey, 378 [M]. Type Schoenicusvestitus Champion, 1884, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Hemeralopius Gistel, viii, 125 [M]. Type species [automatic]: Tenebrioelongatus Herbst, 1797, by monotypy. Status: senior synonym of Belopus Gebien, 1911 in Lagriinae: Belopini. Note: replacement name for Calcar Dejean, 1821; nomen oblitum (see Bouchard and Bousquet 2020b: 6).

Hemerobates Kolbe, 189 [M]. Type Nyctobatesmechowi Kolbe, 1884, by monotypy. Status: junior synonym of Amenophis J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1911a: 441).

Hemicera Laporte & Brullé, 332, 393 [F]. Type Cnodalonsplendens Wiedemann, 1823, by subsequent designation (Gebien 1921a: 335). Status: valid genus and subgenus in Stenochiinae: Cnodalonini. Note: nomenclatural stability in this genus is threatened by the discovery of an older type species designation (Hemiceraarcuata Laporte & Brullé, 1831 [misspelled “Hemiceraarmata”], by subsequent designation by Duponchel (1845b: 528), which is currently the type species of the valid genus Hypocalis Dejean, 1834); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Gebien (1921a: 335).

Hemicistela Blackburn, 331 [F]. Type Hemicisteladiscoidalis Blackburn, 1891, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Hemicyclus Westwood, 44 [M]. Type Hemicyclusgrandis Westwood, 1841 (= Tetraphyllusreaumuri Laporte, 1840), by subsequent designation (Gebien 1941: 1133). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Hemimmedia Gebien, 220, 229 [F]. Type Hemimmediacorpulenta Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hemipraocis Flores & Pizarro-Araya, 66 [M]. Type Praocissellatus Berg, 1889, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 13, 60) without type species designation; the fact that Praocissellatus Berg, 1889 was listed as the “type” of Hemipraocis in the Zoological Record for the year 1958 (Anonymous in Commonwealth Institute of Entomology 1960) does not represent a valid type species designation since the nomenclatural act is anonymous (ICZN 1999, Article 14).

Hemipristis Kolbe, 165, 177 [F]. Type Hemipristisukamia Kolbe, 1903, by present designation. Status: senior synonym of Hemipristula Bouchard & Bousquet, nom. nov. in Lagriinae: Pycnocerini. Note: junior homonym of Hemipristis Agassiz, 1833 [Pisces].

Hemipristula Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Hemipristisukamia Kolbe, 1903, by present designation. Status: valid genus in in Lagriinae: Pycnocerini. Note: replacement name Hemipristis Kolbe, 1903; Hemipristula was proposed earlier by Strand (1935a: 291); however, this name is unavailable because Strand did not designate a type species for the nominal taxon, a mandatory requirement for replacement name without valid typification proposed after 1930 (ICZN 1999; Article 13.3.1).

Hemipterocoma Skopin, 159 [F]. Type Pterocomanikolskii Semenov-Tjan-Shansky, 1910, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Hemitrichestes Reitter, 169 [M]. Type Penthicushirsutus Reitter, 1896, by monotypy. Status: junior synonym of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Löbl et al. (2008b: 267).

Heptaphylla Friedenreich, 375 [F]. Type Heptaphyllafungicola Friedenreich, 1883, by monotypy. Status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: Arrow (1904: 31).

Herbertfranzia Kaszab, 26, 28 [F]. Type Herbertfranzianepalica Kaszab, 1973, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Herbertfranziella Kaszab, 26, 28 [F]. Type Herbertfranziaeutagenoides Kaszab, 1973, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Herlesa Reitter, 298, 301 [F]. Type Herlesaglobicollis Reitter, 1897 (= Micipsacavifrons Fairmaire, 1863), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Herodius Agassiz, 143,179 [M]. Type species [automatic]: Erodiusgibbus Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Erodius Fabricius, 1775 in Pimeliinae: Erodiini. Note: unjustified emendation of Erodius Fabricius, 1775, not in prevailing usage.

Herpiscius Solier, 160, 188 [M]. Type Herpisciusspinolae Solier, 1838, by subsequent designation (Hope 1841: 115). Status: valid genus in Tenebrioninae: Scaurini.

Herpsis Haag-Rutenberg, 4, 66 [F]. Type Adelostomarugosum Guérin-Méneville, 1831, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Herthasida Wilke, 269 [F]. Type Asidaingens Champion, 1892, by monotypy. Status: valid subgenus of Philolithus Lacordaire, 1858 in Pimeliinae: Asidini.

†Hesiodobates Kaszab & Schawaller, 1 [M]. Type Hesiodobatesantiquus Kaszab & Schawaller, 1984, by original designation. Status: junior synonym of Nesocyrtosoma Marcuzzi, 1976 in Stenochiinae: Cnodalonini. Synonymy: Doyen and Poinar (1994: 45). Note: described from Early Miocene amber (Dominican Republic).

Hesiodus Champion, 115 [M]. Type Hesioduslongitarsis Champion, 1885, by subsequent designation (R. Lucas 1920: 323). Status: valid genus in Stenochiinae: Cnodalonini.

Hesperoptorina G.S. Medvedev, 417 [F]. Type Gnaptorinabrucei Blair, 1923, by original designation. Status: valid subgenus of Gnaptorina Reitter, 1887 in Blaptinae: Blaptini: Gnaptorinina.

Hesseodes Ardoin, 92 [M]. Type Hoplonyxkalaharica Hesse, 1935, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: the name Hesseodes used earlier by Ardoin (1962b: 969) is unavailable since it was published after 1930 without a type species designation.

Hesseosis Koch, 76 [F]. Type Hesseosispurpurascens Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Heterarthron Gistel, 190 [N]. Type species [automatic]: Eulabisbicarinata Eschscholtz, 1829, by subsequent designation (Blaisdell 1932: 44). Status: junior synonym of Eulabis Eschscholtz, 1829 in Tenebrioninae: Eulabini. Note: junior homonym of Heterarthron Dejean, 1836 [Coleoptera: Bostrichidae]; unnecessary replacement name for Eulabis Eschscholtz, 1829 (as “Eulabes”).

Heterasida Casey, 76, 165 [F]. Type Pelecyphorusbifurcus J.L. LeConte, 1861, by original designation. Status: valid genus in Pimeliinae: Asidini.

Heterocheira Dejean, 220 [F]. Type Ulomaaustrale Boisduval, 1835, by monotypy. Status: valid genus in Blaptinae: Opatrini: Heterotarsina.

Heterochira Agassiz, 180 [F]. Type species [automatic]: Ulomaaustrale Boisduval, 1835, by monotypy. Status: junior synonym of Heterocheira Dejean, 1836 in Blaptinae: Opatrini: Heterotarsina. Note: unjustified emendation of Heterocheira Dejean, 1836, not in prevailing usage.

Heterogena Froussart, 60, 105 [F]. Type Nesogenagoudotii Fairmaire, 1868, by original designation. Status: valid subgenus of Nesogena Mäklin, 1863 in Tenebrioninae: Praeugenini.

Heterogria Fairmaire, 42 [F]. Type Heterogriapunctatissima Fairmaire, 1896, by monotypy. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Heteromerotylus Pic, 11 [M]. Type Heteromerotylusbicoloripes Pic, 1921, by monotypy. Status: junior synonym of Tearchus Kraatz, 1880 in Stenochiinae: Cnodalonini. Synonymy: Kulzer (1954a: 71).

Heteromira Hölzel, 19 [F]. Type Isomiramoroi Hölzel, 1958, by monotypy. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Heteronicandra Koch, 151 [F]. Type Nicandrazumpti Kulzer, 1951, by original designation. Status: valid subgenus of Nicandra Fairmaire, 1888 in Blaptinae: Pedinini: Helopinina.

Heterophaga Dejean, 199 [F]. Type fixed herein (ICZN 1999, Article 70.3) as Opatrumlaevigatum Fabricius, 1781, misidentified as Tenebriomauritanicus Linnaeus, 1764 in the subsequent designation of Duponchel (1845b: 601). Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Wollaston (1854: 498). Note: see Spilman (1972: 32) for the history of the misidentification.

Heterophyllus Gemminger in Gemminger and Harold, 1955 [M]. Type species [automatic]: Heterophyluschrysomelinus Klug, 1833, by monotypy. Status: junior synonym of Heterophylus Klug, 1833 in Diaperinae: Diaperini: Diaperina. Note: unjustified emendation of Heterophylus Klug, 1833, not in prevailing usage.

Heterophylus Klug, 91 [M]. Type Heterophyluschrysomelinus Klug, 1833, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Heterophylus Mulsant & Rey, 7 [M]. Type Heliopatespicipes Faldermann, 1837, by subsequent designation (Gebien 1938a: 397). Status: senior synonym of Dissonomus Jacquelin du Val, 1861 in Tenebrioninae: Dissonomini. Note: junior homonym of Heterophylus Klug, 1833 [Tenebrionidae: Diaperinae: Diaperini: Diaperina].

Heteropromus Blaisdell, 179 [M]. Type Eleodesvetorator Horn, 1874, by monotypy. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Heteropsectropus Kaszab, 33, 34 [M]. Type Heteropsectropusaenescens Kaszab, 1941, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Heteropus Laporte, 221 [M]. Type Heteropusholosericeus Laporte, 1840, by monotypy. Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Lacordaire (1859a: 250). Note: junior homonym of Heteropus Palisot de Beauvois, 1820 [Mammalia].

Heteroscelis Agassiz, 181, 266 [F]. Type species [automatic]: Adesmiapulcherrima Solier, 1835 (= Adesmiaaudouini Solier, 1835), by subsequent designation (Hope 1841: 118). Status: junior synonym of Oteroscelis Solier, 1835 in Pimeliinae: Adesmiini. Note: unjustified emendation of Oteroscelis Solier, 1835, not in prevailing usage; junior homonym of Heteroscelis Latreille, 1828 [Coleoptera: Tenebrionidae: Blaptinae: Platynotini: Platynotina].

Heteroscelis Latreille, 574 [F]. Type Blapsdentipes Fabricius, 1794, by subsequent designation (Blanchard 1844: pl. 48). Status: senior synonym of Anomalipus Guérin-Méneville, 1831 in Blaptinae: Platynotini: Platynotina. Synonymy: Lacordaire (1859a: 257). Note: nomen oblitum (see Bouchard and Bousquet 2020a: 101); this genus used to be credited to Latreille (1829a: 18) and treated as a junior homonym of Heteroscelis Latreille, 1829 [Hemiptera] until bibliographic research determined that the tenebrionid name was made available earlier by Latreille (1828: 574) and is in fact the senior homonym (see Bouchard and Bousquet 2020a: 101).

Heterostrongylium Kaszab, 10, 27 [N]. Type Strongyliumweiskei Gebien, 1921, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Heterotarsus Latreille, 26 [M]. Type Heterotarsustenebrioides Guérin-Méneville, 1831, by subsequent monotypy (Guérin-Méneville 1831b: pl. 30). Status: valid genus in Blaptinae: Opatrini: Heterotarsina. Note: originally proposed without included nominal species; Guérin-Méneville (1831b: pl. 30), by including the new species Heterotarsustenebrioides Guérin-Méneville, 1831 in association with this name, was the first author to subsequently and expressly include nominal species in Heterotarsus (ICZN 1999, Article 67.2.2).

Hexagonochilus Solier, 168 [M]. Type Hexagonochilusdilaticollis Solier, 1851, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina. Note: Hexagonochilus is an incorrect subsequent spelling of the original spelling Hexagonocheilus, first used by Imhoff (1856: 234), and is in prevailing usage; Hexagonochilus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Hexarhopalus Fairmaire, xix [M]. Type Hexarhopalussculpticollis Fairmaire, 1891, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Hexarhoptrum Fairmaire, 38 [N]. Type species [automatic]: Hexarhopalussculpticollis Fairmaire, 1891, by monotypy. Status: junior synonym of Hexarhopalus Fairmaire, 1891 in Stenochiinae: Cnodalonini. Note: the alternative original spelling Hexaroptrum, used by Fairmaire (1894a: 38), was rejected by Bečvář and Purchart (2008: 39) who acted as the First Revisers; unnecessary replacement name for Hexarhopalus Fairmaire, 1891.

Hicetaon Champion, 111 [M]. Type Hicetaonfrontalis Champion, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hidrosella Koch, 32 [F]. Type Hidrosisincostata Haag-Rutenberg, 1875, by original designation. Status: valid subgenus of Machlopsis Pomel, 1871 in Pimeliinae: Adelostomini.

Hidrosis Haag-Rutenberg, 120 [F]. Type Steiracrenatocostata Redtenbacher, 1868, by subsequent designation (Löbl et al. 2008b: 121). Status: junior synonym of Machlopsis Pomel, 1871 in Pimeliinae: Adelostomini. Synonymy: Bedel (1887: 199).

Himastethe Koch, 286 [F]. Type Cyphostethegigantea Koch, 1950, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini.

Himatismus Erichson, 253 [M]. Type Himatismusmandibularis Erichson, 1843, by subsequent designation (Gebien 1937a: 574). Status: valid subgenus of Imatismus Dejean, 1834 in Pimeliinae: Tentyriini.

Hionthis Miller, 174 [F]. Type Hionthistentyrioides Miller, 1861, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Hionthisoma Gebien, 609 [N]. Type species [automatic]: Hionthisoccidentalis Fairmaire, 1897, by monotypy. Status: junior synonym of Hyonthosoma Reitter, 1900 in Pimeliinae: Tentyriini. Note: unjustified emendation of Hyonthosoma Reitter, 1900, not in prevailing usage.

Hipalmus Bates, 269 [M]. Type Tenebriocostatus Guérin-Méneville, 1831, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini. Note: the original combination of the name of the type species, Tenebriocostatus Guérin-Méneville, 1831, is a junior primary homonym of Tenebriocostatus Pallas, 1781.

Hipomelus Dejean, 181 [M]. Type Sepidium vittatum Fabricius, 1781, by subsequent designation (Hope 1841: 116). Status: junior synonym of Trachynotus Latreille, 1828 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Bousquet and Bouchard (2013a: 47).

Hipponome Laporte, 235 [F]. Type Helopsazureus Brullé, 1832, by monotypy. Status: senior synonym of Raiboscelis Allard, 1876 in Tenebrioninae: Helopini: Helopina. Synonymy: Seidlitz (1895: 754). Note: nomen oblitum (see Nabozhenko and Löbl 2009: 194).Note: the original combination of the name of the type species, Helopsazureus Brullé, 1832, is a junior primary homonym of Helopsazureus Germar, 1823.

Hirsutosora Pic, 32 [F]. Type Nemostirafortithorax Pic, 1922, by monotypy. Status: valid subgenus of Sora Walker, 1859 in Lagriinae: Lagriini: Statirina.

Hirtograbies Koch, 23 [M]. Type Hirtograbiesoograbiesensis Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Hispanomelia Mas-Peinado, Buckley, Ruiz & García-París, 541 [F]. Type Pimeliamanchega Lauffer, 1905, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Histeromimus Gahan, 288 [M]. Type Histeromimusarabicus Gahan, 1895, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Histeromorphus Kraatz, 6, 11 [M]. Type Histeromorphusplicatus Kraatz, 1865, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Histeropsis Chevrolat, 221 [F]. Type Platydemaamericana Laporte & Brullé, 1831, by subsequent designation (Löbl et al. 2008a: 42). Status: junior synonym of Platydema Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Synonymy: Champion (1886: 181).

Histiaea Fairmaire, 107 [F]. Type Histiaeabidentula Fairmaire, 1892, by monotypy. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini.

Histrionotus Koch, 44 [M]. Type Trachynotuslightfooti Péringuey, 1892, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Holaniara Fairmaire, 43 [F]. Type species [automatic]: Ulomapiceum Melsheimer, 1846, by monotypy. Status: junior synonym of Eutochia J.L. LeConte, 1862 in Tenebrioninae: Ulomini. Note: replacement name for Aniara Melsheimer, 1853.

Holdhausia Reitter, 125 [F]. Type Cteniopuscrassus Fairmaire, 1892, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Holeleodes Blaisdell, 132 [M]. Type Eleodesbeameri Blaisdell, 1937 (= Eleodeshepburni Champion, 1884), by original designation. Status: junior synonym of Steneleodes Blaisdell, 1909 in Blaptinae: Amphidorini. Synonymy: Johnston (2015: 12).

Holoblaps Bauer, 233 [F]. Type none designated. Status: undetermined taxon in Blaptinae: Blaptini: Blaptina. Note: this genus was described before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Holoblaps and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Holobrachium Gebien, 256 [N]. Type species [automatic]: Hoplobrachiumasperipenne Fairmaire, 1886 (= Helopsdentipes Fabricius, 1781), by monotypy. Status: junior synonym of Hoplobrachium Fairmaire, 1886 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Hoplobrachium Fairmaire, 1886, not in prevailing usage.

Holobrachys Fairmaire, 233 [M]. Type Holobrachysheterocerus Fairmaire, 1869, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hologenosis Deyrolle, 81, 82 [F]. Type Hologenosislacerata Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Holostrongylium Kaszab, 10, 20 [N]. Type Strongyliumgravidum Mäklin, 1867, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Homala Eschscholtz, 5, 6 [F]. Type Homalapolita Eschscholtz, 1831, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Homalapipleurus Español, 166, 167 [M]. Type Hegetergonzalezi Español, 1957, by original designation. Status: valid subgenus of Hegeter Latreille, 1802 in Pimeliinae: Tentyriini.

Homaleis Rye, 62 [M]. Type species [automatic]: Helopscongener Reiche, 1861, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Note: unjustified emendation of Omaleis Allard, 1877, not in prevailing usage.

Homalinota Koch, 66 [F]. Type species [automatic]: Homalaagona Fairmaire, 1884, by subsequent designation (Gebien 1937a: 610). Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Homalopsis Lesne, 1922.

Homalopsis Lesne, 675 [F]. Type Homalaagona Fairmaire, 1884, by subsequent designation (Gebien 1937a: 610). Status: senior synonym of Homalinota Koch, 1950 in Pimeliinae: Tentyriini. Note: junior homonym of Homalopsis Kuhl & Hasselt, 1822 [Reptilia].

Homalus Rye, 69 [M]. Type species [automatic]: Helopscongener Reiche, 1861, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Note: unjustified emendation of Omaleis Allard, 1876 (as “Omalus”); junior homonym of Homalus Agassiz, 1846 [Hymenoptera].

Homebius Endrödy-Younga, 124 [M]. Type Homebiuskaszabi Endrödy-Younga, 1989, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Homebiina.

Homocyrtus Dejean, 211 [M]. Type species [automatic]: Cyphonotusdromedarius Guérin-Méneville, 1831, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: replacement name for Cyphonotus Guérin-Méneville, 1831; based on a recent assessment of adult and larval morphological characters, Matthews and Lawrence (2015: 290) determined that this genus could not be placed in any known tribe and that the status of Homocyrtus should be regarded as Tenebrionidae incertae sedis.

Homoeocamaria Blair, 75 [F]. Type species [automatic]: Homoeogenuslaticornis C.O. Waterhouse, 1882, by monotypy. Status: junior synonym of Borneocamaria Pic, 1917 in Stenochiinae: Cnodalonini. Note: replacement name for Homoeogenus C.O. Waterhouse, 1882.

Homoeogenus C.O. Waterhouse, 174 [M]. Type Homoeogenuslaticornis C.O. Waterhouse, 1882, by monotypy. Status: senior synonym of Borneocamaria Pic, 1917 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1942a: 323). Note: junior homonym of Homoeogenus C.O. Waterhouse, 1880 [Coleoptera: Psephenidae].

Homoeonota Fairmaire in Fairmaire et al., 1882: 63 [F]. Type Homoeonotasubopaca Fairmaire, 1882, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Homopsis Semenov, 258, 263 [F]. Type Homopsisgrumi Semenov, 1893, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Homoropsis Fairmaire, 450 [F]. Type Homoropsisustulata Fairmaire, 1886, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Homotrysis Pascoe, 489 [F]. Type Alleculatristis Germar, 1848 (= Alleculacarbonaria Germar, 1848), by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: the First Reviser (Homotrysis Pascoe, 1866 versus Hybrenia Pascoe, 1866) is Champion (1897: 158).

Hopatrinus Agassiz, 185, 260 [M]. Type species [automatic]: Opatrumclathratum Fabricius, 1787, by monotypy. Status: junior synonym of Opatrinus Dejean, 1821 in Blaptinae: Platynotini: Platynotina. Note: unjustified emendation of Opatrinus Dejean, 1821, not in prevailing usage.

Hopatroides Agassiz, 185, 260 [M]. Type species [automatic]: Opatroidespunctulatus Brullé, 1832, by monotypy. Status: junior synonym of Opatroides Brullé, 1832 in Blaptinae: Opatrini: Opatrina. Note: unjustified emendation of Opatroides Brullé, 1832, not in prevailing usage.

Hopatromorpha Blackburn, 286 [F]. Type species [automatic]: Opatrummurinum Baudi di Selve, 1876 (= Opatrinussetosus Mulsant & Rey, 1853), by subsequent designation (Gebien 1938a: 399). Status: junior synonym of Mesomorphus Miedel, 1880 in Blaptinae: Opatrini: Opatrina. Note: unnecessary replacement name for Mesomorphus Miedel, 1880 (incorrectly attributed to “Reitter, 1904”).

Hopatropteron Reitter, 701 [N]. Type Hopatropteronsubcostatum Reitter, 1889 (= Heterotarsuscarinula Marseul, 1876), by monotypy. Status: junior synonym of Heterotarsus Latreille, 1829 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Seidlitz (1894: 413), Gebien (1911a: 473).

Hopatrum Agassiz, 185, 260 [N]. Type species [automatic]: Silphasabulosa Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Opatrum Fabricius, 1775 in Blaptinae: Opatrini: Opatrina. Note: unjustified emendation of Opatrum Fabricius, 1775, not in prevailing usage.

Hoplambius Reitter, 114 [M]. Type Hoplarionmelambioides (as “melambioide”) Fairmaire, 1893, by monotypy. Status: valid subgenus of Melambius Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Hoplariobius Reitter, 115 [M]. Type Micrositusdecurtatus Fairmaire, 1884, by subsequent designation (Antoine 1942: 25). Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Hoplarion Mulsant & Rey, 150 [N]. Type Micrositustumidus Mulsant & Rey, 1854, by monotypy. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Hoplaspis Motschulsky, 113 [F]. Type Hoplaspislamellicornis Motschulsky, 1858, by subsequent designation (Crotch 1876: 418). Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: the alternative original spelling Hoplapsis, used by Motschulsky (1858: 113), was rejected by Motschulsky (1868: 69) who acted as the First Reviser (ICZN 1999, Article 24.2.4); transferred from Cucujoidea: Erotylidae by Arrow (1909: 196).

Hoplitoblaps Fairmaire, 26 [F]. Type Hoplitoblapsfallaciosa Fairmaire, 1889, by monotypy. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Hoplobrachium Fairmaire, 74 [N]. Type Hoplobrachiumasperipenne Fairmaire, 1886 (= Helopsdentipes Fabricius, 1781), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Hoplocephala Agassiz, 185, 262 [F]. Type species [automatic]: Ips haemorrhoidalis Fabricius, 1787, by subsequent designation (Motschulsky 1845a: 80). Status: junior synonym of Neomida Latreille, 1829 in Diaperinae: Diaperini: Diaperina. Note: unjustified emendation of Oplocephala Laporte & Brullé, 1831, not in prevailing usage.

Hoplochirus Scudder, 153 [M]. Type species [automatic]: Hoplonyxalleculoides J. Thomson, 1858, by subsequent designation (Gebien 1943: 919). Status: junior synonym of Hoplonyx J. Thomson, 1858 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Oplocheirus Klug, 1835, not in prevailing usage.

Hoploedipinus Kaszab, 355, 356 [M]. Type Hoploedipusheterodoxus Fairmaire, 1898, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Hoploedipus Fairmaire, 395 [M]. Type Camarimenaarmipes Fairmaire, 1882, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Hoplonyx J. Thomson, 98 [M]. Type Hoplonyxalleculoides J. Thomson, 1858, by subsequent designation (Gebien 1943: 919). Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Hoplopeltis Fairmaire, 22 [F]. Type Hoplopeltistricornis Fairmaire, 1894, by monotypy. Status: valid genus in Tenebrioninae: Alphitobiini.

Hoploptera Gemminger in Gemminger and Harold, 2037 [F]. Type species [automatic]: Strongyliumserraticorne Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Oploptera Chevrolat, 1844 in Stenochiinae: Stenochiini. Note: unjustified emendation of Oploptera Chevrolat, 1844, not in prevailing usage.

Hoplostira Borchmann, 217, 225 [F]. Type Hoplostirafemoralis Borchmann, 1921, by original designation. Status: junior synonym of Spinostatira Pic, 1918 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1936: 247).

Hoplostrongylium Ardoin, 1326 [N]. Type Hoplostrongyliumghesquierei Ardoin, 1965, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Horatoma Solier, 210, 264 [364] [F]. Type Horatomaparvula Solier, 1841, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Horatomella Penrith & Endrödy-Younga, 6, 7 [F]. Type Parapachynotelajohni Koch, 1957, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Horatomodes Haag-Rutenberg, 274, 305 [M]. Type Horatomodesbatesi Haag-Rutenberg, 1872, by monotypy. Status: junior synonym of Horatoma Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Synonymy: Penrith and Endrödy-Younga (1994: 10).

Horistelops Gozis, 112 [M]. Type Helopsassimilis Küster, 1850, by original designation. Status: valid subgenus of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina.

Hosohamudama Masumoto, 43 [F]. Type Hosohamudamasasajii Masumoto, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Houaphanica Novák, 470 [F]. Type Houaphanicafera Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Hovacula Fairmaire, 236 [F]. Type Hovaculalineolata Fairmaire, 1898, by monotypy. Status: valid genus in Alleculinae: incertae sedis. Note: placed in Alleculinae by Chatanay (1915a: 526).

Hovadelium Ardoin, 34 [N]. Type Hovadeliumdiscoidale Ardoin, 1961, by original designation. Status: valid genus in Lagriinae: Laenini.

Hovademulus Iwan, 385, 390 [M]. Type Selinuspunctipennis Fairmaire, 1902, by original designation. Status: junior synonym of Clastopus Fairmaire, 1898 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (2001: 500).

Hovademus Iwan, 385, 395 [M]. Type Hovademusandringitrensis Ardoin, 1974, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina. Note: the name Hovademus was proposed earlier by Ardoin (1974a: 165) without a type species designation; the fact that Hovademusandringitrensis Ardoin, 1974 was listed as the “type” of Hovademus in the Zoological Record for the year 1974 (Anonymous in Staff of the Zoological Society of London 1979) does not represent a valid type species designation since the nomenclatural act is anonymous (ICZN 1999, Article 14).

Hovarygmus Fairmaire, 234 [M]. Type Hovarygmusinsularis Fairmaire, 1898, by monotypy. Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Huilamus Koch, 79 [M]. Type Huilamuswelwitschi Koch, 1953, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Hummelinckia Marcuzzi, 19 [F]. Type Hummelinckiacaraibica Marcuzzi, 1954, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Hyalarthrodosis Kaszab, 74 [F]. Type Arthrodosismonodi Pierre, 1974, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Hyalerodius Kaszab, 80 [M]. Type Hyalerodiusjirofti Kaszab, 1979, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Hybocaulus Fairmaire, 27 [M]. Type Hybocauluslaticornis Fairmaire, 1895 (= Porphyrhybaviolaceicolor Fairmaire, 1877), by monotypy. Status: junior synonym of Porphyrhyba Fairmaire, 1877 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1956b: 89).

Hybonotus Dejean, 211 [M]. Type Tetraphyllusformosus Laporte & Brullé, 1831, by monotypy. Status: senior synonym of Damatris Laporte, 1840 in Stenochiinae: Cnodalonini. Synonymy: new synonym [PB]. Note: the type species of Hybonotus Dejean, 1834 is the same as the type species of Damatris Laporte, 1840 and therefore the two genera are objective synonyms; junior homonym of Hybonotus Klug, 1803 [Hymenoptera].

Hyboproctus Kolbe, 241 [M]. Type Hyboproctusnodifer Kolbe, 1897, by subsequent designation (Gebien 1940: 1089). Status: valid genus in Stenochiinae: Cnodalonini.

Hybrenia Pascoe, 489 [F]. Type Hybreniavittata Pascoe, 1866, by subsequent designation (Matthews and Bouchard 2008: 328). Status: junior synonym of Homotrysis Pascoe, 1866 in Alleculinae: Alleculini: Alleculina. Synonymy: Champion (1897: 158).

Hydissus Pascoe, 148 [M]. Type Nyctobatesferonioides Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hydisus Scudder, 167 [M]. Type species [automatic]: Nyctobatesferonioides Pascoe, 1866, by monotypy. Status: junior synonym of Hydissus Pascoe, 1869 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Hydissus Pascoe, 1869, not in prevailing usage.

Hylithus Guérin-Méneville, 12 [M]. Type Hylithustentyrioides Guérin-Méneville, 1834, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Hylocrinus Casey, 289, 331 [M]. Type Eurymetoponlungulum J.L. LeConte, 1851, by original designation. Status: valid genus and subgenus in Pimeliinae: Edrotini.

Hylonoma Macquart, 183 [F]. Type Alegoriadilatata Laporte, 1840, by monotypy. Status: junior synonym of Alegoria Laporte, 1840 in Tenebrioninae: Ulomini. Synonymy: Lacordaire (1859a: 325).

Hyloplonyx Ardoin, 716 [M]. Type Hoplonyxmonophthalmus J. Thomson, 1858, by original designation. Status: valid subgenus of Hoplonyx J. Thomson, 1858 in Tenebrioninae: Amarygmini.

Hymenalia Mulsant, 48 [F]. Type Cistelafusca Illiger, 1794 (= Cistelarufipes Fabricius, 1792), by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Hymenochara Campbell, 435 [F]. Type Mycetophilarufipes J.E. LeConte, 1824, by original designation. Status: valid genus in Alleculinae: Alleculini: Mycetocharina.

Hymenorus Mulsant, 68 [M]. Type Hymenorusdoublieri Mulsant, 1852, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: the alternative original spelling Hymenophorus was corrected to Hymenorus in the “Emendanda” of the same work (p. 188), Hymenorus is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1); see Bousquet et al. (2015: 133).

Hyocis Pascoe, 457 [M]. Type Hyocisbakewellii Pascoe, 1866, by original designation. Status: valid genus and subgenus in Diaperinae: Hyociini: Hyociina.

Hyonthosoma Reitter, 89, 142 [N]. Type Hionthisoccidentalis Fairmaire, 1897, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Hypamarygmus Gebien, 27 [M]. Type Hypamarygmuscoccinelloides Gebien, 1904, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Hypaulax Bates, 259 [F]. Type Hypaulaxmarginata Bates, 1868, by subsequent designation (Gebien 1941: 333). Status: valid genus in Stenochiinae: Cnodalonini. Note: the First Reviser (Hypaulax Bates, 1868 versus Chileone Bates, 1868) is Carter (1914b: 46).

Hyperamarygmus Kaszab, 291 [M]. Type Hyperamarygmusantennalis Kaszab, 1964, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Hyperchalca Fairmaire, 238 [F]. Type Hyperchalcaaenescens Fairmaire, 1869, by monotypy. Status: valid genus and subgenus in Stenochiinae: Stenochiini.

Hypercossyphodes Andreae, 205, 215 [M]. Type Cossyphodesvandami Andreae, 1961, by original designation. Status: junior synonym of Cossyphodes Westwood, 1851 in Pimeliinae: Cossyphodini: Cossyphodina. Synonymy: Schawaller (2013c: 362, implied by inclusion of Cossyphodesvandami Andreae, 1961 in Cossyphodes Westwood, 1851 without use of a subgenus rank).

Hypermicrotelopsis Koch, 743 [F]. Type Microtelopsisthibetana Koch, 1940, by monotypy. Status: valid subgenus of Microtelopsis Koch, 1940 in Pimeliinae: Stenosini: Stenosina. Note: combined description of new genus-group taxon and a single new species (ICZN 1999, Article 13.4); we act as First Revisers and select Microtelopsis Koch, 1940 as the valid name for this genus instead of Extetranosis Koch, 1940 and Hypermicrotelopsis Koch, 1940.

Hyperops Eschscholtz, 5, 9 [M]. Type Hyperopstagenioides Eschscholtz, 1831, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Hyperopsis Bouchard & Bousquet, new subgenus [F]. Type Pachyceracolasi Koch, 1943, by present designation. Status: valid subgenus of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: Koch (1943a: 524, 536) introduced the new subgenus name Hyperopsis for five nominal species, but unfortunately did not designate a type species; the subgenusHyperopsis, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Pachyceracolasi Koch, 1943 as type species and referring to Koch (1943a: 524) for the character states that characterise and differentiate Hyperopsis.

Hypoblaps Kolbe, 200 [F]. Type Blapsrotundata Solier, 1848, by original designation. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Hypocalis Dejean, 206 [F]. Type Hemiceraarcuata Laporte & Brullé, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hypocilibe Bates, 275 [F]. Type Hypocilibemacleayi Bates, 1872, by monotypy. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1911b: 139, with Onosterrhus Pascoe, 1866, a junior synonym of Nyctozoilus Guérin-Méneville, 1831).

Hypocistela Bates, 482 [F]. Type Hypocistelatenuipes Bates, 1879, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Hypogena Dejean, 199 [F]. Type Tenebriobiimpressus Latreille, 1833 (= Peltisbrasilica Perty, 1830), by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Hypolaenopsis Masumoto, 45 [F]. Type Hypolaenopsisuenoi Masumoto, 2001 (= Laenananpingica Schawaller, 2001), by original designation. Status: valid genus in Lagriinae: Laenini.

Hypomelus Solier, 4, 93, 126 [M]. Type Hypomelusbicolor Solier, 1843 (= Helopsperonatus Germar, 1823), by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Hypophlaeus Fabricius, 222 [M]. Type Hypophlaeuscastaneus Fabricius, 1790 (= Corticeusunicolor Piller & Mitterpacher, 1783), by subsequent designation (Curtis 1832: pl. 430). Status: junior synonym of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini. Synonymy: Crotch (1870a: 47).

Hypoprosodes Reitter, 122 [M]. Type Prosodesminuta Kraatz, 1881, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Hypostatira Fairmaire, xlix [F]. Type Hypostatiravariicolor Fairmaire, 1889, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Hypovinsonia Ardoin, 207 [F]. Type Hypovinsoniaalbopilosa Ardoin, 1961, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hypselops Solier, 135 [M]. Type Hypselopsoblongus Solier, 1851, by subsequent designation (R. Lucas 1920: 347). Status: valid genus in Pimeliinae: Epitragini.

Hypsosoma Ménétriés, 30 [N]. Type Hypsosomamongolicum Ménétriés, 1854, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Hypulus Rafinesque, 114 [M]. Type species [automatic]: Tenebriocaeruleus Linnaeus, 1758, by subsequent designation (Hope 1841: 133; see ICZN 2009, Opinion 2237). Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Note: unnecessary replacement name for Helops Fabricius, 1775; junior homonym of Hypulus Paykull, 1798 [Coleoptera: Melandryidae]; see entry for Helops Fabricius, 1775 for information regarding the identity of the type species.

Hysterarthron J. Thomson, 224 [N]. Type Hysterarthroncollare J. Thomson, 1864, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Iberomelia Mas-Peinado, Buckley, Ruiz & García-París, 539 [F]. Type Pimeliacastellana Pérez Arcas, 1865, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Ibnsaudia Koch, 290 [F]. Type Ibnsaudiapropheta Koch, 1941, by monotypy. Status: junior synonym of Thraustocolus Kraatz, 1866 in Pimeliinae: Tentyriini. Synonymy: Kaszab (1981b: 341).

Iccius Champion, 147 [M]. Type Icciuscephalotes Champion, 1886, by subsequent designation (Gebien 1940: 760). Status: valid genus in Diaperinae: Diaperini: Adelinina.

Idahelops Keskin & Nabozhenko, 63 [M]. Type Idahelopsalpagutae Keskin & Nabozhenko, 2012, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Idastrandiella Strand, 23 [F]. Type species [automatic]: Helopsmucoreus Waltl, 1838, by monotypy. Status: junior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Note: replacement name for Apolites Jacquelin du Val, 1861.

Idatius Fairmaire, 276 [M]. Type Idatiusophtalmicus Fairmaire, 1906, by monotypy. Status: valid genus in Alleculinae: incertae sedis. Note: placed in Alleculinae by Chatanay (1915a: 526).

Idiesa Reitter, 203, 245 [F]. Type Diesiafischeri Ménétriés, 1849, by subsequent designation (Gebien 1937a: 807). Status: valid genus in Pimeliinae: Pimeliini.

Idiobates Casey, 62 [M]. Type Tenebriocastaneus Knoch, 1801, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Idiopsis Kaszab, 78 [F]. Type Idiopsisopaca Kaszab, 1981, by original designation. Status: senior synonym of Kocakia Kaszab, 1985 in Pimeliinae: Edrotini. Note: junior homonym of Idiopsis Brauer & Bergenstamm, 1890 [Diptera].

Idisia Pascoe, 452 [F]. Type Idisiaornata Pascoe, 1866, by monotypy. Status: valid genus in Pimeliinae: Idisiini.

Idricus Fairmaire, 199 [M]. Type Idricusdiabolicus Fairmaire, 1888, by monotypy. Status: junior synonym of Ametrocera Fåhraeus, 1870 in Blaptinae: Pedinini: Helopinina. Synonymy: Péringuey (1904: 296).

Iliodera Skopin, 396 [F]. Type Microderadesertoides Skopin, 1961, by original designation. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Ilus Champion, 117 [M]. Type Ilusapicicornis Champion, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ilyxerus Pascoe, 458 [M]. Type Ilyxerusasper Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Imatismus Dejean, 202 [M]. Type Helopsfasciculatus Fabricius, 1798, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Immedia Pascoe, 33 [F]. Type Immediaocculta Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Impressallecula Pic, 12 [F]. Type Impressalleculapurpureipes Pic, 1951, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Impressosora Pic, 254 [F]. Type Impressosoranotaticollis Pic, 1952, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina. Note: new placement [OM], previously included in Lagriinae: Lagriini: Lagriina.

Indenicmosoma Ardoin, 688, 689 [N]. Type Enicmosomaindochinense Kaszab, 1940, by original designation. Status: valid genus in Lagriinae: Lupropini.

Indeucolus Kaszab, 280, 282 [M]. Type Indeucoluscostatus Kaszab, 1975, by original designation. Status: junior synonym of Eucolus Mulsant & Rey, 1853 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (1997: 258).

Indianosis Koch, 297 [F]. Type Stenosiscapitata Koch, 1941, by monotypy. Status: valid subgenus of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina.

Indochillus Koch, 300 [M]. Type Indochilluscristatus Koch, 1941, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Indoprosodes G.S. Medvedev, 690 [M]. Type Prosodesboorpi Kaszab, 1956, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Indostola G.S. Medvedev, 557 [F]. Type Indostolapulchella G.S. Medvedev, 1991, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Indricula Novák, 47 [F]. Type Indriculaargynnis Novák, 2016, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Inscutoheliofugus Freude, 126, 130 [M]. Type Heliofuguskuscheli Freude, 1960, by original designation. Status: valid subgenus of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini.

Insolitoplonyx Bremer, 178, 181 [M]. Type Insolitoplonyxseorsus Bremer, 2014, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Inspinogeton Pic, 174 [M]. Type Cyriogetonimpressipennis Pic, 1937, by monotypy. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Insulasida Escalera, 64 [F]. Type Asidamoraguezi Schaufuss, 1879, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 291).

Iphicorynus Jacquelin du Val, 299 [M]. Type Pentaphyllusmelanophthalmus Mulsant, 1854 (= Nitidulachrysomeloides Rossi, 1792), by monotypy. Status: junior synonym of Pentaphyllus Dejean, 1821 in Diaperinae: Diaperini: Diaperina. Synonymy: Gemminger in Gemminger and Harold (1870: 1956).

Iphius Dejean, 203 [M]. Type Tenebrioserratus Fabricius, 1775, by monotypy. Status: senior synonym of Prioscelis Hope, 1841 in Lagriinae: Pycnocerini. Synonymy: Imhoff (1856: 238). Note: junior homonym of Iphius Schönherr, 1823 [Coleoptera: Curculionidae].

Iphthimera Reitter, 4 [F]. Type Stenocararuficorne Solier, 1835, by present designation. Status: junior synonym of Metriopus Solier, 1835 in Pimeliinae: Adesmiini. Synonymy: new synonym [PB]. Note: the type species of Iphthimera Reitter, 1916 is currently included in the genus Metriopus Solier, 1835 and for that reason Iphthimera is considered a junior synonym of Solier’s name.

Iphthiminus Spilman, 42 [M]. Type Iphthinusitalicus Truqui, 1857, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Iphthimulus Reitter, 16, 17 [M]. Type Iphthinustruquii Marseul, 1869, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Iphthimus Gemminger in Gemminger and Harold, 1977 [M]. Type species [automatic]: Tenebriogigas Linnaeus, 1763, by subsequent designation (Spilman 1973: 42). Status: junior synonym of Mylaris Pallas, 1781 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Iphthinus Dejean, 1834, not in prevailing usage.

Iphthinus Dejean, 203 [M]. Type Tenebriogigas Linnaeus, 1763, by subsequent designation (Spilman 1973: 42). Status: junior synonym of Mylaris Pallas, 1781 in Stenochiinae: Cnodalonini. Synonymy: Chevrolat (1845b: 106, with Nyctobates Guérin-Méneville, 1834, a junior synonym of Mylaris Pallas, 1781), Spilman (1973: 42).

Iranarthrodosis Kaszab, 334 [F]. Type Arthrodosispfaundleri Schuster, 1935, by original designation. Status: junior synonym of Erodiontes Reitter, 1914 in Pimeliinae: Erodiini. Synonymy: Kaszab (1979a: 88).

Iranerodius Kaszab, 334 [M]. Type Arthrodosisrichteri Kaszab, 1957, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Iranolasiostola Pierre, 1020 [F]. Type Iranolasiostoladavatchii Pierre, 1968, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Iranopachyscelis Pierre, 1027 [F]. Type Iranopachysceliseghbali Pierre, 1968 (= Thripterapersica Redtenbacher, 1850), by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Iranosodes G.S. Medvedev, 605 [M]. Type Prosodeskaszabi G.S. Medvedev & Kabakov, 1996, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Irianobates Kaszab, 291 [M]. Type Irianobateskrikkeni Kaszab, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Isaminas Champion, 266 [M]. Type Isaminasgibbipennis Champion, 1887, by subsequent designation (Gebien 1943: 401). Status: valid genus in Stenochiinae: Cnodalonini.

Isarida Pascoe, 456 [F]. Type Isaridatestacea Pascoe, 1866 (= Caediusfulvus Mulsant & Rey, 1859), by monotypy. Status: junior synonym of Caedius Blanchard, 1845 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1939: 466).

Iscanus Fauvel, 176 [M]. Type Iscanuskuniensis Fauvel, 1904, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Ischnarthron Gebien, 47 [N]. Type Ischnarthronlongipes Gebien, 1921, by monotypy. Status: valid genus in Diaperinae: Hypophlaeini.

Ischnodactylus Chevrolat, 173 [M]. Type Ischnodactylusquadrioculatus Chevrolat, 1877, by monotypy. Status: junior synonym of Basides Motschulsky, 1873 in Diaperinae: Diaperini: Diaperina. Synonymy: Schawaller (2004: 47, implied through placement of the type species of Basides Motschulsky, 1873 in Ischnodactylus Chevrolat, 1877). Note: the original spelling of the type species, Ischnodactylusquadridentatus, was corrected to Ischnodactylusquadrioculatus in the “Errata & Notes” of the same work (p. 178), Ischnodactylusquadrioculatus is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Isicerdes Champion, 113 [M]. Type Isicerdesoccultus Champion, 1885, by subsequent designation (R. Lucas 1920: 353). Status: valid genus in Stenochiinae: Cnodalonini.

Ismarus Haag-Rutenberg, 104 [M]. Type Ismarusgodeffroyi Haag-Rutenberg, 1878, by monotypy. Status: senior synonym of Simarus Borchmann, 1909 in Alleculinae: Alleculini: Alleculina. Note: junior homonym of Ismarus Haliday, 1835 [Hymenoptera].

Isocera Borchmann, 713 [F]. Type species [automatic]: Isotomaemarginaticollis Blanchard, 1842, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: replacement name for Isotoma Blanchard, 1842.

Isocerus Dejean, 66 [M]. Type Tenebriopurpurascens Herbst, 1799 (= Helopsferrugineus Fabricius, 1798), by monotypy. Status: senior synonym of Neoisocerus Bouchard, Lawrence, Davies & Newton, 2005 in Blaptinae: Dendarini: Dendarina. Note: junior homonym of Isocerus Illiger, 1802 [Coleoptera: Cerambycidae].

Isomira Mulsant, 52 [F]. Type Chrysomelamurina Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 119). Status: valid genus and subgenus in Alleculinae: Alleculini: Gonoderina.

Isomiropsis Borchmann, 48 [F]. Type Isomiropsiswittei Borchmann, 1942, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Isoncophallus Koch, 55 [M]. Type Isoncophalluszabroides Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Isonota Fairmaire, 171 [F]. Type Isonotaopaca Fairmaire, 1887, by monotypy. Status: junior synonym of Homoeonota Fairmaire, 1882 in Pimeliinae: Tentyriini. Synonymy: Koch (1962c: 254).

Isopedus Stein in Stein and Weise, 121 [M]. Type Helopstenebrioides Germar, 1813, by present designation. Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: Heyden et al. (1883: 136, with Omaleis Allard, 1876, a synonym of Odocnemis Allard, 1876). Note: the original combination of the name of the type species, Helopstenebrioides Germar, 1813, is a junior primary homonym of Helopstenebrioides Palisot de Beauvois, 1812.

Isopteron Hope, 112 [N]. Type Isopteronaustrale Hope, 1841, by original designation. Status: valid genus in Lagriinae: Adeliini. Note: combined description of a new genus-group taxon and a single new species (ICZN 1999, Article 12.2.6).

Isopteroplonyx Bremer, 7 [M]. Type Amarygmustermitophilus Lea, 1910, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Isopterum Agassiz, 197 [N]. Type species [automatic]: Isopteronaustrale Hope, 1841, by original designation. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Note: unjustified emendation of Isopteron Hope, 1841, not in prevailing usage.

Isopus Montrouzier, 299 [M]. Type Isopusblanchardi Montrouzier, 1860, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Isostira Pascoe, 97 [F]. Type Isostiracrenata Pascoe, 1870, by monotypy. Status: junior synonym of Bradymerus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1921a: 253).

Isotoma Blanchard, pl. 15 [F]. Type Isotomaemarginaticollis Blanchard, 1842, by monotypy. Status: senior synonym of Isocera Borchmann, 1909 in Lagriinae: Lagriini: Statirina. Note: junior homonym of Isotoma Bourlet, 1839 [Collembola].

Itagonia Reitter, 362 [F]. Type Itagoniagnaptorinoides Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Italohelops Español, 295 [M]. Type Parablopssubchalybaeus Reitter, 1907, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Italomelia Mas-Peinado, Buckley, Ruiz & García-París, 539 [F]. Type Pimeliarugulosa Germar, 1823, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Itampolis Koch, 31, 143 [F]. Type Itampolisoceanica Koch, 1962, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Iugidorsum Louw, 99, 102 [N]. Type Iugidorsumcumstriis Louw, 1979, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Ixalus Gistel, x [M]. Type species [automatic]: Blapsexcavata Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: junior synonym of Platynotus Fabricius, 1801 in Blaptinae: Platynotini: Platynotina. Note: unnecessary replacement name for Platynotus Fabricius, 1801.

Jaklia Novák, 180 [F]. Type Jakliaserraticornis Novák, 2010, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Japetus Reitter, 160 [M]. Type Japetusmelanarius Reitter, 1904 (= Opatrummelanarium Erichson, 1843), by monotypy. Status: junior synonym of Trichosternum Wollaston, 1861 in Blaptinae: Opatrini: Opatrina. Synonymy: Gebien (1905: 253, with Trichopodus [as Trichopodum] Mulsant & Rey, 1859, a senior synonym of Trichosternum Wollaston, 1861). Note: junior homonym of Japetus Stål, 1863 [Hemiptera].

Javamarygmus Pic, 22 [M]. Type Javamarygmustristis Pic, 1928, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Jintaium Ren in Ren and Yu, 228 [N]. Type Jintaiumsulcatum Ren, 1999, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Jophon Champion, 224 [M]. Type Jophonmyrmecophilus Champion, 1895, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: Jophon is an incorrect subsequent spelling of the original spelling Iophon, first used by Kolbe (1898: 601), in prevailing usage; Jophon is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1), see Matthews and Bouchard (2008: 330).

Julogenius Reitter, 138 [M]. Type Heliotaurusreichii Mulsant, 1856, by subsequent designation (Novák and Pettersson 2008: 333). Status: valid subgenus of Heliotaurus Mulsant, 1856 in Alleculinae: Cteniopodini.

†Jurallecula L.N. Medvedev, 124 [F]. Type Juralleculagrossa L.N. Medvedev, 1969, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: described from Upper Jurassic deposits (Kazakhstan).

Kabakoviella Kaszab, 205 [F]. Type Kabakoviellamenephiloides Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Kaindilagria Merkl, 138 [F]. Type Kaindilagriaforcipata Merkl, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Karroocara Koch, 176 [N]. Type Stenocaragibbipenne Haag-Rutenberg, 1875, by original designation. Status: junior synonym of Stenodesia Reitter, 1916 in Pimeliinae: Adesmiini. Synonymy: Penrith (1979: 70, 85).

Kaszaba Matthews & Doyen, 40 [F]. Type Tenebriocorvinus Erichson, 1842, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Kaszabadelium Watt, 29 [N]. Type Adeliumaucklandicum Broun, 1880, by original designation. Status: valid genus in Lagriinae: Adeliini.

Kaszabiella Koch, 774, 885 [F]. Type Thalpophilasubhemisphaerica Koch, 1943, by monotypy. Status: valid subgenus of Thalpophilodes Strand, 1942, in Pimeliinae: Tentyriini.

Kaszabochillus Fouquè, 228, 240 [M]. Type Indochillusandamanus Kaszab, 1981, by original designation. Status: valid subgenus of Pseudochillus Fouquè, 2015 in Pimeliinae: Stenosini: Dichillina.

Kaszaboscelis Löbl & Merkl, 246 [F]. Type Tenebriohypolithus Pallas, 1781, by original designation. Status: junior synonym of Platyscelis Latreille, 1818 in Blaptinae: Platyscelidini. Synonymy: Egorov (2004: 605).

Kaszabus Freude, 194 [M]. Type Kaszabusaurulentiformis Freude, 1967, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Kawiria Schuster, 26 [F]. Type Kawiriagabrieli Schuster, 1935, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Kershawia Lea, 379 [F]. Type Kershawiarugiceps Lea, 1905, by monotypy. Status: valid genus in Lagriinae: Belopini.

Kirgisomira Weise, 71 [F]. Type Isomiraophthalmica Seidlitz, 1896, by original designation. Status: junior synonym of Asiomira Dubrovina, 1973 in Alleculinae: Alleculini: Gonoderina. Synonymy: Muche (1981: 157).

Klapperichia Kaszab, 249 [F]. Type Klapperichiamirabilis Kaszab, 1954 (= Tenebriocephalonthoracicum Pic, 1925), by original designation. Status: junior synonym of Tenebriocephalon Pic, 1925 in Pimeliinae: Ceratanisini. Synonymy: Kaszab (1983a: 130).

Klewaria Reitter, 20 [F]. Type Klewariacolydiiformis Reitter, 1910, by monotypy. Status: valid genus in Pimeliinae: Klewariini.

Knausia Fall, 15 [F]. Type Knausiacrassicornis Fall, 1931, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Kocakia Kaszab, 54 [F]. Type species [automatic]: Idiopsisopaca Kaszab, 1981, by original designation. Status: valid genus in Pimeliinae: Edrotini. Note: replacement name for Idiopsis Kaszab, 1981.

Kocheria Antoine, 25, 30 [F]. Type Arthrodeisrungsi Español, 1943, by monotypy. Status: valid subgenus of Arthrodeis Solier, 1834 in Pimeliinae: Erodiini.

Kochogaster Kamiński & Raś, 654 [F]. Type species [automatic]: Anchophthalmusimpressicollis Fairmaire, 1897, by original designation. Status: valid subgenus of Anchophthalmus Gerstaecker, 1854 in Blaptinae: Platynotini: Platynotina. Note: replacement name for Cosmogaster Koch, 1956.

Kochotella Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Millotellamicrocornis Koch, 1962, by original designation. Status: valid genus in Pimeliinae: Asidini. Note: replacement name for Millotella Koch, 1962.

Kokeniella Reitter, 41 [F]. Type Kokeniellamesostenoides Reitter, 1906, by subsequent designation (Gebien 1937a: 626). Status: valid genus in Pimeliinae: Tentyriini.

Kombacula Novák, 271 [F]. Type Kombaculakantneri Novák, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Koneus Giraldo-Mendoza & Flores, 94 [M]. Type Prohylithusperuanus Kaszab, 1981, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Kralia Novák, 500 [F]. Type Kraliaminshanica Novák, 2013, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Krollus R. Lucas, 357 [M]. Type species [automatic]: Homoeogenuslaticornis C.O. Waterhouse, 1882, by monotypy. Status: junior synonym of Borneocamaria Pic, 1917 in Stenochiinae: Cnodalonini. Note: replacement name for Homoeogenus C.O. Waterhouse, 1882.

Ksukolcula Novák, 168 [F]. Type Ksukolculahesperia Novák, 2017, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Kuhitangia G.S. Medvedev, 1184 [F]. Type Kuhitangiakryzhanovskii G.S. Medvedev, 1962, by original designation. Status: valid genus in Kuhitangiinae: Kuhitangiini.

Kuschelus Kaszab, 112 [M]. Type Kuscheluslathridioides Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Lupropini.

Labetis C.O. Waterhouse, 267 [F]. Type Labetistibialis C.O. Waterhouse, 1879, by monotypy. Status: valid genus in Alleculinae: Alleculini: Mycetocharina.

Lachna Billberg, 35 [F]. Type Chrysomelahirta Linnaeus, 1758, by subsequent designation (Merkl 2006: 223). Status: junior synonym of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina. Synonymy: Lawrence and Newton (1995: 893).

Lachnodactylus Seidlitz, 838 [M]. Type species [automatic]: Lachnopusdigitalis Seidlitz, 1894, by monotypy. Status: valid genus in Pimeliinae: Lachnogyini: Lachnodactylina. Note: replacement name for Lachnopus Seidlitz, 1894.

Lachnoderes Mulsant & Rey, 70, 102 [M]. Type Pedonoecespubescens G.R. Waterhouse, 1845, by monotypy. Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Gemminger in Gemminger and Harold (1870: 1923, with Pedonoeces G.R. Waterhouse, 1845, a junior synonym of Blapstinus Dejean, 1821); Aalbu and Triplehorn (1991: 170).

Lachnogya Ménétriés, 228 [F]. Type Lachnogyasquamosa Ménétriés, 1849, by monotypy. Status: valid genus in Pimeliinae: Lachnogyini: Lachnogyina.

Lachnopus Seidlitz, 476 [M]. Type Lachnopusdigitalis Seidlitz, 1894, by monotypy. Status: senior synonym of Lachnodactylus Seidlitz, 1898 in Pimeliinae: Lachnogyini: Lachnodactylina. Note: junior homonym of Lachnopus Schönherr, 1840 [Coleoptera: Curculionidae].

Laena Dejean, 64 [F]. Type fixed herein (ICZN 1999, Article 70.3) as Scaurusviennensis Sturm, 1807, misidentified as Helopspimelia Fabricius, 1787 in the original designation by monotypy in Dejean (1821). Status: valid genus in Lagriinae: Laenini. Note: the type species “Helopspimelia Fabricius” was first established by monotypy (ICZN 1999, Article 68.3); Bouchard and Bousquet (2020b: 7) noted that the only valid species originally included in Laena, Helopspimelia Fabricius, 1787, is currently considered a valid species in the genus Penthe Newman, 1838 [Coleoptera: Tetratomidae] and therefore the taxonomic species involved, Helopspimelia Fabricius sensu Dejean 1821 (= Scaurusviennensis Sturm, 1807) is fixed here as the type species of Laena Dejean, 1821 according to the requirements of Article 70.3.2 (ICZN 1999).

Lagria Fabricius, 124 [F]. Type Chrysomelahirta Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Lagriinae: Lagriini: Lagriina.

Lagriallecula Pic, 19 [F]. Type Lagrialleculaaeneipennis Pic, 1920, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Lagriella Borchmann, 61 [F]. Type Lagriamima Borchmann, 1916, by original designation. Status: valid subgenus of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina.

Lagrimina Fairmaire, 675 [F]. Type Lagriminastrigipennis Fairmaire, 1894 (= Porrolagrianuda Kolbe, 1883), by monotypy. Status: junior synonym of Porrolagria Kolbe, 1883 in Lagriinae: Lagriini: Lagriina. Synonymy: Kolbe (1897a: 250).

Lagriocera Fairmaire, 41 [F]. Type Lagrioceracavicornis Fairmaire, 1896, by monotypy. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Lagriodema Borchmann, 442, 524 [F]. Type Nemostiragestroi Borchmann, 1910, by subsequent designation (Borchmann 1936: 458). Status: valid genus in Lagriinae: Lagriini: Statirina.

Lagriodes Borchmann, 405, 432 [M]. Type Heterogriaarmigera Borchmann, 1930, by monotypy. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Lagriogonia Fairmaire, ccxvii [F]. Type Lagriogoniahumerosa Fairmaire, 1891, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lagriola Kirsch, 409 [F]. Type Lagriolaoperosa Kirsch, 1874, by subsequent designation (Bousquet et al. 2018: 33). Status: junior synonym of Paratenetus Spinola, 1845 in Lagriinae: Goniaderini. Synonymy: Gebien (1922b: 268), Matthews and Lawrence (2015: 311).

Lagriomima Pic, 31 [F]. Type Nemostiramaxima Pic, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lagriomima Pic, 11 [F]. Type Lagriomimaalbolineata Pic, 1950, by monotypy. Status: junior synonym of Neogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina. Synonymy: new synonym [OM]. Note: based on the description, the type species belongs to Neogria Borchmann, 1911 and is probably synonymous with the valid species Neogriacyanipennis Borchmann, 1911, therefore, Lagriomima Pic, 1950 is proposed as a new synonym of Neogria; junior homonym of Lagriomima Pic, 1934 [Coleoptera: Tenebrionidae: Lagriinae: Lagriini: Statirina].

Lagriopsis Borchmann, 49, 138 [F]. Type Lagriopsisinsularis Borchmann, 1916, by subsequent designation (Borchmann 1936: 146). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Lagriostira Fairmaire, 103 [F]. Type Statirarufonitens Fairmaire, 1883, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lagriostira Kolbe, 550 [F]. Type Lagriostirahispida Kolbe, 1902 (= Lagriasubseriata Reitter, 1880), by subsequent designation (Borchmann 1936: 224). Status: senior synonym of Alagria Borchmann, 1916 in Lagriinae: Lagriini: Lagriina. Note: junior homonym of Lagriostira Fairmaire, 1883 [Coleoptera: Tenebrionidae: Lagriinae: Lagriini: Statirina].

Lamperos Allard, 4 [M]. Type Helopsmicans Fabricius, 1798, by subsequent designation (Nabozhenko and Löbl 2008: 256). Status: junior synonym of Tarpela Bates, 1870 in Tenebrioninae: Helopini: Helopina. Synonymy: Champion (1887: 288).

Lamprobothris Fairmaire, 302 [F]. Type Lamprobothrisattenuata Fairmaire, 1887 (= Praogenafossulata Müller, 1887), by monotypy. Status: junior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Synonymy: De Moor (1970: 42).

Lamprocrypticus Español, 127 [M]. Type Crypticusalpinus Comolli, 1837, by original designation. Status: valid genus in Diaperinae: Crypticini.

Lanhsia Shibata, 63, 64 [F]. Type Lanhsiabucca Shibata, 1980, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Laonicus Haag-Rutenberg, 100 [M]. Type Laonicuspilosus Haag-Rutenberg, 1878, by subsequent designation (R. Lucas 1920: 361). Status: junior synonym of Platyphanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 454).

Laoscapha Schawaller, 440 [F]. Type Laoscaphaphoupanensis Schawaller, 2016, by original designation. Status: valid genus in Diaperinae: Scaphidemini.

Laosocryptobates Pic, 25 [M]. Type Laosocryptobatestuberculatus Pic, 1928, by monotypy. Status: junior synonym of Hexarhopalus Fairmaire, 1891 in Stenochiinae: Cnodalonini. Synonymy: Bečvář and Purchart (2008: 39).

Laraliprosodes Bogatchev, 513 [M]. Type Prosodeslar Bogatchev, 1947, by original designation. Status: junior synonym of Dineria Motschulsky, 1860 in Blaptinae: Blaptini: Blaptina. Synonymy: G.S. Medvedev and Iwan (2006: 617).

Lariversius Blaisdell, 59 [M]. Type Lariversiustibialis Blaisdell, 1947, by original designation. Status: valid genus in Blaptinae: Amphidorini.

Lasiocnema G.S. Medvedev, 110 [F]. Type Lasiostolaheterogena Fischer von Waldheim, 1844, by original designation. Status: valid subgenus of Lasiostola Dejean, 1834 in Pimeliinae: Pimeliini.

Lasioderus Mulsant & Rey, 13, 42 [M]. Type Lasioderussulcipennis Mulsant & Rey, 1854, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Lasiograna G.S. Medvedev, 111 [F]. Type Lasiostolainterrupta Reitter, 1901, by original designation. Status: valid subgenus of Lasiostola Dejean, 1834 in Pimeliinae: Pimeliini.

Lasiostola Dejean, 179 [F]. Type Tenebriopubescens Pallas, 1781, by subsequent designation (Hope 1841: 118). Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Latacula Campbell, 103 [F]. Type Lataculabeckeri Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Latetribolium Lepesme, 46 [N]. Type Triboliumrisbeci Lepesme, 1943 (= Opatrumlaevigatum Fabricius, 1781), by monotypy. Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Hinton (1948: 13, through synonymy of its type species with Opatrumlaevigatum Fabricius, 1781).

Latheticus C.O. Waterhouse, 147 [M]. Type Latheticusoryzae C.O. Waterhouse, 1880, by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Latipleurosis Penrith, 19, 204 [F]. Type Zophosisbenguelensis Deyrolle, 1867, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Latorhascius Pic, 8 [M]. Type Rhaciusbaeri Pic, 1925, by monotypy. Status: valid subgenus of Adelonia Laporte, 1840 in Lagriinae: Belopini.

Lawrenceus Iwan, 307 [M]. Type Lawrenceuscapensis Iwan, 1998, by original designation. Status: junior synonym of Schelodontes Koch, 1956 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan and Kamiński (2014: 171).

Leanum Uyttenboogaart, 29, 31 [N]. Type Triboliummyrmecophilum Lea, 1904, by monotypy. Status: junior synonym of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini. Synonymy: Hinton (1948: 21).

Leaus Matthews & Lawrence, 312 [M]. Type Leaustasmanicus Matthews & Lawrence, 1992, by original designation. Status: valid genus in Tenebrioninae: Trachelostenini. Note: transferred from Titaenini by Matthews and Lawrence (2015: 293).

Lechinius Blair, 561 [M]. Type Lechiniuscatenulatus Blair, 1922 (= Cistelomorphafossulata Pic, 1913), by monotypy. Status: valid subgenus of Cteniopinus Seidlitz, 1896 in Alleculinae: Cteniopodini.

Lechinius Borchmann, 151 [M]. Type Cistelomorphafossulata Pic, 1913, by subsequent designation (Novák and Pettersson 2008: 330). Status: junior synonym of Lechinius Blair, 1922 in Alleculinae: Cteniopodini. Note: junior homonym of Lechinius Blair, 1922 [Coleoptera: Alleculinae: Cteniopodini].

Lechius Iwan, 404 [M]. Type Selinusabacoides Fairmaire, 1902, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Lechriomus Morawitz, 21 [M]. Type Akislucifuga Adams, 1817, by subsequent designation (Chernei 102). Status: valid subgenus of Cyphogenia Solier, 1837 in Pimeliinae: Akidini.

Leichenum Dejean, 194 [N]. Type Opatrumpictum Fabricius, 1801, by monotypy. Status: valid genus in Blaptinae: Pedinini: Leichenina.

Leichrodomorphus Pic, 6 [M]. Type Leichrodomorphusbrevicornis Pic, 1921, by subsequent designation (Pic 1922e: 135). Status: junior synonym of Stethotrypes Gebien, 1914 in Diaperinae: Leiochrinini. Synonymy: Gebien (1940: 431).

Leiochrinus Westwood, 68 [M]. Type Leiochrinusfulvicollis Westwood, 1883, by subsequent designation (W.F. Kirby 1885a: 89). Status: valid genus in Diaperinae: Leiochrinini.

Leiochrodes Westwood, 69 [M]. Type Leiochrodesdiscoidalis Westwood, 1883, by subsequent designation (W.F. Kirby 1885a: 89). Status: junior synonym of Ades Guérin-Méneville, 1857 in Diaperinae: Leiochrinini. Synonymy: Gebien (1911a: 388).

Leiochrodinus Kaszab, 358, 365 [M]. Type Leiochrodinustetraphyllus Kaszab, 1961, by original designation. Status: valid genus in Diaperinae: Leiochrinini. Note: the alternative original spelling Leichrodinus, used by Kaszab (1961a: 365), was rejected by Papp and Seeno (1981: 53, 71) who acted as the First Revisers.

Leiochrodontes Kaszab, 30, 200 [M]. Type Leiochrodesmadurensis Pic, 1918, by original designation. Status: valid genus in Diaperinae: Leiochrinini. Note: the earlier usage of Leiochrodontes by Gebien (1940: 755) was not accompanied by a description, a definition or a bibliographic reference to such a published statement and therefore is not available from that date.

Leiochromimus Ardoin, 187, 199 [M]. Type Coccimarygmuspunctatus Ardoin, 1966, by monotypy. Status: valid subgenus of Coccimarygmus Ardoin, 1966 in Tenebrioninae: Amarygmini.

Leiochrota Westwood, 70 [F]. Type Leiochrinusuniformis Westwood, 1883, by subsequent designation (W.F. Kirby 1885a: 90). Status: valid genus in Diaperinae: Leiochrinini.

Leiochrotina Westwood, 70, 76 [F]. Type Leiochrinusindicus Westwood, 1883, by monotypy. Status: junior synonym of Crypsis C.O. Waterhouse, 1877 in Diaperinae: Leiochrinini. Synonymy: Gebien (1940: 436).

Leiopeplus Broun, 1160 [M]. Type Helopsexpolitus Broun, 1880, by original designation. Status: senior synonym of Chrysopeplus Gebien, 1942 in Stenochiinae: Cnodalonini. Note: junior homonym of Leiopeplus Murray, 1864 [Coleoptera: Nitidulidae].

Leipopleura Seidlitz, 342, 345 [F]. Type Faustiaintegra Reitter, 1887, by subsequent designation (Kaszab 1940a: 167). Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Lelegeis Champion, 209 [M]. Type Lelegeisaeneipennis Champion, 1886, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Leleupium Kaszab, 106 [N]. Type Leleupiumsubcoecum Kaszab, 1956, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Lemoultia Chatanay, 313 [F]. Type Lemoultiascabripennis Chatanay, 1913, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Lenkous Kaszab, 315 [M]. Type Lenkousmyrmecophilus Kaszab, 1973, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Lepidocaulinus Schawaller, Masumoto & Merkl, 378 [M]. Type Lepidocaulinusmirabilis Schawaller, Masumoto & Merkl, 2013, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Lepidochora Koch, 36 [F]. Type Lepidochoraeberlanzi Gebien, 1938, by original designation. Status: valid genus in Pimeliinae: Adelostomini. Note: the earlier usage of Lepidochora by Gebien (1938b: 71) is unavailable since it was published after 1930 without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Lepidocnemeplatia Bousquet & Bouchard in Bousquet et al., 2018: 20 [F]. Type Cnemeplatiasericea Horn, 1870, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Note: Lepidocnemeplatia was previously described by Kaszab (1938: 80) without original type species fixation and is therefore unavailable from that date (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 245) designated Cnemeplatiasericea Horn, 1870 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Lepidospilus Agassiz, 205 [M]. Type species [automatic]: Pachycoeliasulcicollis Boisduval, 1835, by monotypy. Status: junior synonym of Pachycoelia Boisduval, 1835 in Tenebrioninae: Heleini: Cyphaleina. Note: unjustified emendation of Lepispilus Westwood, 1841, not in prevailing usage.

Lepispilus Westwood, 44 [M]. Type Pachycoeliasulcicollis Boisduval, 1835, by monotypy. Status: junior synonym of Pachycoelia Boisduval, 1835 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Lacordaire (1859b: 413).

Leprocaulinus Kaszab, 75 [M]. Type Leprocaulinuskrikkeni Kaszab, 1982, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: this genus-group name has been considered a junior homonym of Leprocaulinus Uvarov, 1940 [Orthoptera] in literature on Tenebrionidae; however, no type species has yet been designated for orthopteran name Leprocaulus Redtenbacher, 1908 nor its replacement name Leprocaulinus Uvarov, 1940 (see Herwaarden 1998: 65) and therefore Leprocaulinus Uvarov, 1940 is unavailable since it has been proposed as a replacement name after 1930, without a type species designation, for a generic taxon without valid typification (ICZN 1999: Article 13.3.1).

Leprocaulus Fairmaire, 95 [M]. Type Leprocaulusclavipes Fairmaire, 1896, by monotypy. Status: valid subgenus of Hexarhopalus Fairmaire, 1891 in Stenochiinae: Cnodalonini.

Leptasida Chatanay, 3 [F]. Type Leptasidatenuipes Chatanay, 1914, by original designation. Status: valid genus in Pimeliinae: Asidini.

Leptasida Reitter, 40, 60 [F]. Type Asidadiecki Allard, 1869, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 291). Note: junior homonym of Leptasida Chatanay, 1914 [Coleoptera: Tenebrionidae: Pimeliinae: Asidini].

Leptinostethus Borchmann, 238, 466 [M]. Type Leptinostethusmethneri Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Leptocolena Allard, 74 [F]. Type Blapsmucronata Latreille, 1804, by subsequent designation (Harold 1882: 194). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gebien (1910b: 226).

Leptoderis Billberg, 31 [F]. Type Tenebriocollaris Linnaeus, 1767, by monotypy. Status: valid genus in Pimeliinae: Elenophorini: Elenophorina.

Leptoderops Löbl, Bouchard, Merkl & Bousquet, 5 [M]. Type Thraustocoluspriesneri Koch, 1934, by original designation. Status: valid subgenus of Thraustocolus Kraatz, 1866 in Pimeliinae: Tentyriini. Note: name first proposed by Koch (1934: 96) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl et al. (2008a: 40) designated Thraustocoluspriesneri Koch, 1934 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Leptodes Dejean, 181 [M]. Type Sepidium boisduvalii Zubkov, 1833, by monotypy. Status: valid genus and subgenus in Pimeliinae: Leptodini.

Leptodinopsis Kaszab, 353, 359 [F]. Type Sepidium boisduvalii Zubkov, 1833, by original designation. Status: junior synonym of Leptodes Dejean, 1834 in Pimeliinae: Leptodini. Synonymy: G.S. Medvedev and Iljina (2007: 882).

Leptodopsis Haag-Rutenberg, 408 [F]. Type Leptodopsisinsignis Haag-Rutenberg, 1879, by monotypy. Status: valid subgenus of Leptodes Dejean, 1834 in Pimeliinae: Leptodini.

Leptogastrus W.J. MacLeay, 293 [M]. Type Leptogastrusmastersii W.J. MacLeay, 1872, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Leptomorpha Faldermann, 406 [F]. Type Leptomorphachinensis Faldermann, 1835, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Champion (1895: 48).

Leptonychoides Schawaller, 51 [M]. Type Leptonychoidesjuengeri Schawaller, 1990, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Leptonychus Chevrolat, 26, pl. 1 [M]. Type Leptonychuserodioides Chevrolat, 1833, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Leptoscapha Fairmaire, 73 [F]. Type species [automatic]: Stenoscaphaspissicornis Fairmaire, 1885, by monotypy. Status: senior synonym of Brachypophlaeus Fairmaire, 1897 in Tenebrioninae: Ulomini. Synonymy: Alluaud (1899: 342, as “Brachypophloeus”). Note: replacement name for Stenoscapha Fairmaire, 1885; junior homonym of Leptoscapha Fischer, 1883 [Mollusca].

Leptosora Borchmann, 238, 471 [F]. Type Leptosorahamata Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Leptosphena Semenov, 356, 358 [F]. Type Sphenariatomentosa Semenov, 1889 (= Sphenariarubripes Reitter, 1889), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Lepturidea Fauvel, 150 [F]. Type Lepturideadeplanchei Fauvel, 1862, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Leptynoderes Solier, 8, 44 [M]. Type Scotobiusvaricosus Germar, 1823, by original designation. Status: valid genus in Tenebrioninae: Scotobiini.

Lesbidana Reitter, 173 [F]. Type Melanesthessimplex Reitter, 1897, by subsequent designation (G.S. Medvedev 1990: 203). Status: valid subgenus of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina.

Leucolaephus P.H. Lucas, xxii [M]. Type Leucolaephusperrisii P.H. Lucas, 1859 (= Pimelialiliputana P.H. Lucas, 1857), by subsequent designation (Gebien 1937a: 803). Status: valid genus in Pimeliinae: Pimeliini. Note: Leucolaephus is an incorrect subsequent spelling of the original spelling Leucoloephus, first used by Kraatz (1865: 314), and is in prevailing usage; Leucolaephus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1), see Bouchard et al. (2005: 506).

Lichenum Agassiz, 203, 209 [N]. Type species [automatic]: Opatrumpictum Fabricius, 1801, by monotypy. Status: junior synonym of Leichenum Dejean, 1834 in Blaptinae: Pedinini: Leichenina. Note: unjustified emendation of Leichenum Dejean, 1834, not in prevailing usage.

Licinoma Pascoe, 140 [F]. Type Licinomanitida Pascoe, 1869, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Licymnius Bates, 271 [M]. Type Licymniusfoveicollis Bates, 1868, by monotypy. Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Carter (1915a: 60, with Chromomaea Pascoe, 1866, a junior synonym of Lepturidea Fauvel, 1862).

Lindia Blackburn, 275 [F]. Type Lindiaangusta Blackburn, 1888, by monotypy. Status: junior synonym of Lyphia Mulsant & Rey, 1859 in Tenebrioninae: Triboliini. Synonymy: Champion (1894a: 351, 370). Note: junior homonym of Lindia Dujardin, 1841 [Rotifera].

Lineocrypticus Koch, 52 [M]. Type Lineocrypticushessei Koch, 1950, by original designation. Status: valid genus in Diaperinae: Crypticini.

Linio Bouchard & Bousquet, new subgenus [M]. Type Niliolanatus Germar, 1823, by present designation. Status: valid subgenus of Nilio Latreille, 1802 in Nilioninae. Note: Mader (1936: 94) introduced the new subgenus name Linio for several nominal species, but unfortunately did not designate a type species; the subgenusLinio, which has been treated as valid since 1936, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Niliolanatus Germar, 1823 as type species and referring to Mader (1936: 94) for the character states that characterise and differentiate Linio.

Liodema Horn, 378, 385 [F]. Type Platydemalaevis Haldeman, 1848, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Liodocistela Pic, 28 [F]. Type Cistelopsisrufomarginata Pic, 1930, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Lioprosodes Reitter, 121 [M]. Type Prosodesdilaticollis Motschulsky, 1859, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Skopin (1960a: 46).

Lisa Haag-Rutenberg, 134 [F]. Type Lisasingularis Haag-Rutenberg, 1879 (= Alleculaomophloides Hope, 1843), by monotypy. Status: junior synonym of Metistete Pascoe, 1866 in Alleculinae: Alleculini: Alleculina. Synonymy: Carter (1915a: 78).

Litasida Casey, 77, 184 [F]. Type Litasidatownsendi Casey, 1912, by original designation. Status: valid genus in Pimeliinae: Asidini.

Litheleodes Blaisdell, 114 [M]. Type Blaps extricata Say, 1824, by subsequent designation (Triplehorn and Thomas 2015: 11). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Lithoblaps Motschulsky, 532 [F]. Type Tenebriogigas Linnaeus, 1767, by subsequent designation (Skopin, 44). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860). Note: the original combination of the name of the type species, Tenebriogigas Linnaeus, 1767, is a junior primary homonym of Tenebriogigas Linnaeus, 1763.

Litoboriolus Español, 310, 313 [M]. Type Olocratescollaris Mulsant & Rey, 1854, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Litoboromimus Koch, 413 [M]. Type Litoborusparallelus Schuster, 1919, by original designation. Status: valid subgenus of Allophylax Bedel, 1906 in Blaptinae: Dendarini: Melambiina.

Litoborus Mulsant & Rey, 124, 126 [M]. Type Phylaxmoreletii P.H. Lucas, 1846, by subsequent designation (Antoine 1931: 181). Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Litopous Matthews, 1 [M]. Type Litopousbaehri Matthews, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: placed in the subtribe Alleculina by Matthews and Lawrence (2019: 645).

Litororus Reitter, 96 [M]. Type Micrositussemicostatus Mulsant & Rey, 1854, by monotypy. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Lixionica Blackburn, 280 [F]. Type Lixionicacostatipennis Blackburn, 1896, by monotypy. Status: valid genus in Pimeliinae: Vacronini.

Lobatopezus Pic, 2 [M]. Type Lobatopezusdiversecostatus Pic, 1952, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: Bremer (2005: 211) transferred this genus to Chrysomeloidea: Chrysomelidae, but later retained this genus in Tenebrioninae: Amarygmini after further study (Bremer 2013b: 127–128).

Lobetas Motschulsky, 26 [M]. Type Tenebriocostatus Guérin-Méneville, 1831, by original designation. Status: junior synonym of Hipalmus Bates, 1870 in Tenebrioninae: Tenebrionini. Synonymy: C.O. Waterhouse (1876: 288). Note: the original combination of the name of the type species, Tenebriocostatus Guérin-Méneville, 1831, is a junior primary homonym of Tenebriocostatus Pallas, 1781.

Lobodera Mulsant & Rey, 12, 18 [F]. Type Loboderarufescens Mulsant & Rey, 1859, by monotypy. Status: junior synonym of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Synonymy: Gebien (1910b: 333).

Lobometopon Casey, 379, 385 [N]. Type Epitragusfusiformis Casey, 1890, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Lobophilomorphus Pic, 183 [M]. Type Lobophilomorphusrubicollis Pic, 1911, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lobopoda Solier, 233 [F]. Type Lobopodastriata Solier, 1835, by subsequent designation (Bousquet et al. 2015: 134). Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Lobothorax Gemminger, 124 [M]. Type species [automatic]: Loboderarufescens Mulsant & Rey, 1859, by monotypy. Status: junior synonym of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Note: unnecessary replacement name for Lobodera [as “Loboderus”] Mulsant & Rey, 1859.

Locrodes Casey, 332 [M]. Type Emmenastuspiceus Casey, 1890, by subsequent designation (Bousquet et al. 2018: 104). Status: valid subgenus of Hylocrinus Casey, 1907 in Pimeliinae: Edrotini.

Lodinus Mulsant & Rey, 113, 131 [M]. Type Lodinusnigroaeneus Mulsant & Rey, 1859 (= BlapstinuspunctulatusSolier 1851), by monotypy. Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Gemminger in Gemminger and Harold (1870: 1923).

Loensus R. Lucas, 380, 489 [M]. Type species [automatic]: Pedinopsispilipes Gebien, 1910, by monotypy. Status: valid genus and subgenus in Blaptinae: Pedinini: Pedinina. Note: replacement name for Pedinopsis Gebien, 1910.

Lomocnemis Gebien, 287 [F]. Type Lomocnemispolita Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Longuloodescelis Kaszab, 940, 957 [F]. Type Platyscelishirta Seidlitz, 1893, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini. Note: the First Reviser (Longuloodescelis Kaszab, 1940 versus Trichoodescelis Kaszab, 1940) is Egorov (2004: 595).

Lophocnemis Mäklin, 505 [F]. Type Lophocnemisamabilis Mäklin, 1867, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Lopholagria Borchmann, 48, 97 [F]. Type Lagriaamoena Fåhraeus, 1870, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Lagriina.

Lophoma Solier, 253, 285 [N]. Type Pimeliapunctata Fabricius, 1798, by monotypy. Status: junior synonym of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Lacordaire (1859a: 46). Note: the original combination of the name of type species, Pimeliapunctata Fabricius, 1798, is a junior primary homonym of Pimeliapunctata Thunberg, 1787.

Lophophyllus Fairmaire, 71 [M]. Type Lophophylluscostipennis Fairmaire, 1887, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lordodera Gebien, 64 [F]. Type Tenebrioquadrihamatus Fairmaire, 1875, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Lorelopsis Champion, 15 [F]. Type Lorelopsispilosa Champion, 1896, by monotypy. Status: junior synonym of Lorelus Sharp, 1876 in Lagriinae: Lupropini. Synonymy: Doyen (1993: 295).

Lorelus Sharp, 76 [M]. Type Loreluspriscus Sharp, 1876, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Loricula Novák, 45 [F]. Type Alleculasubaeneipennis Pic, 1922, by original designation. Status: senior synonym of Loriculoides Novák, 2020 in Alleculinae: Alleculini: Alleculina. Note: junior homonym of Loricula Curtis, 1833 [Hemiptera].

Loriculoides Novák, 9 [M]. Type species [automatic]: Alleculasubaeneipennis Pic, 1922, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: replacement name for Loricula Novák, 2016.

Lornamus Koch, 191 [M]. Type Lornamusdividiopsis Koch, 1952, by original designation. Status: junior synonym of Cryptocarpes Koch, 1952 in Pimeliinae: Caenocrypticini. Synonymy: Endrödy-Younga (1996: 21).

Lorona Borchmann, 16, 62 [F]. Type Lagriabakeri Borchmann, 1930, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Loubacantus Bonadona, 1033 [M]. Type Loubacantusgiganteus Bonadona, 1959, by original designation. Status: junior synonym of Entypodera Gerstaecker, 1871 in Lagriinae: Lagriini: Statirina. Synonymy: Bonadona (1984: 469).

Louwerensia Kaszab, 104 [F]. Type Louwerensiapapuana Kaszab, 1964, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Loxostethus Triplehorn, 504 [M]. Type Loxostethusfasciatus Triplehorn, 1962, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: new placement [RLA], previously included in Diaperinae: Diaperini: Adelinina.

Lucidolaena Endrödy-Younga & Schawaller, 9, 20 [F]. Type Lucidolaenaamatolensis Endrödy-Younga & Schawaller, 2002, by original designation. Status: valid genus in Lagriinae: Laenini.

Luebbertia Koch, 20, 44 [F]. Type Luebbertiaplana Koch, 1963, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Luprops Hope, 63 [M]. Type Lupropschrysophthalmus Hope, 1833 (= Tageniaindica Wiedemann, 1823), by monotypy. Status: valid genus in Lagriinae: Lupropini.

Luzonoplonyx Bremer, 331 [M]. Type Luzonoplonyxgrandis Bremer, 2009, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Lycanthropa J. Thomson, 20 [F]. Type Eurychoracimicoides Quensel, 1806, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Lycidioides Ando, 107 [M]. Type Lycidioideskaniei Ando, 2003, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Lycogonocnemis Pic, 13 [F]. Type Lycogonocnemisrufa Pic, 1915, by monotypy. Status: valid subgenus of Paragonocnemis Kraatz, 1899 in Tenebrioninae: Amarygmini.

Lycoscelis Blair, 242 [F]. Type Lycoscelisfulva Blair, 1929 (= Plinthochrousgounellei Fairmaire, 1891), by original designation. Status: junior synonym of Plinthochrous Fairmaire, 1891 in Tenebrioninae: Amarygmini. Synonymy: Ardoin (1962b: 957).

Lycula Campbell, 30 [F]. Type Lyculachilensis Campbell, 1976, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Lygestira Pascoe, 470 [F]. Type Prophanessimplex Westwood, 1849, by subsequent designation (R. Lucas 1920: 384). Status: junior synonym of Prophanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 488).

Lygophilus Rafinesque, 114 [M]. Type species [automatic]: Epitragusfuscus Latreille, 1804, by subsequent monotypy (Latreille 1804: 322). Status: junior synonym of Epitragus Latreille, 1802 in Pimeliinae: Epitragini. Note: unnecessary replacement name for Epitragus Latreille, 1802.

Lyphia Mulsant & Rey, 166 [F]. Type Lyphiaficicola Mulsant & Rey, 1859 (= Biustetraphyllus Fairmaire, 1857), by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Lyprochelyda Fairmaire, 214 [F]. Type Lyprochelydapurpurina Fairmaire, 1899, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Lyprosodes Reitter, 116 [M]. Type Prosodesquadricostata Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Lystronychus Latreille, 41 [M]. Type Helopsequestris Fabricius, 1775, by subsequent designation (Saunders 1837: 154). Status: valid genus and subgenus in Alleculinae: Alleculini: Xystropodina. Note: Lystronychus is an incorrect subsequent spelling of the original spelling Lystronichus, first used by Perty (1832: 63), in prevailing usage; Lystronychus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Macellocerus Solier, 154, 263 [M]. Type species [automatic]: Dolichoderusacuminatus Klug, 1833, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Nycteropina. Note: replacement name for Dolichoderus Klug, 1833; discovery of the older unused name Dillacerus Solier, 1835 threatens the stability of Macellocerus Solier, 1848; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to conserve usage of Macellocerus Solier, 1848.

Machla Herbst, 152 [F]. Type Opatrumvillosum G.-A. Olivier, 1795, by subsequent designation (R. Lucas 1920: 386). Status: valid genus in Pimeliinae: Asidini. Note: the older name Machla Lichtenstein, 1796 was published in a work that was suppressed for nomenclatural purposes (ICZN 1995, Opinion 1820).

Machlasida Escalera, 336 [F]. Type Asidamuleyhafidi Escalera, 1907, by subsequent designation (Escalera 1928: 137). Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini.

Machleida Fåhraeus, 256 [F]. Type Machleidanodulosa Fåhraeus, 1870, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Machloida Rye, 286 [F]. Type species [automatic]: Machleidanodulosa Fåhraeus, 1870, by monotypy. Status: junior synonym of Machleida Fåhraeus, 1870 in Pimeliinae: Asidini. Note: unjustified emendation of Machleida Fåhraeus, 1870, not in prevailing usage.

Machlomorpha Péringuey, 257 [F]. Type Machlomorphaaltitudinis Péringuey, 1899, by subsequent designation (Gebien 1937a: 739). Status: valid genus and subgenus in Pimeliinae: Asidini.

Machlophila Wilke, 521 [F]. Type Machlophilavogti Wilke, 1924, by monotypy. Status: junior synonym of Machla Herbst, 1799 in Pimeliinae: Asidini. Synonymy: Koch (1962a: 121, 144).

Machloplasta Wilke, 263 [F]. Type Opatrumvillosum G.-A. Olivier, 1795, by original designation. Status: junior synonym of Machla Herbst, 1799 in Pimeliinae: Asidini. Synonymy: Koch (1962a: 119).

Machlopsis Pomel, 236 [F]. Type Eurychoralevaillantii P.H. Lucas, 1850, by original designation. Status: valid genus and subgenus in Pimeliinae: Adelostomini. Note: Machlopsis is an incorrect subsequent spelling of the original spelling Maclopsis, first used by Bedel (1887: 199), in prevailing usage; Machlopsis is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Macradesmia Kaszab, 402 [F]. Type Adesmiaservillei Solier, 1835, by monotypy. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: name first proposed by Koch (1944b: 146) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Macradesmia Löbl & Merkl in Löbl et al., 2020: 1 [F]. Type Pimeliacancellata Klug, 1830, by original designation. Status: junior synonym of Macradesmia Kaszab, 1959 in Pimeliinae: Adesmiini. Note: name first proposed by Koch (1944b: 146) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); not knowing about the availability of Macradesmia Kaszab, 1959, Löbl & Merkl (2003: 244) designated Pimeliacancellata Klug, 1830 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1) and subsequently Macradesmia was proposed as new by Löbl and Merkl in Löbl et al. (2020: 1); junior homonym of Macradesmia Kaszab, 1959 [Coleoptera: Tenebrionidae: Pimeliinae: Adesmiini].

Macrarthra Agassiz, 35, 219 [F]. Type species [automatic]: Arthromacradonacioides W. Kirby, 1837 (= Lagriaaenea Say, 1824), by monotypy. Status: junior synonym of Arthromacra W. Kirby, 1837 in Lagriinae: Lagriini: Statirina. Note: unjustified emendation of Arthromacra W. Kirby, 1837, not in prevailing usage; the original combination of the accepted name of the type species, Lagriaaenea Say, 1824, is a junior primary homonym of Lagriaaenea Fabricius, 1775.

Macroartactes Pic, 24 [M]. Type Macroartactescostulatus Pic, 1924, by subsequent designation (Gebien 1935: 64). Status: junior synonym of Artactes Pascoe, 1868 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1935: 64).

Macrocasnonidea Pic, 31 [F]. Type Nemostiraquadrimaculata Pic, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Macrocistela Pic, 13 [F]. Type Macrocistelastriata Pic, 1941, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Macrocistelopsis Pic, 89 [F]. Type Macrocistelopsistestaceicolor Pic, 1956, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Macrohyperchalca Pic, 98 [F]. Type Hyperchalcastriata Pic, 1935, by monotypy. Status: valid subgenus of Hyperchalca Fairmaire, 1869 in Stenochiinae: Stenochiini.

Macrolagria Lewis, 422 [F]. Type Statirarufobrunnea Marseul, 1876, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Macropachylesthus Pic, 28 [M]. Type Macropachylesthusgigas Pic, 1923, by subsequent designation (Gebien 1941: 350). Status: valid genus in Stenochiinae: Cnodalonini.

Macroperas Carter, 393 [N]. Type Macroperasantennale Carter, 1914, by monotypy. Status: junior synonym of Daedrosis Bates, 1868 in Lagriinae: Adeliini. Synonymy: Matthews (1998: 756).

Macrophanes Iablokoff-Khnzorian, 165 [M]. Type Hedyphanescorax Iablokoff-Khnzorian, 1957 (= Helopsamandanus Reitter, 1902), by original designation. Status: junior synonym of Delonurops Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko (2002b: 689).

Macrophtalmus Montrouzier, 33 [M]. Type Macrophtalmuscoeruleus Montrouzier, 1855, by monotypy. Status: senior synonym of Macrophthalmata Strand, 1935 in Tenebrionidae: incertae sedis. Note: this genus has been treated as a junior homonym of Macrophtalmus Laporte, 1832 [Hemiptera] in the literature; however, the hemipteran name was originally proposed with two spellings, Macrophthalmus (Laporte, 6, 11, 87) and Macrophtalmus (Laporte, 78); the alternative original spelling Macrophtalmus of the hemipteran name, used by Laporte (1832: 78), was rejected by Neave (1940: 17) who acted as First Reviser; therefore Macrophtalmus Montrouzier, 1855 is the valid name for the tenebrionid genus since it is not a junior homonym.

Macrophthalmata Strand, 302 [F]. Type species [automatic]: Macrophtalmuscoeruleus Montrouzier, 1855, by monotypy. Status: junior synonym of Macrophtalmus Montrouzier, 1855 in Tenebrionidae: incertae sedis. Note: replacement name for Macrophtalmus Montrouzier, 1855.

Macropoda Solier, 512, 515 [F]. Type Pimeliavariolaris G.-A. Olivier, 1795, by subsequent designation (Hope 1841: 118). Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Macropodesmia Löbl & Merkl in Löbl et al., 2020: 1 [F]. Type Pimeliareticulata Klug, 1830, by original designation. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: name first proposed by Koch (1944b: 147) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 244) designated Pimeliareticulata Klug, 1830 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Macrostethus Wollaston, 504 [M]. Type Macrostethustuberculatus Wollaston, 1854, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Macrosynopticus Pic, 25 [M]. Type Macrosynopticuscostatus Pic, 1922, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Macrotrachyscelis Pic, 12 [F]. Type Macrotrachyscelisrufa Pic, 1925, by monotypy. Status: valid genus in Diaperinae: Trachyscelini.

Macrozophobas Pic, 6 [M]. Type Macrozophobasgracilicornis Pic, 1913, by monotypy. Status: valid subgenus of Zophobas Dejean, 1834 in Tenebrioninae: Tenebrionini.

Macruloma Pic, 20 [N]. Type Macrulomagigas Pic, 1921, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Madagassa Koch, 67 [F]. Type species [automatic]: Pycnaaphodina Fairmaire, 1894, by monotypy. Status: valid genus in Phrenapatinae: Penetini. Note: replacement name for Pycna Fairmaire, 1894.

Madobalus Fairmaire, 73 [M]. Type Madobalusrotundicollis Fairmaire, 1901, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Madreallecula Kanda, 587 [F]. Type Madrealleculamcclevei Kanda, 2013, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Maerodes C.O. Waterhouse, 72 [M]. Type Prophaneswestwoodi W.J. MacLeay, 1872 (= Prophanesaculeatus Westwood, 1849), by monotypy. Status: junior synonym of Prophanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1918: 717).

Magdanalia Novák, 496 [F]. Type Prionychusdenticulatus Muche, 1982, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Magela G.S. Medvedev & Lawrence, 578 [F]. Type Magelauptoni G.S. Medvedev & Lawrence, 1986, by original designation. Status: valid genus in Diaperinae: Hyociini: Brittonina.

Magrebmelia Mas-Peinado, Buckley, Ruiz & García-París, 540 [F]. Type Pimeliaxauenensis Escalera, 1923, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Mahena Gebien, 315 [F]. Type Mahenacuprea Gebien, 1922, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Makicula Novák, 275 [F]. Type Makiculaphoupaneica Novák, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Malacova Fairmaire, 479 [F]. Type Malacovabicolor Fairmaire, 1898, by present designation. Status: junior synonym of Damatris Laporte, 1840 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1942a: 316).

Malaiseum Borchmann, 13 [N]. Type Malaiseumsingulare Borchmann, 1941, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Malayaplamius Masumoto, 17 [M]. Type Malayaplamiussakaii Masumoto, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Malaymira Novák, 56 [F]. Type Malaymirajenisi Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Malayoscelis Schawaller, 198 [F]. Type Malayoscelisgebieni Schawaller, 2003, by original designation. Status: valid genus in Lagriinae: Pycnocerini.

Malaysphena Bečvář & Purchart, 38 [F]. Type Laosocryptobatesrotundipennis Kaszab 1960, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Mallogria Borchmann, 20, 165 [F]. Type Lagrialongipilis Fairmaire, 1875, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Mamorina Antoine, 98 [F]. Type Mamorinasulcaticeps Antoine, 1951, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Mantichorula Reitter, 695 [F]. Type Mantichorulasemenowi Reitter, 1889, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Maracia Gebien, 27, 34 [F]. Type Camariafemoralis Kirsch, 1866, by subsequent designation (Gebien 1942a: 319). Status: valid genus in Stenochiinae: Cnodalonini.

Margus Dejean, 200 [M]. Type fixed herein (ICZN 1999, Article 70.3) as Colydiumcastaneum Herbst, 1797, misidentified as Tenebrioferrugineus Fabricius, 1781 in the original designation by monotypy in Dejean (1834). Status: junior synonym of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini. Synonymy: Guérin-Méneville (1846: cxvii). Note: the type species “Tenebrioferrugineus Fabricius” was first established by monotypy (ICZN 1999, Article 68.3); as noted by C.O. Waterhouse (1896: 230) and Blair (1913: 223) the Tenebrioferrugineus Fabricius of authors, including Stephens (1829: 19, as “ferruginea, Oliv.”), was misidentified; Blair (1913: 223) noted that the species to which the authors referred is in fact Colydiumcastaneum Herbst, 1797; we follow currently accepted concepts (e.g., Bousquet et al. 2018: 224) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Tenebrioferrugineus Fabricius, 1781 is a valid species in the genus Tribolioides Blair, 1913 [Coleoptera: Cucujidae].

Mariepskopia Schawaller, 218 [F]. Type Mariepskopiaalbomaculata Schawaller, 2012, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Martianus Fairmaire, 540 [M]. Type Martianuscastaneus Fairmaire, 1893 (= Palembusocularis Casey, 1891), by original designation. Status: junior synonym of Ulomoides Blackburn, 1888 in Diaperinae: Diaperini: Diaperina. Synonymy: Halstead (1975: 241, with Palembus Casey, 1891, a junior synonym of Ulomoides Blackburn, 1888).

Massadraamelia Mas-Peinado, Buckley, Ruiz & García-París, 541 [F]. Type Pimeliagranulithorax Escalera, 1914, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Mateuina Español, 22, 30 [F]. Type Mateuinakaszabi Español, 1944, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Matthewsotys Bouchard & Bousquet, new genus [M]. Type Otysarmatus Champion, 1895, by present designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: Matthews and Bouchard (2008: 333) selected Otysarmatus Champion, 1895 as the type species of Otys Champion, 1895 and treated it as a valid genus; however, an earlier type species designation by R. Lucas (1920: 468) confirms Otys Champion, 1895 as a junior synonym of Scaletomerus Blackburn, 1891; we hereby establish Matthewsotys as a new genus and refer to Matthews and Bouchard (2008: 205) for the character states that characterise and differentiate this taxon.

Mauritianopidium Dajoz, 244 [N]. Type Mauritianopidiumoculatum Dajoz, 1977, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Mayidicistela Pic, 259 [F]. Type Mayidicistelapaulostriata Pic, 1954, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Mechanetes C.O. Waterhouse, 448 [M]. Type Mechanetescornutus C.O. Waterhouse, 1887, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mecocerus Solier, 241 [M]. Type Xystropusdejeanii Solier, 1835, by monotypy. Status: junior synonym of Prostenus Klug, 1829 in Alleculinae: Alleculini: Xystropodina. Synonymy: Lacordaire (1859b: 513). Note: junior homonym of Mecocerus Schönherr, 1833 [Coleoptera: Anthribidae].

Mecopisthopus Karsch, 46 [M]. Type Mecopisthopusrohlfsi Karsch, 1881, by original designation. Status: junior synonym of Leucolaephus P.H. Lucas, 1859 in Pimeliinae: Pimeliini. Synonymy: Semenov (1893: 261).

Mecysmus Horn, 349 [M]. Type Blapstinusangustus J.L. LeConte, 1851, by monotypy. Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Lumen et al. (2020: 342).

Mederis Motschulsky, 24 [M]. Type Upisangulata Erichson, 1842, by original designation. Status: junior synonym of Promethis Pascoe, 1869 in Stenochiinae: Cnodalonini. Synonymy: C.O. Waterhouse (1876: 288).

Medvedevia Chigray, 915 [F]. Type Medvedeviaglebi Chigray, 2019, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Medvedevoblaps Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Protoblapskashkarovi G.S. Medvedev, 1998, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina. Note: replacement name for Protoblaps G.S. Medvedev, 1998.

Megacantha Westwood, 121 [F]. Type Megacanthatenebrosa Westwood, 1843 (= Helopsdentatus Fabricius, 1801), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: genus also described as new by Westwood (1844: 228).

Megadasus Reitter, 146 [M]. Type Opatrumlefranci Fairmaire, 1863, by original designation. Status: junior synonym of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Gebien (1939: 443).

Megagenius Solier, 512, 513 [M]. Type Megageniusfrioli Solier, 1835, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Megalophrys G.R. Waterhouse, 321, 322 [F]. Type Megalophryspatagonica G.R. Waterhouse, 1845, by monotypy. Status: senior synonym of Peltolobus Lacordaire, 1859 in Pimeliinae: Trilobocarini. Note: although the older name Megalophrys Wagler, 1830 [Amphibia] was placed on the Official Index of Rejected and Invalid Generic Names in Zoology by the ICZN (1994b, Opinion 1763), it was not suppressed for the Principle of Homonymy and therefore remains available and the senior homonym.

Megaprosodes Reitter, 118 [M]. Type Prosodesstriata Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Megasattus Casey, 56, 62 [M]. Type Eusattuserosus Horn, 1870, by original designation. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: Triplehorn (1968: 379).

Megascythis Keleinikova, 622 [F]. Type Megascythispanfilovi Keleinikova, 1963, by monotypy. Status: junior synonym of Scythis Kraatz, 1865 in Pimeliinae: Tentyriini. Synonymy: Skopin (1979: 171).

Megasida Casey, 77, 202 [F]. Type Asidaobliterata Champion, 1892, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Megatenebrio Gridelli, 221 [M]. Type Tenebriogiganteus Fairmaire, 1897, by monotypy. Status: valid subgenus of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini.

Megatlasion Koch, 427 [N]. Type Micrositusatlantis Escalera, 1914, by original designation. Status: junior synonym of Atlasion Koch, 1948 in Blaptinae: Dendarini: Melambiina. Synonymy: Antoine (1957: 351).

Megeleates Casey, 623 [M]. Type Megeleatessequoiarum Casey, 1895, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Megelenophorus Gebien, 121 [M]. Type species [automatic]: Elenophorusamericanus Lacordaire, 1830, by monotypy. Status: valid genus in Pimeliinae: Elenophorini: Megelenophorina. Note: replacement name for Cacicus Dejean, 1834.

Megischia Solier, 245, 247 [F]. Type Cistelacurvipes Brullé, 1832, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Megischina Reitter, 118, 171 [F]. Type Cistelaarmillata Brullé, 1832, by subsequent designation (R. Lucas 1920: 399). Status: valid genus in Alleculinae: Cteniopodini.

Meglyphus Motschulsky, 38 [M]. Type Meglyphuslaenoides Motschulsky, 1872, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina. Note: we act as First Revisers and reject the alternative original spelling Megliphus, used by Motschulsky (1872: 41).

Meladeras Mulsant & Rey, 191, 219 [N]. Type Meladerasquadratulum Mulsant & Rey, 1854, by subsequent designation (Viñolas 1990: 61). Status: valid subgenus of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina.

Meladiesia Reitter, 309 [F]. Type Meladiesiamiritarsis Reitter, 1909, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Meladocrates Reitter, 96 [M]. Type Olocratesplaniusculus Mulsant & Rey, 1854, by subsequent designation (Viñolas 1990: 60). Status: valid subgenus of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina.

Melambasida Reitter, 10, 21 [F]. Type Alphasidainterjecta Reitter, 1917, by monotypy. Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Melambatlasus Koch, 433 [M]. Type Micrositushebes Antoine, 1933, by original designation. Status: valid subgenus of Melambius Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Melambiophylax Schuster, 48, 49 [M]. Type Phylaxsardous Baudi di Selve, 1876, by monotypy. Status: junior synonym of Allophylax Bedel, 1906 in Blaptinae: Dendarini: Melambiina. Synonymy: Leo (1994: 134).

Melambius Mulsant & Rey, 124 [M]. Type Opatrumbarbarum Erichson, 1841, by monotypy. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Melanastus Casey, 289, 353 [M]. Type Eurymetoponatrum J.L. LeConte, 1851, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Melancrus Reiche & Saulcy, 190 [N]. Type Melancruslaevigatum Reiche & Saulcy, 1857, by subsequent designation (Löbl et al. 2008a: 42). Status: junior synonym of Oxycara Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Lacordaire (1859a: 57). Note: authors have assigned a masculine gender for this name in the literature; however, the correct gender is neuter; Melancrus has two melan- and -crus, the first root is clearly Greek: melas, melaina, melan (an irregular adjective meaning dark or black) while the second is Latin: crus (the name for a leg, shank, or shin); the second word is a neuter noun and therefore the gender of Melancrus is neuter (ICZN 1999, Article 30.1.1).

Melaneleodes Blaisdell, 36 [M]. Type Blapscarbonaria Say, 1824, by subsequent designation (Triplehorn and Thomas 2012: 254). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Melanesthes Dejean, 191 [F]. Type Opatrumsibiricum Faldermann, 1833, by subsequent designation (Gebien 1939: 462). Status: valid genus and subgenus in Blaptinae: Opatrini: Opatrina.

Melanimon Motschulsky, 78 [N]. Type Microzoumcollare Motschulsky, 1839, by monotypy. Status: senior synonym of Platynosum Mulsant & Rey, 1859 in Blaptinae: Opatrini: Sclerina. Synonymy: Seidlitz (1894: 413, 454). Note: junior homonym of Melanimon Steven, 1828 [Coleoptera: Tenebrionidae: Tenebrioninae: Melanimonini].

Melanimon Steven, 98 [N]. Type Opatrumtibiale Fabricius, 1781, by monotypy. Status: valid genus in Tenebrioninae: Melanimonini.

Melanochrus Wollaston, 467 [M]. Type Melanochruslacordairii Wollaston, 1864, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Melanocoma Wollaston, 181 [F]. Type Melanocomavestita Wollaston, 1868, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Melanocratus Fairmaire, 21 [M]. Type Melanocratusvalidipes Fairmaire, 1895, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Melanolophus Fairmaire in Fairmaire et al., 1882: 69 [M]. Type Melanolophusseptemcostatus Fairmaire, 1882, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Melanophorus Lacordaire, 74 [M]. Type species [automatic]: Melaphorusreichii Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Melaphorus Guérin-Méneville, 1834 in Pimeliinae: Evaniosomini. Note: unjustified emendation of Melaphorus Guérin-Méneville, 1834, not in prevailing usage.

Melanopterus Mulsant & Rey, 13, 14 [M]. Type Melanopterusporcatus Mulsant & Rey, 1854, by subsequent designation (Gebien 1938a: 292). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Melanostola Solier, 123 [F]. Type Pimeliasimplex Solier, 1836, by monotypy. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Note: the name Melanostola was listed as synonym of Pimelia Fabricius, 1775 by Solier (1836; published by 16 May), being treated before 1961 as an available name and adopted as the name of a taxon (e.g., Dejean 1836 (published by 30 July), Hope 1841: 118); Melanostola was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Melansis Wollaston, 491 [F]. Type Phylaxcostatus Brullé, 1839, by subsequent designation (Dallas 1865: 401). Status: valid genus in Blaptinae: Dendarini: Melambiina.

Melaphorus Guérin-Méneville, 13 [M]. Type Melaphorusreichii Guérin-Méneville, 1834, by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Melaps Carter, 409 [M]. Type Melapscisteloides Carter, 1908, by monotypy. Status: junior synonym of Oocistela Borchmann, 1908 in Alleculinae: Alleculini: Alleculina. Synonymy: Carter (1915a: 78). Note: although this name has been used as valid, with Oocistela Borchmann, 1908 as a synonym, in recent literature (e.g., Matthews and Lawrence 2019: 645), data on the date of publication of both names reveals that Oocistela was published first and has priority over Melaps.

Melarachnica Kraatz, 80, 174 [F]. Type Melarachnicawestermanni Kraatz, 1865 (= Microderacoromandelensis Solier, 1835), by monotypy. Status: junior synonym of Nemapus Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Koch (1943b: 790).

Melasia Perroud & Mulsant, 160 [F]. Type Melasiagagatina Perroud & Mulsant, 1856 (= Tenebrioculinaris Linnaeus, 1758), by subsequent designation (G.S. Medvedev 1990: 227). Status: junior synonym of Uloma Dejean, 1821 in Tenebrioninae: Ulomini. Synonymy: Lacordaire (1859a: 332).

Melasma Wollaston, 484 [N]. Type Phylaxlineatus Brullé, 1838, by monotypy. Status: senior synonym of Melasmana Strand, 1935 in Blaptinae: Dendarini: Melambiina. Note: junior homonym of Melasma Adams, 1854 [Mollusca].

Melasmana Strand, 297 [F]. Type species [automatic]: Phylaxlineatus Brullé, 1838, by monotypy. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina. Note: replacement name for Melasma Wollaston, 1864.

Melasmocara Reitter, 95 [N]. Type Melanochruslacordairii Wollaston, 1864, by subsequent monotypy (Löbl et al. 2008a: 42). Status: junior synonym of Melanochrus Wollaston, 1864 in Pimeliinae: Tentyriini. Synonymy: Jakobson (1924: 243), Löbl et al. (2008a: 42). Note: originally proposed without included nominal species; Löbl et al. (2008a: 42), by including the species Melanochruslacordairii Wollaston, 1864 in association with this name, were the first authors to subsequently and expressly include nominal species in Melasmocara (ICZN 1999, Article 67.2.2).

Melaxumia Reitter, 280 [F]. Type Melaxumiaacutangula Reitter, 1895 (= Tentyriaangulosa Gebler, 1832), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Melobates Kaszab, 4, 23 [M]. Type Melobatesbiroi Kaszab, 1941, by original designation. Status: junior synonym of Rehumius Fairmaire, 1893 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983d: 84).

Melobrachys Kaszab, 273 [M]. Type Melobrachyssarawakensis Kaszab, 1960, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Menandris Haag-Rutenberg, 103 [F]. Type Menandrisaenea Haag-Rutenberg, 1878, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mencheres Champion, 5 [M]. Type Mencheresnicaraguensis Champion, 1884, by subsequent designation (R. Lucas 1920: 403). Status: valid genus in Pimeliinae: Edrotini.

Menearchus Carter, 229 [M]. Type Menearchusimpressosulcatus Carter, 1920 (= Helopsnigritus Fabricius, 1777), by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina. Note: as mentioned by Matthews and Bouchard (2008: 351) the type species was originally described from Australia in error, as the genus Menearchus Carter, 1920 is endemic to India and Sri Lanka.

Menechides Motschulsky, 26 [M]. Type Helopscalcaratus Fabricius, 1798, by original designation. Status: valid subgenus of Centronopus Solier, 1848 in Tenebrioninae: Centronopini.

Menederes Solier, 152, 203 [M]. Type Menederesrufilabris Solier, 1848, by original designation. Status: valid genus and subgenus in Blaptinae: Platynotini: Eurynotina.

Menederopsis Koch, 16 [F]. Type Menederopsisconstricta Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Menedrio Motschulsky, 27 [M]. Type Tenebrioobscurus Fabricius, 1792, by original designation. Status: junior synonym of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini. Synonymy: Heyden in Heyden et al. (1883: 134).

Menephilus Mulsant, 291 [M]. Type Tenebriocurvipes Fabricius, 1792 (= Tenebriocylindricus Herbst, 1784), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Meneristes Pascoe, 150 [M]. Type Meneristeslaticollis Pascoe, 1869 (= Tenebrioaustralis Boisduval, 1835), by original designation. Status: valid genus in Tenebrioninae: Heleini: Asphalina.

Menes Champion, 442 [M]. Type Menesmeridanus Champion, 1888, by subsequent designation (Bousquet et al. 2015: 134). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Menimoides Kaszab, 19 [M]. Type Menimoidestarandus Kaszab, 1946, by original designation. Status: junior synonym of Micropeneta Pic, 1921 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Kaszab (1978a: 172).

Menimopsis Champion, 16 [F]. Type Menimopsisexcaecus Champion, 1896, by monotypy. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Menimus Sharp, 73 [M]. Type Menimusbatesi Sharp, 1876, by subsequent designation (Gebien 1940: 429). Status: valid genus and subgenus in Diaperinae: Gnathidiini: Gnathidiina.

Meniscophorus Champion, 64 [M]. Type Meniscophorusamazonicus Champion, 1889, by subsequent designation (R. Lucas 1920: 404). Status: valid genus in Lagriinae: Lagriini: Statirina.

Menoeceus Champion, 443 [M]. Type Menoeceuscrassicornis Champion, 1888, by subsequent designation (Casey 1891: 122). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Menoncotus Koch, 34 [M]. Type Oncotuschaetotaxicus Koch, 1954, by original designation. Status: valid subgenus of Oncotus Blanchard, 1845 in Blaptinae: Platynotini: Eurynotina.

Mentariobius Koch, 425 [M]. Type Micrositusdistinguendus Mulsant & Rey, 1854, by original designation. Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Mentes Champion, 559 [M]. Type Mentesruficollis Champion, 1893, by subsequent designation (R. Lucas 1920: 404). Status: valid genus in Stenochiinae: Stenochiini.

Meracantha W. Kirby, 237 [F]. Type Meracanthacanadensis W. Kirby, 1837 (= Helopscontractus Palisot de Beauvois, 1811), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Meracanthoides Linell, 698 [M]. Type Meracanthoidescupreolineatus Linell, 1896, by monotypy. Status: valid subgenus of Paramarygmus Quedenfeldt, 1885 in Tenebrioninae: Amarygmini.

Merinus J.L. LeConte, 230 [M]. Type Tenebriolaevis G.-A. Olivier, 1795, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: the original combination of the name of the type species, Tenebriolaevis G.-A. Olivier, 1795, is a junior primary homonym of Tenebriolaevis Forskål, 1775.

Merklia Chen, 307 [F]. Type Merkliabimaculata Chen, 1997, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Meropersina Reitter, 117 [F]. Type Prosodescordicollis Allard, 1884, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Meropria Borchmann, 217, 228 [F]. Type Statiraglabrata Mäklin, 1863, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Merotemnus Horn, 364, 367 [M]. Type Merotemnuselongatus Horn, 1870 (= Ulomafiliforme Laporte, 1840), by monotypy. Status: junior synonym of Adelonia Laporte, 1840 in Lagriinae: Belopini. Synonymy: Spilman (1961: 49).

Meroxys Ardoin, 132 [M]. Type Meroxyshaafi Ardoin, 1963, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: name first proposed by Ardoin (1962b: 970) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Mesabates Champion, 3 [M]. Type Mesabateslatifrons Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Mesabatodes Casey, 517 [M]. Type Mesabatesinaequalis Champion, 1892, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Mesoblaps Bauer, 231 [F]. Type Blaps rugulipennis Fairmaire, 1891, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Mesohelops Reitter, 31 [M]. Type Helopscyanipes Allard, 1877, by subsequent designation (Nabozhenko and Löbl 2008: 251). Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko and Keskin (2017: 41). Note: the original combination of the name of the type species, Helopscyanipes Allard, 1877, is a junior primary homonym of Helopscyanipes Fabricius, 1801.

Mesoleptodes G.S. Medvedev & Iljina, 881 [M]. Type Leptodessemenowi Reitter, 1892, by original designation. Status: valid subgenus of Leptodes Dejean, 1834 in Pimeliinae: Leptodini.

Mesolobopoda Campbell, 34 [F]. Type Alleculasocia J.L. LeConte, 1854, by original designation. Status: valid subgenus of Lobopoda Solier, 1835 in Alleculinae: Alleculini: Alleculina.

Mesomorphus Miedel, 140 [M]. Type Opatrummurinum Baudi di Selve, 1876 (= Opatrinussetosus Mulsant & Rey, 1853), by subsequent designation (Gebien 1938a: 399). Status: valid genus in Blaptinae: Opatrini: Opatrina.

Mesopatrum Broun, 1355 [N]. Type Mesopatrumgranulosum Broun, 1893, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Mesopraocis Flores & Pizarro-Araya, 60 [M]. Type Praociscalderanus Kulzer, 1958, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 12, 31) without type species designation.

Mesoprosodes G.S. Medvedev, 827 [M]. Type Prosodesnuratensis Semenov, 1894, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Mesopterocoma Skopin, 147 [F]. Type Pterocomasemicarinata Bates, 1879, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Mesostena Eschscholtz, 5, 9 [F]. Type Mesostenapunctata Eschscholtz, 1831 (= Pimeliaangustata Fabricius, 1775), by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Mesostenopa Kraatz, 80, 179 [F]. Type Mesostenopapicea Kraatz, 1865, by subsequent designation (Gebien 1937a: 608). Status: valid subgenus of Mesostena Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Mesosternoplax Skopin, 110, 142 [F]. Type Trigonoscelislaeviuscula Kraatz, 1882, by original designation. Status: valid subgenus of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini.

Mesotretis Bates, 151 [F]. Type Mesotretisferruginea Bates, 1872, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Messalia Pascoe, 442 [F]. Type Messaliavarians Pascoe, 1883, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Messoricolum Koch, 384 [N]. Type Messoricolumscotti Koch, 1960, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Metablapylis Blaisdell, 391 [F]. Type Eleodesnigrina J.L. LeConte, 1858, by subsequent designation (Bousquet et al. 2018: 159). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Metabolocerus Bates, 259 [M]. Type Metaboloceruspilosus Bates, 1873, by subsequent designation (R. Lucas 1920: 407). Status: valid genus in Tenebrioninae: Ulomini.

Metaclisa Jacquelin du Val, 296 [F]. Type Platydemaparallela Fairmaire, 1855 (= Diaperisazurea Waltl, 1838), by original designation. Status: valid genus in Tenebrioninae: Metaclisini. Note: nomen protectum (see Bouchard et al. 2007: 393).

Metacorticeus Bremer & Lillig, 71 [M]. Type Corticeusornatus Bremer, 1993, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Metacossyphodes Andreae, 205, 215 [M]. Type Cossyphodeskundelunguensis Basilewsky, 1950, by original designation. Status: junior synonym of Cossyphodes Westwood, 1851 in Pimeliinae: Cossyphodini: Cossyphodina. Synonymy: Schawaller (2013c: 362, implied by inclusion of Cossyphodeskundelunguensis Basilewsky, 1950 in Cossyphodes Westwood, 1851 without use of a subgenus rank).

Metallonotus Gray in Griffith and Pidgeon, 790 [M]. Type Metallonotusdenticollis Gray, 1832, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Methistamena Gebien, 28, 151 [F]. Type Methistamenaclavipes Gebien, 1919, by original designation. Status: junior synonym of Camariomorpha Pic, 1915 in Stenochiinae: Cnodalonini. Synonymy: Kulzer (1954a: 71).

Metisopus Bates, 371 [M]. Type Metisopuspurpureipennis Bates, 1873, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Metistete Pascoe, 489 [F]. Type Tanychilusgibbicollis Newman, 1838, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Metonites Gistel, viii, 126 [M]. Type species [automatic]: Blapidaokeni Perty, 1830, by monotypy. Status: junior synonym of Blapida Perty, 1830 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Blapida Perty, 1830.

Metopoloba Casey, 379 [F]. Type Epitraguspruinosus Horn, 1870, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Metoponiopsis Casey, 290 [F]. Type Eurymetoponbicolor Horn, 1870, by monotypy. Status: valid subgenus of Metoponium Casey, 1907 in Pimeliinae: Edrotini.

Metoponium Casey, 288 [N]. Type Eurymetoponabnorme J.L. LeConte, 1851, by original designation. Status: valid genus and subgenus in Pimeliinae: Edrotini.

Metriolagria Merkl, 124, 138 [F]. Type Lagriaaffinis Boisduval, 1835, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Metriopus Solier, 512, 570 [M]. Type Metriopushoffmanseggii Solier, 1835, by monotypy. Status: valid genus and subgenus in Pimeliinae: Adesmiini.

Metulosonia Bates, 261 [F]. Type Metulosoniahorni Bates, 1873, by subsequent designation (Gebien 1940: 1061). Status: valid genus in Tenebrioninae: Triboliini.

Micipsa P.H. Lucas, xxxiv [M]. Type Micipsarufitarsis P.H. Lucas, 1855 (= Pimeliamulsanti Levrat, 1853), by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Micipsina Reitter, 94, 188 [F]. Type Micipsinarolphi Reitter, 1900, by monotypy. Status: junior synonym of Thalpobia Fairmaire, 1871 in Pimeliinae: Tentyriini. Synonymy: Escalera (1914: 279).

Micrantereus Solier, 151, 175 [M]. Type Acanthomerusanomalus Guérin-Méneville, 1834, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Micrarmalia Casey, 516 [F]. Type Emmenastusconstrictus Champion, 1892, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Micrasida Smith, 608 [F]. Type Micrasidaobrienorum Smith, 2013, by original designation. Status: valid genus in Pimeliinae: Asidini.

Micrectyche Bates, 362 [F]. Type Micrectycheintermedia Bates, 1873, by subsequent designation (R. Lucas 1920: 412). Status: valid genus in Diaperinae: Ectychini.

Micreuphlaeus Fairmaire, 223 [M]. Type Micreuphlaeusasperipellis Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Micrisomira Pic, 30 [F]. Type Micrisomiraruficollis Pic, 1930, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Microamarygmus Pic, 8 [M]. Type Microamarygmusmadurensis Pic, 1915, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Microanaedus Pic, 16 [M]. Type Microanaedusnotatus Pic, 1923, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Microatasthalus Ando, 153 [M]. Type Microatasthalushadrocerus Ando, 2010, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Microbasanus Pic, 1 [M]. Type Microbasanusjureceki Pic, 1921, by monotypy. Status: junior synonym of Scaphidema Redtenbacher, 1848 in Diaperinae: Scaphidemini. Synonymy: Löbl et al. (2008b: 318), see also Schawaller (2008: 384).

Microblattellus Ferrer, 78 [M]. Type Microblattelluslecongmani Ferrer, 2006, by original designation. Status: valid genus in Tenebrioninae: Falsocossyphini.

Microblemma Semenov, 213 [N]. Type Microblemmasimplex Semenov, 1889, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Platamodina.

Microbolitonaeus Grimm, 194 [M]. Type Microbolitonaeusarmatus Grimm, 2014, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Microbradymerus Schawaller, 144 [M]. Type Microbradymerusmerkli Schawaller, 1999, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Microcalcar Pic, 9 [N]. Type Belopusinstriatus Pic, 1925, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Microcalydonis Pic, 17 [F]. Type Microcalydonismetallica Pic, 1923, by monotypy. Status: valid subgenus of Osternus Fairmaire, 1895 in Stenochiinae: Cnodalonini.

Microcameria Ren, 108, 113 [F]. Type Microcameriapygmaea Ren, 1998, by original designation. Status: junior synonym of Foochounus Pic, 1921 in Stenochiinae: Cnodalonini. Synonymy: Ando (2008: 39).

Microcatomus Pic, 7 [M]. Type Microcatomuslongipilis Pic, 1925, by monotypy. Status: valid genus in Tenebrioninae: Helopini: incertae sedis. Note: Nabozhenko (2018: 183) mentioned that the position of this genus within the tribe Helopini is unclear.

Microcenoscelis Schawaller, 438 [F]. Type Microcenosceliscaeca Schawaller, 2015, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Microcilibe Carter, 147 [F]. Type Microcilibecastanea Carter, 1919, by monotypy. Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Kaszab (1978a: 173).

Microcistela Pic, 26 [F]. Type Microcistelarosinae Pic, 1904, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Microcistela Pic, 6 [F]. Type Microcistelaobscura Pic, 1919, by monotypy. Status: senior synonym of Microcistelopsis Pic, 1922 in Alleculinae: Alleculini: Alleculina. Note: junior homonym of Microcistela Pic, 1904 [Coleoptera: Tenebrionidae: Alleculinae: Alleculini: Gonoderina].

Microcistelopsis Pic, 20 [F]. Type species [automatic]: Microcistelaobscura Pic, 1919, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: replacement name for Microcistela Pic, 1919.

Microcrypticus Gebien, 7 [M]. Type Diaperisvariegata Klug, 1833, by original designation. Status: valid genus and subgenus in Diaperinae: Crypticini.

Microdendarus Escalera, 84 [M]. Type Dendarusschusteri Español, 1937, by original designation. Status: junior synonym of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina. Synonymy: Español (1961a: 44).

Microdera Eschscholtz, 5, 6 [F]. Type Tentyriadeserta Tauscher, 1812, by subsequent designation (Gebien 1937a: 619). Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Microderopsis Haag-Rutenberg, 86 [F]. Type Microderopsisbenguelensis Haag-Rutenberg, 1876, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Microdisema Pic, 2, 3 [F]. Type Disemagounellei Pic, 1912, by monotypy. Status: valid subgenus of Barsenis Pascoe, 1887 in Lagriinae: Lagriini: Statirina. Note: this genus was established without included nominal species; Disemagounellei Pic, 1912 was first and expressly included in Microdisema by Pic (1937c: 36).

Microdocnemis Nabozhenko & Keskin, 841 [F]. Type Microdocnemisxerophilica Nabozhenko & Keskin, 2010, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Microeucyrtus Pic, 47 [M]. Type Microeucyrtuscoomani Pic, 1926, by monotypy. Status: junior synonym of Eucyrtus Lacordaire, 1859 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1941: 1139).

Microgauromaia Pic, 23 [F]. Type Microgauromaiaminuta Pic, 1921, by monotypy. Status: junior synonym of Phaedis Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Ando (2007: 164).

Microgoniadera Pic, 9 [F]. Type Microgoniaderahirsuta Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Microgonocnemis Pic, 16 [F]. Type Microgonocnemiscarinata Pic, 1936, by monotypy. Status: valid subgenus of Paragonocnemis Kraatz, 1899 in Tenebrioninae: Amarygmini.

Microhedyphanes Nabozhenko & Lillig, 189 [M]. Type Hedyphaneschikatunovi Nabozhenko & Lillig, 2013, by original designation. Status: valid subgenus of Hedyphanes Fischer, 1820 in Tenebrioninae: Helopini: Helopina.

Microhemicera Pic, 30 [F]. Type Hemicerahumeralis Pic, 1921, by monotypy. Status: junior synonym of Simalura Gebien, 1914 in Stenochiinae: Cnodalonini. Synonymy: Löbl et al. (2008b: 347).

Microhionthis Blair, 120 [F]. Type Microhionthispatriciae Blair, 1923 (= Falsocatomuluseuphraticus Pic, 1914), by monotypy. Status: junior synonym of Falsocatomulus Pic, 1914 in Pimeliinae: Tentyriini. Synonymy: Koch (1941: 267).

Microlagria Seidlitz, 336, 339 [F]. Type Lagriapoupillieri Reiche, 1864, by monotypy. Status: junior synonym of Adynata Fåhraeus, 1870 in Lagriinae: Lagriini: Lagriina. Synonymy: Borchmann (1936: 23).

Microleichenum G.S. Medvedev, 648 [N]. Type Microleichenumchoresmense G.S. Medvedev, 1973, by original designation. Status: junior synonym of Apsheronellus Bogatchev, 1967 in Blaptinae: Pedinini: Leichenina. Synonymy: G.S. Medvedev and Iwan (2006: 614).

Microlyprops Kaszab, 108 [M]. Type Microlypropsceylonicus Kaszab, 1939, by original designation. Status: valid genus in Lagriinae: Goniaderini.

Micromenandris Kaszab, 511, 513 [F]. Type Micromenandrismirabilis Kaszab, 1955, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Micromes Casey, 432, 441 [M]. Type Stibiaovipennis Horn, 1874, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Micromophlus Znojko in Ogloblin and Znojko, 124 [M]. Type Omophlussubtilis Solsky, 1881, by monotypy. Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini.

Micronilio Pic, 198 [M]. Type Niliopunctatus Pic, 1918 (= Niliogounellei Ihering, 1914), by monotypy. Status: valid subgenus of Nilio Latreille, 1802 in Nilioninae.

Microomopheres Freude, 214 [M]. Type Omopheresbrendelli Freude, 1993, by monotypy. Status: valid subgenus of Omopheres Casey, 1907 in Pimeliinae: Epitragini.

Micropedinus Lewis, 379 [M]. Type Micropedinusalgae Lewis, 1894 (= Heterophagapullulus Boheman, 1858), by subsequent designation (R. Lucas 1920: 415). Status: valid genus in Lagriinae: Lupropini.

Micropeltoides Pic, 16 [M]. Type Micropeltoidescrypticoides Pic, 1916, by monotypy. Status: valid subgenus of Peltoides Laporte, 1833 in Tenebrioninae: Alphitobiini.

Micropeneta Pic, 19 [F]. Type Micropenetatestacea Pic, 1921, by subsequent designation (Riley 1923: 128). Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Microphenus Gebien, 324, 338 [M]. Type Microphenuscordicollis Gebien, 1921 (= Espitesobscurus Blair, 1915), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Microphligra Koch, 47 [F]. Type Phligraminuta Péringuey, 1904, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Microphyes W.J. MacLeay, 286 [M]. Type Microphyesrufipes W.J. MacLeay, 1872 (= Opatrumlaevigatum Fabricius, 1781), by monotypy. Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Blair (1914: 486).

Microphylacinus Iwan, Kamiński & Aalbu, 2 [M]. Type Microphylacinusverendus Iwan, Kamiński & Aalbu, 2011, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Microplatyscelis Kaszab, 142, 144 [F]. Type Faustiaseriepunctata Reitter, 1890, by original designation. Status: valid genus in Blaptinae: Platyscelidini.

Microprostenus Pic, 14 [M]. Type Microprostenuslongicornis Pic, 1921, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Micropseudochillus Fouquè, 230, 240 [M]. Type Pseudochilluspalawanus Fouquè, 2015, by original designation. Status: valid subgenus of Pseudochillus Fouquè, 2015 in Pimeliinae: Stenosini: Dichillina.

Microschatia Solier, 406, 474 [F]. Type Microschatiapunctata Solier, 1836, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Microselinus Koch, 214 [M]. Type Microselinusmuelleri Koch, 1956, by original designation. Status: junior synonym of Glyptopteryx Gebien, 1910 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 91).

Microsis Koch, 76, 82 [F]. Type Microsisvilhenai Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Micrositus Mulsant & Rey, 131, 148 [M]. Type Micrositusorbicularis Mulsant & Rey, 1854, by subsequent designation (Gebien 1938a: 415). Status: valid genus in Blaptinae: Dendarini: Dendarina.

Microsphaerotus Pic, 10 [M]. Type Microsphaerotusruficornis Pic, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Microstenogena Pic, 30 [F]. Type Microstenogenaruficornis Pic, 1924, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Microsthes Novák, 322 [M]. Type Microsthesbarborae Novák, 2011, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Microstizopus Koch, 36 [M]. Type Microstizopusciliatus Koch, 1963, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Microstrongylium Pic, 13 [N]. Type Microstrongyliumcyanicolle Pic, 1917, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Kaszab (1977b: 30).

Microtelopsis Koch, 742 [F]. Type Tetranillussimplicifrons Gridelli, 1934, by monotypy. Status: valid genus and subgenus in Pimeliinae: Stenosini: Stenosina. Note: we act as First Revisers and select Microtelopsis Koch, 1940 as the valid name for this genus instead of Extetranosis Koch, 1940 and Hypermicrotelopsis Koch, 1940.

Microtelus Solier, 7, 9 [M]. Type Microtelusasiaticus Solier, 1838, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Microthelecta Pic, 65 [F]. Type Cylindrothorusbraunsi Pic, 1952, by monotypy. Status: valid subgenus of Cylindrothorus Solier, 1843 in Alleculinae: Alleculini: Mycetocharina.

Microtocerus Pic, 11 [M]. Type Microtocerusgrandicornis Pic, 1918, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Microzoon Hope, 112 [N]. Type species [automatic]: Opatrumtibiale Fabricius, 1781, by monotypy. Status: junior synonym of Melanimon Steven, 1828 in Tenebrioninae: Melanimonini. Note: unjustified emendation of Microzoum Dejean, 1834, not in prevailing usage.

Microzophobas Pic, 7 [M]. Type Microzophobasluteomaculatus Pic, 1944, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Microzoum Dejean, 193 [N]. Type Opatrumtibiale Fabricius, 1781, by monotypy. Status: junior synonym of Melanimon Steven, 1828 in Tenebrioninae: Melanimonini. Synonymy: Seidlitz (1894: 452).

Mictopsis Fairmaire, 538 [F]. Type Mictopsislaticollis Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Miglica Reitter, 171 [F]. Type Pedinuslaticollis Gebler, 1830, by subsequent designation (Iwan and Löbl 2008: 267). Status: junior synonym of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Reichardt (1936: 175).

Millotella Koch, 116, 130 [F]. Type Millotellamicrocornis Koch, 1962, by original designation. Status: senior synonym of Kochotella Bouchard & Bousquet, nom. nov. in Pimeliinae: Asidini. Note: junior homonym of Millotella Poisson, 1949 [Hemiptera].

Millstreamia G.S. Medvedev & Lawrence, 579 [F]. Type Csiroparodoxa G.S. Medvedev & Lawrence, 1984, by original designation. Status: valid subgenus of Csiro G.S. Medvedev & Lawrence, 1984 in Diaperinae: Hyociini: Hyociina.

Miltoprepes Gerstaecker, 65 [M]. Type Miltoprepeslaetus Gerstaecker, 1871, by monotypy. Status: valid genus in Tenebrioninae: Praeugenini.

Mimelasida Reitter, 10, 21 [F]. Type Asidapuncticollis Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Mimoborchmania Pic, 31 [F]. Type Nemostiracoloripes Pic, 1922, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Mimocellus Wasmann, 11 [M]. Type Mimocellustrechoides Wasmann, 1904, by subsequent designation (R. Lucas 1920: 419). Status: valid genus in Lagriinae: Lupropini.

Mimocistela Borchmann, 124 [F]. Type Mimocistelazumpti Borchmann, 1938, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Mimocossyphus Pic, 5 [M]. Type Mimocossyphusminor Pic, 1923, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: incertae sedis. Note: Schawaller (2013c: 362) mentioned that the position of this taxon within Cossyphodini is doubtful.

Mimogoueum Pic, 10 [N]. Type Mimogoueumhutteli Pic, 1952, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Ardoin (1962a: 64).

†Mimohelops Haupt, 114, 130 [M]. Type Mimohelopsvenosus Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Mimolaena Ardoin, 36 [F]. Type Mimolaenapauliani Ardoin, 1961, by monotypy. Status: valid genus in Lagriinae: Laenini.

Mimolagria Pic, 44 [F]. Type Mimolagriabruchi Pic, 1927, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Mimopeus Pascoe, 477 [M]. Type Mimopeusamaroides Pascoe, 1866 (= Celibeelongata Brême, 1842), by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina. Note: as mentioned by Matthews and Bouchard (2008: 350) the type species was originally described from Australia in error, the genus Mimopeus Pascoe, 1866 is endemic to New Zealand; Matthews and Lawrence (2019: 629) mentioned that this genus seems indistinguishable from the Australian genus Celibe Boisduval, 1835.

Mimopraogena Pic, 10 [F]. Type Alleculametallicipennis Pic, 1942, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Mimosynopticus Pic, 25 [M]. Type Mimosynopticusparvulus Pic, 1922, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Mimothydemus Pic, 20 [M]. Type Mimothydemusangustatus Pic, 1923 (= Lophocnemiscyaneus Kraatz, 1880), by monotypy. Status: junior synonym of Lophocnemis Mäklin, 1867 in Stenochiinae: Stenochiini. Synonymy: Kulzer (1966: 352).

Mimoxenotermes Pic, 106 [M]. Type Mimoxenotermesduporti Pic, 1931, by monotypy. Status: valid genus in Tenebrioninae: Rhysopaussini.

Mimuroplatopsis Borchmann, 485 [F]. Type Mimuroplatopsisatricolor Pic, 1931, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: this genus was proposed earlier by Pic (1931b: 32) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Minasius Pic, 18 [M]. Type Minasiusopacipennis Pic, 1932, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Minorus Mulsant & Rey, 41 [M]. Type Eurynotusrugicollis Mulsant & Rey, 1854, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

†Miostenosis Wickham,1913: 297 [F]. Type Miostenosislacordairei Wickham, 1913, by original designation. Status: valid genus in Pimeliinae: Stenosini: incertae sedis. Note: described from Upper Eocene deposits (USA).

Miotodera Fairmaire, 190 [F]. Type Miotoderafuneraria Fairmaire, 1901, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Mireanopidium Dajoz, 240 [N]. Type Mireanopidiumcamerunense Dajoz, 1977, by original designation. Status: valid genus in Diaperinae: Gnathidiini: incertae sedis. Note: placed in “Gnathidiini incertae sedis” by Schawaller and Purchart (2012: 312).

Miripronotum Louw, 117, 118 [N]. Type Miripronotumprominoculatum Louw, 1979, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Misolampidius Solsky, 292 [M]. Type Misolampidiustentyrioides Solsky, 1876, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Misolampomorphus Kaszab, 2, 6 [M]. Type Misolampomorphuskochi Kaszab, 1941 (= Leprocaulusreitteri Pic, 1934), by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Misolampus Latreille, 160 [M]. Type Misolampushoffmannseggii Latreille, 1806 (= Pimeliagibbula Herbst, 1799), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mithippia Pascoe, 288, 293 [F]. Type Mithippiaaurita Pascoe, 1869, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Mitotagenia Reitter, 138, 153 [F]. Type Stenosisarabs Baudi di Selve, 1881, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Mitragardhus Koch, 379 [M]. Type Tragardhusnodosus Koch, 1956, by monotypy. Status: valid subgenus of Tragardhus Koch, 1956 in Blaptinae: Dendarini: Melambiina. Note: combined description of a new genus-group taxon and a single new species (ICZN 1999, Article 13.4).

Mitragenius Solier, 307, 328 [M]. Type Mitrageniusdejeanii Solier, 1836, by monotypy. Status: valid genus in Pimeliinae: Nycteliini. Note: the alternative original spelling Mitrogenius, used by Solier (1836: 330), was rejected by Solier (1838a: 487) who acted as First Reviser (ICZN 1999, Article 24.2.4).

Mitrephorus Carter, 83 [M]. Type Mitrephorusconvexicollis Carter, 1913, by monotypy. Status: senior synonym of Mitrothorax Carter, 1914 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1914b: 78, with Ctimene Bates, 1873, a senior synonym of Mitrothorax Carter, 1914). Note: junior homonym of Mitrephorus Schönherr, 1837 [Coleoptera: Curculionidae].

Mitrothorax Carter, 78 [M]. Type species [automatic]: Mitrephorusconvexicollis Carter, 1913, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: replacement name for Mitrephorus Carter, 1913. Note: the First Reviser (Mitrothorax Carter, 1914 versus Timeneca Carter, 1914) is Matthews (1992: 456)

Mitua Hope, 56 [F]. Type Mituabidwelli Hope, 1848 (= Opatrumtuberculicostatum White, 1846), by original designation. Status: senior synonym of Pseudopatrum Sharp, 1886 in Lagriinae: Adeliini. Synonymy: Blair (1919a: 531). Note: Mitua Hope, 1848 has been used as valid in recent literature despite the fact that it is a junior homonym of Mitua Strickland, 1841, an unjustified emendation for Mitu Lesson, 1831 [Aves]; Strickland's name has been used as valid since 1899 (e.g., Heinroth 1931: 278) and therefore reversal of precedence cannot be used to conserve the younger name as valid in Tenebrionidae.

Mitys Champion, 97 [M]. Type Mitysinflatus Champion, 1885, by subsequent designation (Gebien 1943: 402). Status: valid genus in Stenochiinae: Cnodalonini.

Modicodisema Pic, 3 [F]. Type Disemasubopaca Pic, 1912, by present designation. Status: valid subgenus of Barsenis Pascoe, 1887 in Lagriinae: Lagriini: Statirina. Note: originally proposed without included nominal species; based on personal communications with Maurice Pic, Borchmann (1936: 510) was the first author to subsequently and expressly include nominal species in Modicodisema (ICZN 1999, Article 67.2.2), by including the species Disemaserraticornis Mäklin, 1875 and Disemasubopaca Pic, 1912, in association with this name.

Moeon Champion, 251 [N]. Type Moeonisthmicum Champion, 1886, by subsequent designation (Gebien 1942a: 330). Status: valid genus in Stenochiinae: Cnodalonini. Note: as pointed out by M.A. Alonso-Zarazaga (in Bousquet et al. 2018: 307) the name Moeon is the Latinization of the Greek noun moion (privy parts) and is neuter.

Moerodes Rye, 62 [M]. Type species [automatic]: Prophaneswestwoodi W.J. MacLeay, 1872 (= Prophanesaculeatus Westwood, 1849), by monotypy. Status: junior synonym of Prophanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Note: unjustified emendation of Maerodes C.O. Waterhouse, 1877, not in prevailing usage.

Mogadoria Escalera, 467 [F]. Type Tentyriasubelegans Fairmaire, 1871, by monotypy. Status: junior synonym of Eulipus Wollaston, 1864 in Pimeliinae: Tentyriini. Synonymy: Gebien (1937a: 634).

Mokalagria Pic, 163 [F]. Type Lopholagriaangustata Pic, 1953, by original designation. Status: valid subgenus of Lopholagria Borchmann, 1916 in Lagriinae: Lagriini: Lagriina.

Molion Champion, 142 [M]. Type Penetataurus Lacordaire, 1859, by subsequent designation (Gebien 1940: 758). Status: valid genus in Phrenapatinae: Penetini.

Moluris Latreille, 169 [F]. Type Pimeliagibba Fabricius, 1787 (= Tenebriogibbus Pallas, 1781), by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina. Note: the original combination of the accepted name of the type species, Tenebriogibbus Pallas, 1781, is a junior primary homonym of both Tenebriogibbus Linnaeus, 1767 and Tenebriogibbus DeGeer, 1778.

Monatrum Reichardt, 81, 208 [N]. Type Opatrumcarinatum Gebler, 1830, by original designation. Status: junior synonym of Scleropatrum Reitter, 1887 in Blaptinae: Opatrini: Opatrina. Synonymy: Löbl and Merkl (2003: 250).

Mongolesthes Reitter, 174 [F]. Type Melanesthesheydeni Csiki, 1901, by monotypy. Status: valid subgenus of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina.

Mongolopterocoma Skopin, 146 [F]. Type Pterocomareitteri Frivaldszky, 1890, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Monodius Koch, 180 [M]. Type Selinusconvexipennis Gebien, 1904, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Monoloba Solier, 235 [F]. Type Lobopodadircaeoides Solier, 1835, by monotypy. Status: valid subgenus of Lobopoda Solier, 1835 in Alleculinae: Alleculini: Alleculina.

Montagona G.S. Medvedev, 176 [F]. Type Tagonoidesasperula Fairmaire, 1896, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Montaguea Kaszab, 227 [F]. Type Montagueacaledonica Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Adeliini.

Montanocatomus Nabozhenko, 842 [M]. Type Catomusgrandis G.S. Medvedev, 1978, by original designation. Status: valid subgenus of Catomus Allard, 1876 in Tenebrioninae: Helopini: Helopina.

Montanoodescelis Egorov, 592 [F]. Type Platyscelissahlbergi Reitter, 1900, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Monteithium Matthews, 704, 803 [N]. Type Monteithiumascetum Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Montiprosodes G.S. Medvedev, 83 [M]. Type Prosodesalaiensis Kraatz, 1885, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Mophis Champion, 168 [M]. Type Mophismarginicollis Champion, 1886, by subsequent designation (Gebien 1940: 1061). Status: valid genus in Diaperinae: Diaperini: Adelinina.

Mophon Champion, 247 [M]. Type Mophontinctipennis Champion, 1886, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Moragacinella Español, 162 [F]. Type species [automatic]: Moralesialongepilosa Kaszab, 1944, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: replacement name for Moralesia Kaszab, 1944.

Moralesia Kaszab in Español, 18, 29 [F]. Type Moralesialongepilosa Kaszab, 1944, by original designation. Status: senior synonym of Moragacinella Español, 1954 in Blaptinae: Opatrini: Ammobiina. Note: junior homonym of Moralesia Fowler, 1943 [Pisces].

Morica Dejean, 182 [F]. Type Akisplanata Fabricius, 1801, by subsequent designation (Hope 1841: 122). Status: valid genus in Pimeliinae: Akidini.

Morocaulus Fairmaire, 215 [M]. Type Morocaulusremipes Fairmaire, 1899 (= Bratynaapicalis Westwood, 1875), by monotypy. Status: junior synonym of Bratyna Westwood, 1875 in Alleculinae: incertae sedis. Synonymy: Borchamann (1938: 120, through synonymy of the type species with the type species of Bratyna Westwood, 1875).

Moromelas Fairmaire, 481 [M]. Type Moromelasfoveipennis Fairmaire, 1898, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Morphostenophanes Pic, 7 [M]. Type Morphostenophanesaenescens Pic, 1925, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mrazius Pic, 86 [M]. Type Mraziusnodulosus Pic, 1925, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mucheimira Novák, 188 [F]. Type Isomirastoetzneri Muche, 1981, by original designation. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Mutiloxicum Nabozhenko & Ivanov, 546 [N]. Type Toxicumelvirae Nabozhenko & Ivanov, 2018, by original designation. Status: valid subgenus of Toxicum Latreille, 1802 in Tenebrioninae: Toxicini: Toxicina.

Myatis Bates, 480 [F]. Type Myatishumeralis Bates, 1879, by subsequent designation (R. Lucas 1920: 425). Status: valid genus in Blaptinae: Platyscelidini.

Mycetochara Guérin-Méneville, 346 [F]. Type species [automatic]: Cistelaflavipes Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 118). Status: valid genus and subgenus in Alleculinae: Alleculini: Mycetocharina. Note: replacement name for Mycetophila Gyllenhal, 1810; Mycetochara was also proposed later in the same year by Berthold (1827: 371), see Bouchard and Bousquet (2020a: 101); nomenclatural stability in this genus is threatened by the discovery that 1) the accepted type species, Cistelascapularis Illiger, 1805 (= Cistelahumeralis Fabricius, 1787) by subsequent designation by Westwood (1838: 32), is currently placed in the valid subgenusErnocharis C.G. Thomson, 1859 (e.g., Novák 2020i: 436) and 2) there are two other species currently placed in subgenusErnocharis C.G. Thomson, 1859 that were selected as type species of Mycetochara before the currently accepted designation in Westwood (1838: 32): Cistelahumeralis Fabricius, 1787 was chosen as the type species by Guérin-Méneville (1827: 346) when he proposed the replacement name Mycetochara, and Cistelalinearis Illiger, 1794 (= Cistelamaura Fabricius, 1792) was selected as the type species for Mycetocharis Gyllenhal, 1827, another replacement name for Mycetophila Gyllenhal, 1810, by Stephens (1832a: 375); of the nominal species originally included in Mycetophila Gyllenhal, 1810 that are currently placed in the subgenusMycetochara Guérin-Méneville, 1827, C.G. Thomson (1859: 118) was the first to select Cistelaflavipes Fabricius, 1792 as the type species; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to set aside previous type species designations for Mycetochara Guérin-Méneville, 1827 and select the type species designation proposed by C.G. Thomson (1859: 118).

Mycetochares Latreille, 42 [M]. Type species [automatic]: Cistelaflavipes Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 118). Status: junior synonym of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: replacement name for Mycetophila Gyllenhal, 1810.

Mycetocharina Seidlitz, 136 [F]. Type Alleculaorientalis Faust, 1877, by monotypy. Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Mycetocharis Gyllenhal, 510 [F]. Type species [automatic]: Cistelaflavipes Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 118). Status: junior synonym of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: replacement name for Mycetophila Gyllenhal, 1810.

†Mycetocharoides Schaufuss, 269 [M]. Type Mycetocharoidesbaumeisteri Schaufuss, 1889, by monotypy. Status: valid genus in Alleculinae: Alleculini: Mycetocharina. Note: described from Eocene Baltic amber.

Mycetocula Novák, 78 [F]. Type Mycetocharinasubcruciata Pic, 1922, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Mycetophila Gyllenhal, 541 [F]. Type Cistelaflavipes Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 118). Status: senior synonym of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: junior homonym of Mycetophila Meigen, 1803 [Diptera].

Mychestes Pascoe, 96 [M]. Type Mychesteslignarius Pascoe, 1870, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Mycotrogus Horn, 364, 367 [M]. Type Mycotroguspiceus Horn, 1870, by subsequent designation (R. Lucas 1920: 427). Status: valid genus in Tenebrioninae: Triboliini.

Myladanesthes Skopin, 202 [F]. Type Myladinafortidens Reitter, 1915, by original designation. Status: valid subgenus of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina. Note: the alternative original spelling Myladansthes, used by Skopin (1961b: 202), was rejected by Skopin (1967: 205) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Myladina Reitter, 706 [F]. Type Myladinaunguiculina Reitter, 1889, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Myladion Reitter, 386 [N]. Type Myladionacuticolle Reitter, 1887, by monotypy. Status: valid subgenus of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Note: the alternative original spelling Miladion, used by Reitter (1887a: 385), was rejected by Reitter (1896b: 166) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Mylaris Pallas, 37 [F]. Type Tenebriogigas Linnaeus, 1763, by subsequent designation (Guérin-Méneville 1844: 120). Status: valid genus in Stenochiinae: Cnodalonini. Note: see Bousquet et al. (2018: 307, footnote 82) regarding a possible nomenclatural problem with this genus name.

Myonophloeus Bremer & Lillig, 68 [M]. Type Corticeustuberculatus Triplehorn, 1979, by original designation. Status: valid genus in Diaperinae: Hypophlaeini.

Myrmechixenus Chevrolat, 267 [M]. Type Myrmechixenussubterraneus Chevrolat, 1835, by monotypy. Status: valid genus in Diaperinae: Myrmechixenini.

Myrmechoxenus Gaubil, 288 [M]. Type species [automatic]: Myrmechixenussubterraneus Chevrolat, 1835, by monotypy. Status: junior synonym of Myrmechixenus Chevrolat, 1835 in Diaperinae: Myrmechixenini. Note: unjustified emendation of Myrmechixenus Chevrolat, 1835, not in prevailing usage; while both Myrmechixenus (p. 71) and Myrmechoxenus (p. 266) were used by Gaubil (1849), the fact that Myrmechixenus was written in italics in the Index (p. 282) indicates that the original spelling was treated as a synonym.

Myrmecocatops Wasmann, 268 [M]. Type Myrmecocatopslatus Wasmann, 1897, by monotypy. Status: valid genus in Diaperinae: Crypticini.

Myrmecodema Gebien, 402 [F]. Type species [automatic]: Myrmecosomanycterinoides Germain, 1855, by monotypy. Status: valid genus in Tenebrioninae: Trachelostenini. Note: replacement name for Myrmecosoma Germain, 1855. Note: transferred from Stenochiinae: Cnodalonini by Matthews and Lawrence (2015: 293).

Myrmecodichillus Kaszab, 6, 7, 14 [M]. Type Dichillusreichenspergeri Kaszab, 1960, by original designation. Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Myrmecopeltoides Kaszab, 318 [M]. Type Myrmecopeltoidescamponoti Kaszab, 1973, by original designation. Status: valid genus in Lagriinae: Goniaderini.

Myrmecophosis Koch, 72 [F]. Type Zophosispedinoides Gebien, 1920, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Myrmecosoma Germain, 403 [N]. Type Myrmecosomanycterinoides Germain, 1855, by monotypy. Status: senior synonym of Myrmecodema Gebien, 1943 in Tenebrioninae: Trachelostenini. Note: junior homonym of Myrmecosoma Mannerheim, 1846 [Coleoptera: Anthicidae].

Myrmecoxenus Märkel, 253 [M]. Type species [automatic]: Myrmechixenussubterraneus Chevrolat, 1835, by monotypy. Status: junior synonym of Myrmechixenus Chevrolat, 1835 in Diaperinae: Myrmechixenini. Note: unjustified emendation of Myrmechixenus Chevrolat, 1835, not in prevailing usage.

Nalassus Mulsant, 323 [M]. Type Helopsdryadophilus Mulsant, 1854, by subsequent designation (Nabozhenko 2001a: 630). Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina.

Nalepa Reitter, 366 [F]. Type Blapscylindracea Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Namaphosis Penrith, 127, 143 [F]. Type Zophosissolivaga Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Namaquaeon Koch, 305, 341 [N]. Type Phaeotribonaustralis Péringuey, 1908, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Namazopus Koch, 34 [M]. Type Namazopusarachnipes Koch, 1963, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Namibismus Koch, 107 [M]. Type Namibismuscastaneus Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Namibomodes Koch, 223 [M]. Type Psammodesserrimargo Gebien, 1938, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Nannalcyon Koch, 68 [F]. Type species [automatic]: Nannoceruscylindrus Fairmaire, 1887, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Nannocerus Fairmaire, 1887.

Nannocerus Fairmaire, 292 [M]. Type Nannoceruscylindrus Fairmaire, 1887, by monotypy. Status: senior synonym of Nannalcyon Koch, 1950 in Stenochiinae: Cnodalonini. Note: junior homonym of Nannocerus Mayr, 1885 [Hymenoptera].

Nannohyocis G.S. Medvedev & Lawrence, 569 [M]. Type Hyocisinquilinus Carter, 1921, by original designation. Status: valid subgenus of Hyocis Pascoe, 1866 in Diaperinae: Hyociini: Hyociina.

Nanoblaps Semenov-Tjan-Shansky & Bogatchev, 565 [F]. Type Blaps jakovlevi Semenov-Tjan-Shansky & Bogatchev, 1936, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Löbl et al. (2008c: 219).

Nanocaecus Schawaller & Purchart, 305, 310 [M]. Type Nanocaecushlavaci Schawaller & Purchart, 2012, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Nanocalcar Skopin, 67, 102 [N]. Type Nanocalcarnanum Skopin, 1974, by monotypy. Status: valid subgenus of Centorus Mulsant, 1854 in Lagriinae: Belopini.

Nanohemicera Pic, 25 [F]. Type Hemicerarufomaculata Pic, 1923, by monotypy. Status: valid subgenus of Hemicera Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini.

Nanotagalus Gebien, 118, 120 [M]. Type Afrotagalususambaricus Gebien, 1942, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Narses Champion, 423 [M]. Type Narsessubalatus Champion, 1888, by monotypy. Status: junior synonym of Charisius Champion, 1888 in Alleculinae: Alleculini: Alleculina. Synonymy: Campbell (2014: 271).

Narsodes Campbell, 32 [M]. Type Narsodesbrachypterus Campbell, 1976, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Nataloplonyx Ardoin, 716 [M]. Type Hoplonyxmicans Schaufuss, 1870, by original designation. Status: valid subgenus of Hoplonyx J. Thomson, 1858 in Tenebrioninae: Amarygmini.

Natalostira Pic, 143 [F]. Type Natalostirabrevithorax Pic, 1913, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Nautes Pascoe, 475 [M]. Type Nautesfervidus Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Neacisba Peyerimhoff, 53 [F]. Type species [automatic]: Pachychiladissecta Kraatz, 1865, by monotypy. Status: junior synonym of Neocisba Reitter, 1900 in Pimeliinae: Tentyriini. Note: unjustified emendation of Neocisba Reitter, 1900, not in prevailing usage.

Neandrosus Pic, 12 [M]. Type Neandrosussingularipes Pic, 1921, by monotypy. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1987: 44). Note: we act as First Revisers and reject the alternative original spelling Neoandrosus, used by Pic (1921b: 12).

Neanopidium Dajoz, 93 [N]. Type Neanopidiummexicanum Dajoz, 1975, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Neatus J.L. LeConte, 233 [M]. Type Helopstenebrioides Palisot de Beauvois, 1812, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Necrobioides Fairmaire, 234 [M]. Type Necrobioidescoeruleatus Fairmaire, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Neglectophloeus Bremer & Lillig, 71 [M]. Type Hypophlaeusluteomaculatus Pic, 1914, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Nelites J.L. LeConte, 232 [M]. Type Nelitesaeneolus J.L. LeConte, 1850, by monotypy. Status: junior synonym of Scaphidema Redtenbacher, 1848 in Diaperinae: Scaphidemini. Synonymy: J.L. LeConte (1862: 237).

Nemanes Fairmaire, 195 [M]. Type Nemanesexpansicollis Fairmaire, 1888, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Nemapus Solier, 313 [M]. Type Microderacoromandelensis Solier, 1835, by original designation. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: the name Nemapus was listed as synonym of Microdera Eschscholtz, 1831 by Solier (1835b: 313); it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Koch 1943b: 860); therefore, Nemapus was made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Nemoplonyx Ardoin, 715, 734 [M]. Type Hoplonyxgallanus Gridelli, 1939, by original designation. Status: valid subgenus of Hoplonyx J. Thomson, 1858 in Tenebrioninae: Amarygmini.

Nemostira Fairmaire, 815 [F]. Type Nemostiracoquerelii Fairmaire, 1869, by subsequent designation (R. Lucas 1920: 434). Status: junior synonym of Sora Walker, 1859 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1929a: 5).

Nemostiromorpha Pic, 2,3 [F]. Type Disemalongicornis Mäklin, 1875, by subsequent monotypy (Borchmann 1936: 510). Status: valid subgenus of Barsenis Pascoe, 1887 in Lagriinae: Lagriini: Statirina. Note: originally proposed without included nominal species; based on personal communications with Maurice Pic, Borchmann (1936: 510) was the first author to subsequently and expressly include nominal species in Nemostiromorpha (ICZN 1999, Article 67.2.2), by including the species Disemalongicornis Mäklin, 1875, in association with this name.

Nemostiropsis Borchmann, 341 [F]. Type Sorapurpureipennis Borchmann, 1930, by original designation. Status: valid subgenus of Sora Walker, 1859 in Lagriinae: Lagriini: Statirina.

Neoabantis Gebien, 341 [F]. Type species [automatic]: Abantisaenescens Fairmaire, 1892, by monotypy. Status: junior synonym of Diphyrrhynchus Fairmaire, 1849 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Gebien (1938a: 407). Note: replacement name for Abantiades Fairmaire, 1894.

Neoadelium Carter, 259 [N]. Type Adeliumfairmairei Bates, 1873, by subsequent designation (Gebien 1942a: 742). Status: valid genus in Lagriinae: Adeliini.

Neoatractus Borchmann, 713 [M]. Type species [automatic]: Atractusviridis Boisduval, 1835, by subsequent designation (Duponchel and Chevrolat 1841: 312). Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324). Note: unnecessary replacement name for Atractus Boisduval, 1835.

Neobaphion Blaisdell, 390 [N]. Type Eleodesplanipennis J.L. LeConte, 1866, by monotypy. Status: valid genus in Blaptinae: Amphidorini.

Neoblaps Ren & Li, 310 [F]. Type Neoblapshuizensis Ren & Li, 2001, by monotypy. Status: junior synonym of Coelocnemodes Bates, 1879 in Blaptinae: Blaptini: Blaptina. Synonymy: Ren et al. (2016: 432).

Neocabirutus Kulzer, 221 [M]. Type Neocabirutusindicus Kulzer, 1964, by monotypy. Status: valid subgenus of Cabirutus Strand, 1929 in Blaptinae: Pedinini: Pedinina.

Neocaedius Pierre, 960, 967 [M]. Type Caediushalli Kaszab, 1949, by original designation. Status: junior synonym of Cyptus Gerstaecker, 1871 in Blaptinae: Opatrini: Ammobiina. Synonymy: Ferrer (1996: 254, 2003: 297).

Neocamaria Kulzer, 52 [F]. Type Neocamariatibialis Kulzer, 1954, by original designation. Status: junior synonym of Robustocamaria Pic, 1922 in Stenochiinae: Cnodalonini. Synonymy: Masumoto (1993b: 222).

Neocisba Reitter, 154 [F]. Type Pachychiladissecta Kraatz, 1865, by monotypy. Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Neocistela Borchmann, 713 [F]. Type species [automatic]: Pseudocistelaovalis Blackburn, 1891, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: replacement name for Pseudocistela Blackburn, 1891.

Neodissonomus G.S. Medvedev, 235 [M]. Type Heterophylusangustitarsis Reitter, 1896, by original designation. Status: valid subgenus of Dissonomus Jacquelin du Val, 1861 in Tenebrioninae: Dissonomini.

Neoeutrapela Bousquet & Bouchard, 61 [F]. Type species [automatic]: Crioceriselongata Fabricius, 1781 (= Chrysomelaunifasciata DeGeer, 1778), by subsequent designation (Duponchel 1844b: 533). Status: valid subgenus (stat. nov. [OM]) of Impressosora Pic, 1952 in Lagriinae: Lagriini: Statirina. Note: replacement name for Eutrapela Dejean, 1834; new placement [OM], previously included in Lagriinae: Lagriini: Lagriina.

Neognathosia Kaszab, 383 [F]. Type Gnathosiapseudanemia Reitter, 1915, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Neogria Borchmann, 222 [F]. Type Neogriasulcipennis Borchmann, 1911, by subsequent designation (R. Lucas 1920: 436). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Neohelops Dajoz, 356 [M]. Type Neohelopstexanus Dajoz, 2001, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina. Note: placed in the subtribe Helopina by Nabozhenko and Purchart (2019: 156).

Neohyocis G.S. Medvedev & Lawrence, 571 [M]. Type Hyociswaterhousei G.S. Medvedev & Lawrence, 1983, by original designation. Status: valid subgenus of Hyocis Pascoe, 1866 in Diaperinae: Hyociini: Hyociina.

Neoisocerus Bouchard, Lawrence, Davies & Newton, 510 [M]. Type species [automatic]: Tenebriopurpurascens Herbst, 1799 (= Helopsferrugineus Fabricius, 1798), by monotypy. Status: valid genus in Blaptinae: Dendarini: Dendarina. Note: replacement name for Isocerus Dejean, 1821.

Neomenimus Kaszab, 190 [M]. Type Neomenimusclavatus Kaszab, 1939, by original designation. Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: G.S. Medvedev (2007b: 665).

Neomida Latreille, 29 [F]. Type Ips haemorrhoidalis Fabricius, 1787, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: Neomida was used earlier by Dahl (1823: 44) but Dahl’s work was suppressed for the purposes of zoological nomenclature by the ICZN (1964, Opinion 710).

Neooligocara Guerrero, Vidal & Moore, 407 [N]. Type Oligocarabucki Kulzer, 1962, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Neopachypterus Bouchard, Löbl & Merkl, 386 [M]. Type species [automatic]: Pachypterusmauritanicus P.H. Lucas, 1847, by monotypy. Status: valid genus in Blaptinae: Opatrini: Neopachypterina. Note: replacement name for Pachypterus P.H. Lucas, 1847.

Neophaleria Español, 74, 76 [F]. Type Phaleriaardoini Español, 1963, by monotypy. Status: valid subgenus of Phaleria Latreille, 1802 in Diaperinae: Phaleriini.

Neophylax Bedel, 92 [M]. Type species [automatic]: Phylaxlittoralis Mulsant, 1854 (= Opatrumpicipes G.-A. Olivier, 1812), by subsequent designation (Gebien 1938a: 412). Status: senior synonym of Allophylax Bedel, 1906 in Blaptinae: Dendarini: Melambiina. Note: junior homonym of Neophylax McLachlan, 1871 [Trichoptera].

Neoplamius Löbl, Bouchard, Merkl & Bousquet, 4 [M]. Type Neoplamiuszoltani Masumoto, 1981, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: name first proposed by Masumoto (1981: 15) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Neoplateia Marcuzzi, 177 [F]. Type Neoplateiakaszabi Marcuzzi, 1986, by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Neoplonyx Ardoin, 309, 352 [M]. Type Gonocnemissulcicollis Fairmaire, 1899, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Neoporphyrhyba Ardoin, 89 [F]. Type Porphyrhybacyaneocuprea Fairmaire, 1894, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Neopraocis Kulzer, 3, 6 [M]. Type Praocisreflexicollis Solier, 1851, by monotypy. Status: valid genus in Pimeliinae: Praociini.

Neopsectropus Kaszab, 30 [M]. Type Neopsectropusgebieni Kaszab, 1941, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Neopterocoma Skopin, 158 [F]. Type Pterocomabalchashensis Skopin, 1974, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Neosolenopistoma Bouchard & Bousquet, new subgenus. [N]. Type Eurynotusdenticosta Mulsant & Rey, 1854, by present designation. Status: valid subgenus of Eurynotus W. Kirby, 1819 in Blaptinae: Platynotini: Eurynotina. Note: taxon first proposed by Mulsant and Rey (1854: 29; as “Solenopistoma Solier”); however, this is treated as an incorrect subsequent spelling of Selenepistoma Dejean, 1834 (see Bousquet and Bouchard 50); the subgenusSolenopistoma Mulsant & Rey, 1854, which is currently used as valid, is therefore unavailable (ICZN 1999, Articles 33.3); we hereby make the name Neosolenopistoma available by selecting Eurynotusdenticosta Mulsant & Rey, 1854 as type species and referring to Mulsant and Rey (1854: 29) for the character states that characterise and differentiate Neosolenopistoma.

Neotagalus Kaszab, 471, 477 [M]. Type Neotagalustuberculiger Kaszab, 1955, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Neotheca Carter, 540 [F]. Type Neothecafusca Carter, 1930, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Neozophobas Ferrer, 235 [M]. Type Zophobaslaticollis Kraatz, 1880, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini.

Nepalindia G.S. Medvedev, 187 [F]. Type Tagonoidesalpina Kaszab, 1965, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Nepalofranziella Fouquè, 194, 196 [F]. Type Nepalofranziellakaszabi Fouquè, 2013, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Nepalolaena Schawaller, 277 [F]. Type Nepalolaenakira Schawaller, 2001, by original designation. Status: valid genus in Lagriinae: Laenini.

Nepaloplonyx Bremer, 175, 181 [M]. Type Nepaloplonyxcaelebs Bremer, 2014, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Nephodes Blanchard, 34 [M]. Type Nephodesvilliger Rosenhauer, 1856, by subsequent monotypy (Rosenhauer 218). Status: senior synonym of Nephodinus Gebien, 1943 in Tenebrioninae: Helopini: Helopina. Note: originally proposed without included nominal species; Rosenhauer (1856: 218), by including the new species Nephodesvilliger Rosenhauer, 1856 in association with this name, was the first author to subsequently and expressly include nominal species in Nephodes (ICZN 1999, Article 67.2.2); the older name Nephodes Schönherr, 1840 [Coleoptera: Curculionidae] is available as it was originally proposed as a synonym and subsequently treated as a senior homonym by Gebien (1943: 900; see ICZN 1999, Article 11.6.1), therefore Nephodes Blanchard, 1845 is a junior homonym of Nephodes Schönherr, 1840.

Nephodinus Gebien, 900 [M]. Type species [automatic]: Nephodesvilliger Rosenhauer, 1856, by subsequent monotypy (Rosenhauer 218). Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina. Note: replacement name for Nephodes Blanchard, 1845.

Nerina Lacordaire, 70 [F]. Type Nerinadispar Lacordaire, 1859, by monotypy. Status: senior synonym of Afrinus Fairmaire, 1888 in Pimeliinae: Tentyriini. Synonymy: Blair (1935a: 103). Note: junior homonym of Nerina Robineau-Desvoidy, 1830 [Diptera].

Nerinodon Koch, 138 [M]. Type Nerinodoncaviceps Koch, 1952, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Nesioticus Westwood, 120 [M]. Type Nesioticusflavopictus Westwood, 1843, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: redescribed as new by Westwood (1844: 227).

Nesocaedius Kolbe, 262 [M]. Type Nesocaediusschultzei Kolbe, 1915, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Nesocyrtosoma Marcuzzi, 137 [N]. Type Cyrtosomainflatum Marcuzzi, 1976, by plenary powers (ICZN 2017, Opinion 2398). Status: valid genus in Stenochiinae: Cnodalonini. Note: following an application by Hopp et al. (2014) to conserve the generic name Nesocyrtosoma Marcuzzi, 1976, the ICZN (2017, Opinion 2398) determined that this name is nomenclaturally available despite not having been accompanied by a type species fixation in the original publication, placed it on the Official List of Generic Names in Zoology and, designated Cyrtosomainflatum Marcuzzi, 1976 as its type species.

Nesogena Mäklin, 557 [F]. Type Nesogenahybrida Mäklin, 1863, by subsequent designation (Gebien 1948: 549). Status: valid genus and subgenus in Tenebrioninae: Praeugenini.

Nesogenomorpha Pic, 18 [F]. Type Nesogenomorphasemiviolacea Pic, 1917, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Nesopatrum Gebien, 20 [N]. Type Opatrinusjosephi Karsch, 1881, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Nesophaerotus Ardoin, 66, 67 [M]. Type species [automatic]: Nesosphaerotusaeneus Gebien, 1921, by subsequent designation (Gebien 1943: 403). Status: junior synonym of Nesosphaerotus Gebien, 1921 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Nesosphaerotus Gebien, 1921, not in prevailing usage.

Nesosphaerotus Gebien, 115 [M]. Type Nesosphaerotusaeneus Gebien, 1921, by subsequent designation (Gebien 1943: 403). Status: valid genus in Stenochiinae: Cnodalonini.

Nesostes Casey, 56, 58 [M]. Type Eusattusrobustus J.L. LeConte, 1866, by original designation. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: Triplehorn (1968: 379).

Nesotaurus Fairmaire, 354 [M]. Type Nesotaurussericans Fairmaire, 1896, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Nesotes Allard, 5 [M]. Type Helopsasper Küster, 1850, by subsequent designation (Nabozhenko 2008: 38). Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina. Note: the First Reviser (Nesotes Allard, 1876 versus Diastixus Allard, 1876) is Antoine (1949: 134).

Netopha Fairmaire, 299 [F]. Type Netophapallidipes Fairmaire, 1893, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Netuschilia Reitter, 34, 35 [F]. Type Lachnopushauseri Reitter, 1897, by monotypy. Status: valid genus in Pimeliinae: Lachnogyini: Netuschiliina.

Nevermanniella Borchmann, 235, 332 [F]. Type Statiraalbolineata Champion, 1889, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: the alternative original spelling Nevermannia (pp. 11, 235) was corrected to Nevermanniella in the “Corrigenda” of the same work (p. 541), Nevermanniella is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1)

Nevisia Marcuzzi, 179 [F]. Type Diastolinusbardudensis Marcuzzi, 1962, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Nicandra Fairmaire, 191 [F]. Type Nicandracostulipennis Fairmaire, 1888, by monotypy. Status: valid genus and subgenus in Blaptinae: Pedinini: Helopinina.

Nikomenalia Dubrovina, 166 [F]. Type Hymenaliakaszabi Muche, 1972, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: elevated from subgenus of Hymenalia Mulsant, 1856 by Novák (2020c: 507).

Nilio Latreille, 179 [M]. Type Coccinellavillosa Fabricius, 1787, by monotypy. Status: valid genus and subgenus in Nilioninae. Note: Nilio is an incorrect subsequent spelling of the original spelling Nilion, first used by Latreille (1804: 333), in prevailing usage; Nilio is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1); the original combination of the name of the type species, Coccinellavillosa Fabricius, 1787, is a junior primary homonym of Coccinellavillosa Fourcroy, 1785.

Nipponalassus Nabozhenko & Ando, 306 [M]. Type Tarpelaandoi Masumoto, 1993, by original designation. Status: valid subgenus of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina.

Nipponohelops Masumoto, Ando & Akita, 33 [M]. Type Nipponohelopsishikawai Masumoto, Ando & Akita, 2006, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Nocar Blackburn, 328 [N]. Type Cisteladepressiuscula W.J. MacLeay, 1872, by subsequent designation (R. Lucas 1920: 441). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Nochelius Gistel, xi [M]. Type species [automatic]: Evaniosomusorbignianus Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Evaniosomus Guérin-Méneville, 1834 in Pimeliinae: Evaniosomini. Note: unnecessary replacement name for Evaniosomus Guérin-Méneville, 1834, not in prevailing usage.

Nocibiotes Casey, 617 [M]. Type Notibiusgranulatus J.L. LeConte, 1851, by subsequent designation (Gebien 1938a: 407). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Nodosogylium Pic, 12 [N]. Type Nodosogyliuminaequale Pic, 1951, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: we act as First Revisers and reject the alternative original spelling Nodosogilium, used by Pic (1951: 12).

Nodotelus Koch, 67 [M]. Type species [automatic]: Eutelusrequieni Solier, 1843, by subsequent designation (R. Lucas 1920: 291). Status: junior synonym of Eutelonotus Fairmaire in Alluaud, 1902 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Eutelus Solier, 1843.

Nolicima Matthews, 708, 714 [F]. Type Cardiothoraxangusticollis Carter, 1906, by original designation. Status: valid genus in Lagriinae: Adeliini.

Notacula Campbell, 107 [F]. Type Notaculahowdenae Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Nothogria Borchmann, 49, 142 [F]. Type Nothogrianodipennis Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Nothrocerus Fairmaire, 174 [M]. Type Nothroceruscylindricornis Fairmaire, 1887, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Notiasida Casey, 76, 124 [F]. Type Notiasidaabstrusa Casey, 1912, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Notibius J.L. LeConte, 144 [M]. Type Notibiuspuberulus J.L. LeConte, 1851, by subsequent designation (Gebien 1938a: 406). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Notiolesthus Motschulsky, 25 [M]. Type Notiolesthusnatalensis Motschulsky, 1872, by original designation. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1921b: 69). Note: we act as First Revisers and reject the alternative original spelling Notiolosthus, used by Motschulsky (1872: 34).

Notioscythis Fairmaire, 31 [F]. Type Notioscythispunctoseriata Fairmaire, 1883, by monotypy. Status: junior synonym of Stenosida Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Chatanay (1917: 231).

Notoblaps Bauer, 232 [F]. Type Blaps juliae Allard, 1881, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Notocistela Carter, 78, 102 [F]. Type Notocistelatibialis Carter, 1915, by subsequent designation (Matthews and Bouchard 2008: 333). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Notocorax Dejean, 191 [M]. Type Opatrumjavanum Wiedemann, 1819, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Notoprataeus Carter, 37 [M]. Type Notoprataeuslitoralis Carter, 1924 (= Mesotertisinconstans Lea, 1917), by monotypy. Status: junior synonym of Micropedinus Lewis, 1894 in Lagriinae: Lupropini. Synonymy: G.S. Medvedev (1992: 656), Matthews and Lawrence (2005: 534).

Notostrongylium Carter, 523 [N]. Type Notostrongyliumrugosicolle Carter, 1915, by subsequent designation (Kulzer 1966: 387). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Kaszab (1977b: 3, 29).

Nototrintus Carter, 40 [M]. Type Otrintusjacksoni Carter, 1905, by original designation. Status: valid genus in Lagriinae: Adeliini.

Nudoplatyscelis Kaszab, 149, 222 [F]. Type Platynoscelisturanica Reitter, 1896, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Nuptis Motschulsky, 25 [M]. Type Nuptistenuis Motschulsky, 1872, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Nyctalops Gistel, 125 [M]. Type species [automatic]: Pseudoblapssubstriata Guérin-Méneville, 1834, by subsequent designation (Hope 1841: 124). Status: junior synonym of Pseudoblaps Guérin-Méneville, 1834 in Blaptinae: Platynotini: Platynotina. Note: unnecessary replacement name for Pseudoblaps Guérin-Méneville, 1834; junior homonym of Nyctalops Wagler, 1832 [Aves].

Nyctelia Berthold, 367 [F]. Type Zophosisnodosa Germar, 1823 (= Zophosispicipes Billberg, 1815), by monotypy. Status: valid genus in Pimeliinae: Nycteliini. Note: nomen protectum (see Silvestro and Flores 2016: 658).

Nyctelioma Casey, 163 [F]. Type Nycteliomaexplanata Casey, 1908, by original designation. Status: junior synonym of Nyctelia Berthold, 1827 in Pimeliinae: Nycteliini. Synonymy: Gebien (1910a: 141).

Nyctelius Guérin-Méneville, 21 [M]. Type Zophosisnodosa Germar, 1823 (= Zophosispicipes Billberg, 1815), by monotypy. Status: senior synonym of Nyctelia Berthold, 1827 in Pimeliinae: Nycteliini. Synonymy: Silvestro and Flores (2016: 658). Note: nomen oblitum (see Silvestro and Flores 2016: 658).

Nycterinus Eschscholtz, 9.13 [M]. Type Nycterinusthoracicus Eschscholtz, 1829, by subsequent designation (Hope 1841: 124). Status: valid genus in Blaptinae: Amphidorini.

Nycteropus Klug, 89 [M]. Type Nycteropusebeninus Klug, 1833, by subsequent designation (Hope 1841: 124). Status: valid genus in Tenebrioninae: Toxicini: Nycteropina.

Nyctipates Gebler, 373 [M]. Type Nyctipatesrugulosus Gebler, 1841, by monotypy. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Erichson (1845a: 116).

Nyctipates Solier, 154, 285 [M]. Type Nyctipatescoriaceus Solier, 1848 (= Blapsangustata Zubkov, 1833), by subsequent designation (Löbl et al. 2008c: 236). Status: senior synonym of Prosodopria Reitter, 1909 in Blaptinae: Blaptini: Prosodina. Synonymy: Löbl et al. (2008a: 43). Note: junior homonym of Nyctipates Gebler, 1841 [Coleoptera: Tenebrionidae: Blaptinae: Blaptini: Prosodina].

Nyctobates Guérin-Méneville, 33 [M]. Type Tenebriogigas Linnaeus, 1763, by original designation. Status: junior synonym of Mylaris Pallas, 1781 in Stenochiinae: Cnodalonini. Synonymy: Spilman (1973: 42), Ferrer and Siliansky (2008: 186).

Nyctopetus Guérin-Méneville, pl. 4 [M]. Type Nyctopetustenebrioides Guérin-Méneville, 1831, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Nyctoporis Eschscholtz, 10, 11 [F]. Type Nyctoporiscristata Eschscholtz, 1831, by subsequent designation (Hope 1841: 124). Status: valid genus in Pimeliinae: Nyctoporini.

Nyctozoilus Guérin-Méneville, pl. 4 [M]. Type Nyctozoilusobesus Guérin-Méneville, 1831, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Nypsius Champion, 219 [M]. Type Nypsiusaeneopiceus Champion, 1895, by subsequent designation (R. Lucas 1920: 447). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Oatesius Westwood, 376 [M]. Type species [automatic]: Derosphaeriusanthracinus Westwood, 1881, by monotypy. Status: junior synonym of Derosphaerius Westwood, 1881 in Pimeliinae: Tentyriini. Note: unnecessary replacement name for Derosphaerius Westwood, 1881.

Obenbergeria Strand, 24 [F]. Type species [automatic]: Diaperisriederii Faldermann, 1833, by subsequent designation (Jakobson 1924: 243). Status: junior synonym of Emypsara Pascoe, 1866 in Diaperinae: Phaleriini. Synonymy: Gebien (1939: 743). Note: replacement name for Callicomus Motschulsky, 1860.

Obesacula Campbell, 109 [F]. Type Obesaculaaptera Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Oblongoodescelis Kaszab, 941, 958 [F]. Type Platyscelisoblonga Ballion, 1878, by original designation. Status: junior synonym of Clavatoodescelis Kaszab, 1940 in Blaptinae: Platyscelidini. Synonymy: Egorov (2020: 380).

Oblongoplatyscelis Kaszab, 910, 916 [F]. Type Platyscelisganglbaueri Seidlitz, 1893, by original designation. Status: valid subgenus of Platyscelis Latreille, 1818 in Blaptinae: Platyscelidini.

Obriomaia Gebien, 45 [F]. Type Eucyrtussubcostatus Fairmaire, 1893, by original designation. Status: junior synonym of Tetragonomenes Chevrolat, 1878 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 133).

Occidentophosis Penrith, 18, 157 [F]. Type Zophosisdamarina Péringuey, 1908, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Ochrolamus Reitter, 73 [M]. Type Dilamuspictus Baudi di Selve, 1881, by monotypy. Status: valid subgenus of Dilamus Jacquelin du Val, 1861 in Blaptinae: Opatrini: Ammobiina.

Ocnera Fischer, 169 [F]. Type Pimeliaimbricata Fischer, 1820, by subsequent designation (Wilke 1922: 260). Status: valid genus in Pimeliinae: Pimeliini. Note: as mentioned by Bouchard and Bousquet (2020b: 7) the nomenclatural stability of this name is threatened by the discovery of an older type species designation (Tenebriocephalotes Pallas, 1781, by subsequent designation by Crotch (1870b: 241), currently the type species of the valid subgenusChaetotoma Motschulsky, 1860 in Pimelia Fabricius, 1775); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Gebien (1937a: 814).

Ocnodes Fåhraeus, 270 [F]. Type Ocnodesscrobicollis Fåhraeus, 1870, by subsequent designation (Kamiński et al. 2019b: 55). Status: valid genus and subgenus in Pimeliinae: Sepidiini: Molurina.

Ocularisora Pic, 32 [F]. Type Nemostirabenitensis Pic, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Oculochara Novák, 80 [F]. Type Mycetocharaocularis Reitter, 1884, by original designation. Status: valid subgenus of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina.

Oculosis Penrith, 18, 126 [F]. Type Zophosisboei Solier, 1834, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Odocnemis Allard, 4 [F]. Type Odocnemiscaudata Allard, 1876 (= Helopspraelongus Baudi di Selve, 1876), by subsequent designation (Nabozhenko 2001a: 662). Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina. Note: the First Reviser (Odocnemis Allard, 1876 versus Omaleis Allard, 1876) is Nabozhenko (2001a: 628).

Odontocera Chen & Yuan, 183 [F]. Type Odontoceraqinlingensis Chen & Yuan, 1996, by original designation. Status: senior synonym of Odontocerostira Merkl, 2007 in Lagriinae: Lagriini: Statirina. Note: junior homonym of Odontocera Audinet-Serville, 1834 [Coleoptera: Cerambycidae].

Odontocerostira Merkl, 269 [F]. Type species [automatic]: Odontoceraqinlingensis Chen & Yuan, 1996, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: replacement name for Odontocera Chen & Yuan, 1996.

Odontocnemis Rye, 69 [F]. Type species [automatic]: Odocnemiscaudata Allard, 1876 (= Helopspraelongus Baudi di Selve, 1876), by subsequent designation (Nabozhenko 2001a: 662). Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Note: unjustified emendation of Odocnemis Allard, 1876, not in prevailing usage.

Odontogria Borchmann, 16, 62 [F]. Type Lagriablairi Borchmann, 1925, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Odontomophlus Seidlitz, 240 [M]. Type Omophlusochraceipennis Faldermann, 1837, by subsequent designation (Bousquet et al. 2015: 141). Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini. Note: as pointed out by Bousquet et al. (2015: 141) the type species of this genus was first designated by Novák and Pettersson (2008: 335) who selected Cistelaarmillata Brullé, 1832, a species originally included in Odontomophlus Seidlitz, 1896 but also the type species of the valid genus Megischina Reitter, 1906; Löbl and Smetana (2011: 34) “retracted” the act proposed earlier by Novák and Pettersson (2008: 335); in order to conserve nomenclatural stability Bousquet et al. (2015: 141) recommended usage of Omophlusochraceipennis Faldermann, 1837 as the type species of this genus until an application be submitted to the Commission to rule on the Case.

Odontopezus Alluaud, xlv [M]. Type species [automatic]: Odontopuscostatus Silbermann, 1833, by subsequent designation (Hope 1841: 126). Status: junior synonym of Pezodontus Dejean, 1834 in Lagriinae: Pycnocerini. Note: unnecessary replacement name for Odontopus Silbermann, 1833.

Odontopus Silbermann, no 3 [M]. Type Odontopuscostatus Silbermann, 1833, by subsequent designation (Hope 1841: 126). Status: senior synonym of Pezodontus Dejean, 1834 in Lagriinae: Pycnocerini. Note: junior homonym of Odontopus Say, 1831 [Coleoptera: Curculionidae].

Odrotes La Rivers, 320 [M]. Type Edrotesarens La Rivers, 1947, by monotypy. Status: valid subgenus of Edrotes J.L. LeConte, 1851 in Pimeliinae: Edrotini.

Oeatus Champion, 111 [M]. Type Oeatuschevrolati Champion, 1885, by subsequent designation (Gebien 1941: 342). Status: valid genus in Stenochiinae: Cnodalonini.

Oectosis Pascoe, 149 [F]. Type Upiscylindrica Germar, 1848, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Oedemutes Pascoe, 51 [M]. Type Oedemutestumidus Pascoe, 1860, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Oedenocera Reiche, 372 [F]. Type species [automatic]: fixed herein (ICZN 1999, Article 70.3) as Tenebriobuprestoides Fabricius, 1781, misidentified as Akislaevigata Fabricius, 1801 in the original designation by monotypy in Eschscholtz (1831). Status: junior synonym of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Löbl et al. (2008b: 192). Note: replacement name for Pachycera Eschscholtz, 1831. Note: see additional information in the entry for Pachycera Eschscholtz, 1831.

Oenomia Pascoe, 441 [F]. Type Oenomiafemorata Pascoe, 1883, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Oenopion Champion, 98 [M]. Type Oenopiongibbosus Champion, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ogoueum Pic, 27 [N]. Type Ogoueumsemirufum Pic, 1923, by monotypy. Status: junior synonym of Eccoptostoma Gebien, 1913 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1962a: 64).

Ograbies Péringuey, 293 [M]. Type Ograbiessingularis Péringuey, 1899 (= Oncotustestaceus Solier, 1848), by monotypy. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Ohyonthis Reitter, 347 [F]. Type Ohyonthismicroderoides Reitter, 1898, by monotypy. Status: junior synonym of Stegastopsis Kraatz, 1865 in Pimeliinae: Tentyriini. Synonymy: Reitter (1900c: 89, 139).

Oligocara Solier, 153, 224 [N]. Type Oligocaranitidum Solier, 1848, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Oligorus Dejean, 206 [M]. Type Tageniaindica Wiedemann, 1823, by monotypy. Status: junior synonym of Luprops Hope, 1833 in Lagriinae: Lupropini. Synonymy: Lacordaire (1859a: 397, as “Lyprops”).

Oliprosodes Reitter, 118 [M]. Type Prosodestrisulcata Bates, 1879, by subsequent designation (G.S. Medvedev 1999a: 851). Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Olisthaena Erichson, 177 [F]. Type Olisthaenanitida Erichson, 1842, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Olocrates Mulsant, 150, 383 [M]. Type Opatrumgibbum Fabricius, 1775, by subsequent designation (C.G. Thomson 1859: 115, as “Omocrates”). Status: junior synonym of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina. Synonymy: Seidlitz (1898a: 828). Note: the original spelling Omocrates (p. 150) was corrected to Olocrates in the “Errata” of the same work (p. 383), Olocrates is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1); see Bouchard and Bousquet (2020a: 100).

Ologlyptus Lacordaire [in LeConte], 19 [M]. Type species [automatic]: Stenosidesgraciliformis Solier, 1836, by monotypy. Status: junior synonym of Stenosides Solier, 1836 in Pimeliinae: Asidini. Note: unnecessary replacement name for Stenosides Solier, 1836 (as Stenorides).

Omala Agassiz, 184 [F]. Type species [automatic]: Homalapolita Eschscholtz, 1831, by monotypy. Status: junior synonym of Homala Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: unjustified emendation of Homala Eschscholtz, 1831, not in prevailing usage.

Omaleis Allard, 4 [M]. Type Helopscongener Reiche, 1861, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: Nabozhenko (2001a: 628). Note: Allard’s original spelling Omalus, which is a junior homonym of Omalus Panzer, 1801 [Hymenoptera], was corrected to “Omaloïs” [= Omalois] in the errata for volume 14 of L’Abeille, Journal d’Entomologie (at the end of page 36 of the section “Table alphabétique”) and therefore Omalois should be considered the correct original spelling (ICZN 1999, Article 32.5.1.1); however, the incorrect subsequent spelling Omaleis, which was introduced by Allard (1877: 36), is in prevailing usage and attributed to the original author, it is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Omandelostoma Purchart, 282 [N]. Type Adelostomamuqalensis Purchart, 2017, by original designation. Status: valid subgenus of Adelostoma Duponchel, 1827 in Pimeliinae: Adelostomini.

Omedes Broun, 1169 [M]. Type Omedesnitidus Broun, 1893, by subsequent designation (R. Lucas 1920: 456). Status: valid genus in Alleculinae: Alleculini: incertae sedis.

Omegeleodes Triplehorn & Thomas, 253 [M]. Type Eleodesdebilis J.L. LeConte, 1858, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Ommatochara Borchmann, 347 [F]. Type Ommatocharatibialis Borchmann, 1932, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Ommatophorus W.J. MacLeay, 304 [M]. Type Ommatophorusmastersii W.J. MacLeay, 1872, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Omocula Borchmann, 222 [F]. Type Alleculacollaris Borchmann, 1930, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Omolepta Fåhraeus, 320 [F]. Type Omoleptaelegans Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Omolipus Pascoe, 127 [M]. Type Omolipuscorvus Pascoe, 1860, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Omopheres Casey, 519 [M]. Type Omopheresfarctus Casey, 1907, by original designation. Status: valid genus and subgenus in Pimeliinae: Epitragini.

Omophlina Reitter, 34 [F]. Type Omophluspodontoides Reitter, 1890, by subsequent designation (Novák and Pettersson 2008: 334). Status: valid genus in Alleculinae: Cteniopodini.

Omophlus Dejean, 213 [M]. Type Cistelalepturoides Fabricius, 1787, by subsequent designation (Solier 1835a: 246). Status: valid genus and subgenus in Alleculinae: Cteniopodini. Note: the earlier usage of the name Omophlus by Dahl (1823: 46) was suppressed for the purposes of zoological nomenclature by the ICZN (1964, Opinion 710).

Oncopterus Fairmaire, 178 [M]. Type Oncopterusacantholophus Fairmaire, 1887, by monotypy. Status: senior synonym of Oncopteryx Gebien, 1943 in Blaptinae: Pedinini: Helopinina. Note: junior homonym of Oncopterus Steindachner, 1875 [Pisces].

Oncopteryx Gebien, 905 [F]. Type species [automatic]: Oncopterusacantholophus Fairmaire, 1887, by monotypy. Status: valid genus in Blaptinae: Pedinini: Helopinina. Note: replacement name for Oncopterus Fairmaire, 1887.

Oncosoma Westwood, 121 [N]. Type Oncosomagranulare Westwood, 1843 (= Pimeliagemmata Fabricius, 1801), by monotypy. Status: junior synonym of Amatodes Dejean, 1834 in Blaptinae: Pedinini: Helopinina. Synonymy: Erichson (1844: 282). Note: unjustified emendation of the original spelling Ogcosoma, introduced by Agassiz (1846b: 257, 259), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1), see Bouchard et al. (2005: 509); redescribed by Westwood (1844: 227) under the name Ogcoosoma; the original combination of the accepted name of the type species, Pimeliagemmata Fabricius, 1801, is a junior primary homonym of Pimeliagemmata Herbst, 1799.

Oncoosoma Gebien, 563 [N]. Type species [automatic]: Oncosomagranulare Westwood, 1843 (= Pimeliagemmata Fabricius, 1801), by monotypy. Status: junior synonym of Amatodes Dejean, 1834 in Blaptinae: Pedinini: Helopinina. Note: unjustified emendation of Oncosoma Westwood, 1843 (as “Ogcosoma”), not in prevailing usage.

Oncotiphallops Koch, 162 [M]. Type Oncotiphallopsbarbosai Koch, 1956, by original designation. Status: junior synonym of Anchophthalmus Gerstaecker, 1854 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (2002a: 53).

Oncotopsis Koch, 152 [F]. Type Nicandrabicolor Kulzer, 1951, by original designation. Status: valid subgenus of Nicandra Fairmaire, 1888 in Blaptinae: Pedinini: Helopinina.

Oncotus Blanchard, 13, 24 [M]. Type Oncotusfarctus Solier, 1848, by subsequent designation (Gebien 1938a: 72). Status: valid genus and subgenus in Blaptinae: Platynotini: Eurynotina. Note: originally proposed without included nominal species; Solier (1848: 216–222), by including five new species in association with the genus “Oncotus Dejean”, was the first author to subsequently and expressly include nominal species in Oncotus (ICZN 1999, Article 67.2.2).

Onocera Borchmann, 139 [F]. Type Ecnolagriasecurigera Borchmann, 1916, by original designation. Status: valid subgenus of Ecnolagria Borchmann, 1916 in Lagriinae: Lagriini: Lagriina.

Onoglypta Carter, 144 [F]. Type species [automatic]: Aglyptaoctocostata Gebien, 1908, by monotypy. Status: junior synonym of Aglypta Gebien, 1908 in Tenebrioninae: Heleini: Cyphaleina. Note: unnecessary replacement name for Aglypta Gebien, 1908.

Ononyctus Carter, 382 [M]. Type Ononyctussulcatus Carter, 1914, by monotypy. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 473).

Onosterrhus Pascoe, 451 [M]. Type Onosterrhuslaevis Pascoe, 1866, by monotypy. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 473).

Onotrichus Carter, 138, 164 [M]. Type Onotrichuslateralis Carter, 1911, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Onychomira Campbell, 289 [F]. Type Onychomirafloridensis Campbell, 1984, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Onychosis Deyrolle, 226 [F]. Type Onychosisgracilipes Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Note: the name Anisosis, also described in the same publication, was used in error for Onychosis in the “Tableau des genres” on p. 81 (see Dallas 1868: 265–266).

Onymacris Allard, 157 [F]. Type Adesmiacandidipennis Brême, 1840, by subsequent designation (Gebien 1937a: 655). Status: valid genus in Pimeliinae: Adesmiini.

Oochila J.L. LeConte, 220 [F]. Type Asbolusinfaustus J.L. LeConte, 1854, by original designation. Status: junior synonym of Cryptoglossa Solier, 1837 in Pimeliinae: Cryptoglossini. Synonymy: Horn (1870: 278, with Centrioptera Mannerheim, 1843, a junior synonym of Cryptoglossa Solier, 1837).

Oochrotus P.H. Lucas, xxix [M]. Type Oochrotusunicolor P.H. Lucas, 1852, by monotypy. Status: valid genus in Diaperinae: Crypticini.

Oocistela Borchmann, 356 [F]. Type Oocistelaconvexa Borchmann, 1908, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Ooconibius Casey, 618 [M]. Type Notibiusopacus J.L. LeConte, 1866, by monotypy. Status: junior synonym of Conibius J.L. LeConte, 1851 in Blaptinae: Opatrini: Blapstinina. Synonymy: Aalbu in Bousquet et al. (2018: 202).

Oodeoscelis Agassiz, 260 [F]. Type species [automatic]: Blapspolita Sturm, 1807, by subsequent designation (Kaszab 1940b: 937; see ICZN 1993b, Opinion 1729). Status: junior synonym of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini. Note: unjustified emendation of Oodescelis Motschulsky, 1845, not in prevailing usage.

Oodescelis Motschulsky, 76 [F]. Type Blapspolita Sturm, 1807, by subsequent designation (Kaszab 1940b: 937; see ICZN 1993b, Opinion 1729). Status: valid genus and subgenus in Blaptinae: Platyscelidini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1993b, Opinion 1729).

Oogaster Faldermann, 30 [F]. Type Oogastermenetriesii Faldermann, 1837 (= Tageniapicea Ménétriés, 1832), by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Oogeton Kaszab, 69 [M]. Type Oogetonnigrocoeruleus Kaszab, 1941 (= Amarygmusmakii Miwa, 1939), by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Note: name first used by Miwa (1939: 412) without type species fixation (see Masumoto 1989a: 116; Löbl et al. 2008a: 44).

Oopiestus Chevrolat, 30, pl. 2 [M]. Type Oopiestusovalis Chevrolat, 1833 (= Peltoidessenegalensis Laporte, 1833), by monotypy. Status: senior synonym of Peltoides Laporte, 1833 in Tenebrioninae: Alphitobiini. Synonymy: Audoin and Milne-Edward (1835: 188). Note: the alternative original spelling Opiestus, used by Chevrolat (1833b: 30), was rejected by Chevrolat (1847a: 118) who acted as the First Reviser (ICZN 1999, Article 24.2.4); bibliographic evidence indicates that Oopiestus Chevrolat, 1833 was published by 16 March 1833 before Peltoides Laporte, 1833, issued by 1 April 1833, and should be treated as the valid name instead; an application to the ICZN is necessary to conserve usage of Peltoides Laporte, 1833 as the valid name.

Opacoplonyx Bremer, 35 [M]. Type Plesiophthalmusdavidis Fairmaire, 1878, by original designation. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Opatrasida Escalera, 69 [F]. Type Asidajurinei Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Polasida Reitter, 1917 in Pimeliinae: Asidini. Synonymy: Gebien (1937a: 727).

Opatresthes Gebien, 192 [F]. Type Opatresthesbinodosa Gebien, 1928, by subsequent designation (Gebien 1941: 817). Status: valid subgenus of Goniadera Perty, 1832 in Lagriinae: Goniaderini.

Opatrinus Dejean, 66 [M]. Type Opatrumclathratum Fabricius, 1787, by monotypy. Status: valid genus and subgenus in Blaptinae: Platynotini: Platynotina.

Opatroides Brullé, 219 [M]. Type Opatroidespunctulatus Brullé, 1832, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Opatronesthes Reitter, 174 [F]. Type Melanesthespunctipennis Reitter, 1889, by subsequent designation (Iwan and Löbl 2007: 734). Status: valid subgenus of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina.

Opatropis Reitter, 134, 159 [F]. Type Opatrumhispidum Brullé, 1839 (= Opatrumaffine Billberg, 1815), by monotypy. Status: valid subgenus of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina.

Opatrum Fabricius, 76 [N]. Type Silphasabulosa Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Blaptinae: Opatrini: Opatrina.

Ophthalmosis Deyrolle, 81, 229 [F]. Type Ophthalmosislongipes Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Note: the original spelling Ophtalmosis (pp. 77, 229, 231) was corrected to Ophthalmosis in a footnote in the “Table des matières” of the same work (p. clxiv), Ophthalmosis is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Opigenia Pascoe, 288 [F]. Type Opigeniairidescens Pascoe, 1869 (= Platyphanesvittatus Westwood, 1849), by monotypy. Status: junior synonym of Platyphanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 454).

Opisthoblaps Kolbe, 201 [F]. Type Blaps sulcifera Seidlitz, 1893, by subsequent designation (Nabozhenko and Chigray 2020: 10). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Oplocephala Laporte & Brullé, 332, 338 [F]. Type Ips haemorrhoidalis Fabricius, 1787, by subsequent designation (Motschulsky 1845a: 80). Status: junior synonym of Neomida Latreille, 1829 in Diaperinae: Diaperini: Diaperina. Synonymy: Dejean (1834: 197).

Oplocheirus Klug, 40 [M]. Type Helopscarbonarius Klug, 1835 (= Acanthomerusstriatus Guérin-Méneville, 1834), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Oploptera Chevrolat in Guérin-Méneville, 126 [F]. Type Strongyliumserraticorne Guérin-Méneville, 1834, by monotypy. Status: valid genus and subgenus in Stenochiinae: Stenochiini.

Opostirus Kirsch, 45 [M]. Type Opostirusexsectus Kirsch, 1865, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina. Note: transferred from Zopheridae: Colydiinae by Ivie et al. (2016: 759).

Oppenheimeria Koch, 110 [F]. Type Oppenheimeriabombophthalma Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Evaniosomini.

Oracula Novák, 54 [F]. Type Oraculabicolor Novák, 2019, by original designation. Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Orarabion Leo & Liberto, 157 [N]. Type Orarabiondominici Leo & Liberto, 2011, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Orchesiolobopoda Pic, 6 [F]. Type Orchesiolobopodaminutissima Pic, 1919, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Orcopagia Pascoe, xii [F]. Type Orcopagiamonstrosa Pascoe, 1868, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Oremasis Pascoe, 470 [M]. Type Adeliumcupreum Gray, 1831, by original designation. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 490).

Oreogria Merkl, 248 [F]. Type Oreogriakaszabi Merkl, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Oreomelasma Español, 238, 240 [N]. Type Oreomelasmaoromii Español, 1975, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Orghidania Ardoin, 383 [F]. Type Orghidaniatorrei Ardoin, 1977, by monotypy. Status: senior synonym of Spelaebiosis Bousquet & Bouchard, 2018 in Tenebrioninae: Triboliini. Note: junior homonym of Orghidania Capuse, 1971 [Lepidoptera].

Orientocara Koch, 176 [N]. Type Stenocaraarachnoides Gerstaecker, 1854, by original designation. Status: valid genus in Pimeliinae: Adesmiini.

Orientochile Penrith & Endrödy-Younga, 82 [F]. Type Cryptochileelegans Gerstaecker, 1854, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Orobychus Pascoe, xii [M]. Type Orobychuslacordairii Pascoe, 1868 (= Taphrosomadohrnii Kirsch, 1866), by monotypy. Status: junior synonym of Taphrosoma Kirsch, 1866 in Stenochiinae: Cnodalonini. Synonymy: Champion (1885: 108).

Orocina Reitter, 303 [F]. Type Orocinacapnisiceps Reitter, 1897, by subsequent designation (R. Lucas 1920: 463). Status: junior synonym of Syachis Bates, 1879 in Pimeliinae: Tentyriini. Synonymy: Bogdanov-Katjkov (1916: 68).

Orophylaxus Koch, 417 [M]. Type Phylaxincertus Mulsant & Godart, 1866, by original designation. Status: valid subgenus of Otinia Antoine, 1942 in Blaptinae: Dendarini: Melambiina.

Oroptera Borchmann, 48, 104 [F]. Type Oropteraphysoptera Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Orostegastopsis Koch, 255 [F]. Type Stegastopsisscorteccii Koch, 1962, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Orphelops Gozis, 102 [M]. Type Helopsimpressicollis Faldermann, 1837 (= Helopsfaldermanni Faldermann, 1837), by original designation. Status: junior synonym of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: new synonym [YB]. Note: this genus-group taxon has been forgotten in the literature; its type species is currently placed in the nominotypical subgenus of Nalassus Mulsant, 1854 and for that reason Gozis’s name is regarded as a new junior synonym of Nalassus.

Ortheolus Casey, 380 [M]. Type Schoenicusoculatus Champion, 1884, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Orthogonoderes Solier, 233 [M]. Type Praocissubreticulatus Solier, 1841, by subsequent designation (Flores and Pizarro-Araya 2014: 68). Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini.

Orthonychius Gebien, 83 [M]. Type Orthonychiusdigitatus Gebien, 1926, by monotypy. Status: junior synonym of Trilobocara Solier, 1851 in Pimeliinae: Trilobocarini. Synonymy: Doyen (1994: 501).

Orthostibia Blaisdell, 235 [F]. Type Orthostibiafrontalis Blaisdell, 1923, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Osdara Walker, 284 [F]. Type Osdarapicipes Walker, 1858, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Osdaroides Kaszab, 324 [M]. Type Osdaroidesmetallicus Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: Osdaroides was used earlier by Kaszab (1979a: 108) without a description, a definition or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Ospidus Pascoe, 467 [M]. Type Ospiduschrysomeloides Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Ossiporis Pascoe, 451 [F]. Type Ossiporisterrena Pascoe, 1866, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Osternus Fairmaire, 446 [M]. Type Osternusopacicollis Fairmaire, 1895, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Ostorius Fairmaire, xxxiii [M]. Type Ostoriusmulticostatus Fairmaire, 1889, by monotypy. Status: junior synonym of Adelonia Laporte, 1840 in Blaptinae: Pedinini: Leichenina. Synonymy: Kamiński et al. (2019a: 92).

Oterophloeus Desbrochers des Loges, 52 [M]. Type Oterophloeuspicipes Desbrochers des Loges, 1881 (= Pachychilahumerosus Fairmaire, 1875), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Oteroscelis Solier, 546 [F]. Type Adesmiapulcherrima Solier, 1835 (= Adesmiaaudouini Solier, 1835), by subsequent designation (Hope 1841: 118). Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Oteroscelopsis Löbl & Merkl in Löbl et al., 2020: 1 [F]. Type Pimeliadilatata Klug, 1830, by original designation. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: name first proposed by Koch (1944b: 147) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 244) designated Pimeliadilatata Klug, 1830 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Othelecta Pascoe, 488 [F]. Type Othelectatorrida Pascoe, 1866, by monotypy. Status: junior synonym of Cylindrothorus Solier, 1843 in Alleculinae: Alleculini: Mycetocharina. Synonymy: Haag-Rutenberg (1879c: 412).

Othryades Champion, 72 [M]. Type Othryadesfragilicornis Champion, 1889, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Othryoneus Champion, 245 [M]. Type Othryoneuserotyloides Champion, 1886, by subsequent designation (Gebien 1942a: 315). Status: valid genus in Stenochiinae: Cnodalonini.

Otinia Antoine, 22, 44 [F]. Type Otiniaiblanensis Antoine, 1942, by monotypy. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Otoceromorphus Pic, 11 [M]. Type Otoceromorphusgounellei Pic, 1915, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Otocerus Mäklin, 484 [M]. Type species [automatic]: Strongyliumserraticorne Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Oploptera Chevrolat, 1844 in Stenochiinae: Stenochiini. Note: unnecessary replacement name for Oploptera Chevrolat, 1844 (as “Hoploptera”).

Otrintus Pascoe, 483 [M]. Type Prosodesbehrii Germar, 1848, by original designation. Status: junior synonym of Cardiothorax Motschulsky, 1860 in Lagriinae: Adeliini. Synonymy: Bates (1879c: 30).

Otys Champion, 221 [M]. Type Otysharpalinus Champion, 1895 (= Scaletomerusproximus Blackburn, 1891), by subsequent designation (R. Lucas 1920: 468). Status: junior synonym of Scaletomerus Blackburn, 1891 in Alleculinae: Alleculini: Alleculina. Synonymy: Carter (1915a: 78). Note: Matthews and Bouchard (2008: 333) selected Otysarmatus Champion, 1895 as the type species of this genus (and treated Otys Champion, 1895 as valid) not knowing of the earlier valid designation by R. Lucas (1920: 468).

Oubanghinum Pic, 4 [N]. Type Oubanghinumatrum Pic, 1933, by monotypy. Status: junior synonym of Heterotarsus Latreille, 1829 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Ardoin (1969b: 126).

Ovalobioramix Egorov, 603 [F]. Type Platyscelismolesta Bogatchev, 1947, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Ovaloodescelis Kaszab, 940, 947 [F]. Type Platyscelisaffinis Seidlitz, 1893, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Overlaetia Pic, 304 [F]. Type Overlaetiagracilitarsis Pic, 1937, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: as pointed out by Bremer and Lillig (2014: 118) the older name Overlaetia Schouteden, 1932 [Hemiptera] is not nomenclaturally available and therefore Overlaetia Pic, 1937 is not a junior homonym.

Oxidates Champion, 263 [M]. Type Oxidatesplanicollis Champion, 1886, by subsequent designation (Gebien 1943: 402). Status: valid genus in Stenochiinae: Cnodalonini.

Oxinthas Champion, 72 [M]. Type Oxinthaspraocioides Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Branchini.

Oxura W. Kirby, 413 [F]. Type Oxurasetosa W. Kirby, 1819, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Oxycara Solier, 253, 254 [N]. Type Oxycarablapsoides Solier, 1835, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Oxycarops Reitter, 94 [M]. Type Hegeterfuscipes Brullé, 1839, by subsequent designation (Gebien 1937a: 640). Status: valid genus in Pimeliinae: Tentyriini.

Oxycerus Koch, 46 [M]. Type Trachynotusresolutus Péringuey, 1904, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Oxyge Chatanay, 7 [F]. Type Oxygerugosa Chatanay, 1914, by original designation. Status: valid genus in Pimeliinae: Asidini.

Oxygonodera Casey, 433, 444 [F]. Type Oxygonoderavillosa Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Oxypistoma Löbl, Bouchard, Merkl & Bousquet, 5 [N]. Type Prochomabucculentum Koch, 1940, by original designation. Status: valid subgenus of Prochoma Solier, 1835 in Pimeliinae: Tentyriini. Note: name first proposed with two alternative spellings, as Oxypistoma (Koch 1940a: 258, 259, 260, 262) and Oxipistoma (Koch 1940a: 259), without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl et al. (2008a: 40) designated Prochomabucculenta Koch, 1940 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Oxythorax Fåhraeus, 288 [M]. Type Oxythoraxclathratus Fåhraeus, 1870, by monotypy. Status: junior synonym of Anchophthalmus Gerstaecker, 1854 in Blaptinae: Platynotini: Platynotina. Synonymy: Péringuey (1904: 238).

Oxyura Agassiz, 267, 268 [F]. Type species [automatic]: Oxurasetosa W. Kirby, 1819, by monotypy. Status: junior synonym of Oxura W. Kirby, 1819 in Pimeliinae: Sepidiini: Oxurina. Note: unjustified emendation of Oxura W. Kirby, 1819, not in prevailing usage; junior homonym of Oxyura Bonaparte, 1831 [Aves].

Oyanus Pic, 23 [M]. Type Oyanuscurticornis Pic, 1921, by monotypy. Status: junior synonym of Cleomis Fairmaire, 1892 in Stenochiinae: Cnodalonini. Synonymy: Löbl et al. (2008b: 340).

Ozaenimorphus Fairmaire, 127 [M]. Type Ozaenimorphuscostulipennis Fairmaire, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ozolais Pascoe, 457 [F]. Type Ozolaisscruposa Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Ozotypoides Kaszab, 222 [M]. Type Ozotypoidesgranulatus Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Adeliini.

Ozotypus Pascoe, 328 [M]. Type Ozotypussetosus Pascoe, 1862, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pachacamacius Flores & Giraldo-Mendoza in Giraldo-Mendoza and Flores, 84 [M]. Type Pachacamaciusaguilari Giraldo-Mendoza & Flores, 2019, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Pachycera Eschscholtz, 5, 7 [F]. Type fixed herein (ICZN 1999, Article 70.3) as Tenebriobuprestoides Fabricius, 1781, misidentified as Akislaevigata Fabricius, 1801 in the original designation by monotypy in Eschscholtz (1831). Status: junior synonym of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Löbl et al. (2008b: 192). Note: junior homonym of Pachycera Billberg, 1820 [Hemiptera]; the type species “Akislaevigata Fabricius” was first established by monotypy; Koch (1943: 530) noted that Akislaevigata Fabricius of Eschscholtz (1831) was misidentified and corresponded to the species Tenebriobuprestoides Fabricius, 1781; we follow currently accepted concepts (e.g., Löbl et al. 2008b: 192) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Akislaevigata Fabricius, 1801 also belongs to the genus Hyperops Eschscholtz, 1831; the original combination of the accepted name of the type species, Tenebriobuprestoides Fabricius, 1781, is a junior primary homonym of Tenebriobuprestoides Scopoli, 1763 and Tenebriobuprestoides Pallas, 1773.

Pachycerops Koch, 524, 533 [M]. Type Pachycerainsidiosa Fairmaire, 1896, by monotypy. Status: valid subgenus of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Pachycerus Montrouzier, 291 [M]. Type Pachycerusdomesticus Montrouzier, 1860, by monotypy. Status: senior synonym of Sciophagus Sharp, 1885 in Diaperinae: Diaperini: Diaperina. Synonymy: Fauvel (1904: 185). Note: junior homonym of Pachycerus Schönherr, 1823 [Coleoptera: Curculionidae].

Pachychila Eschscholtz, 5 [F]. Type Pimeliapunctata Fabricius, 1798, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: the original combination of the name of the type species, Pimeliapunctata Fabricius, 1798, is a junior primary homonym of Pimeliapunctata Thunberg, 1787.

Pachychile Lacordaire, 46 [F]. Type species [automatic]: Pimeliapunctata Fabricius, 1798, by monotypy. Status: junior synonym of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: unjustified emendation of Pachychila Eschscholtz, 1831, not in prevailing usage. Note: the original combination of the name of the type species, Pimeliapunctata Fabricius, 1798, is a junior primary homonym of Pimeliapunctata Thunberg, 1787.

Pachychilina Reitter, 91, 145 [F]. Type Tentyriadejeani Besser, 1832, by subsequent designation (R. Lucas 1920: 473). Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Pachycoelia Boisduval, 248 [F]. Type Pachycoeliasulcicollis Boisduval, 1835, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: name incorrectly listed as preoccupied in the literature (e.g., Matthews and Bouchard 2008: 296) but used as valid recently (Matthews and Lawrence 2019: 632).

Pachycossyphodes Andreae, 203, 214 [M]. Type Cossyphodesmachadoi Basilewsky, 1952, by original designation. Status: junior synonym of Cossyphodes Westwood, 1851 in Pimeliinae: Cossyphodini: Cossyphodina. Synonymy: Schawaller (2013c: 362, implied by inclusion of Cossyphodesmachadoi Basilewsky, 1952 in Cossyphodes Westwood, 1851 without use of a subgenus rank).

Pachycyrtosoma Marcuzzi, 81 [N]. Type Cyrtosomamerkli Marcuzzi, 1999, by original designation. Status: junior synonym of Nesocyrtosoma Marcuzzi, 1976 in Stenochiinae: Cnodalonini. Synonymy: Hopp and Ivie (2009: 13).

Pachylagria Borchmann, 17 [F]. Type Pachylagriaovata Borchmann, 1912, by subsequent designation (Merkl 2008: 115). Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1916a: 163, with Heterogria Fairmaire, 1896, a junior synonym of Xanthalia Fairmaire, 1894).

Pachylesthus Fairmaire, 219 [M]. Type Pachylesthusvalidus Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pachylocerus Hope, 186 [M]. Type Pachyloceruswestermanni Hope, 1841, by monotypy. Status: senior synonym of Pycnocerus Westwood, 1841 in Lagriinae: Pycnocerini. Note: junior homonym of Pachylocerus Hope, 1834 [Coleoptera: Cerambycidae].

Pachylodera Quedenfeldt, 399 [F]. Type Pachyloderabrevicornis Quedenfeldt, 1890, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Pachymastus Fairmaire, 350 [M]. Type Pachymastusasperulus Fairmaire, 1896, by monotypy. Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Pachynotelus Solier, 210, 267 [367] [M]. Type Pachynotelusalbiventris Solier, 1841, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Pachyphaleria Gebien, 136 [F]. Type Phaleriacapensis Laporte, 1840, by monotypy. Status: valid genus in Diaperinae: Phaleriini.

Pachypterocoma Skopin, 157 [F]. Type Pterocomapallasi Semenov-Tjan-Shansky, 1910, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Pachypterus P.H. Lucas, pl. 29 [M]. Type Pachypterusmauritanicus P.H. Lucas, 1847, by monotypy. Status: senior synonym of Neopachypterus Bouchard, Löbl & Merkl, 2007 in Blaptinae: Opatrini: Neopachypterina. Note: junior homonym of Pachypterus Swainson, 1839 [Pisces].

Pachyscelina Kwieton, 29, 30 [F]. Type Pachyscelismicros Kaszab, 1970, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Pachyscelis Solier, 9, 54 [F]. Type species [automatic]: Pimeliamusiva Ménétriés, 1832, by subsequent designation (Bouchard et al. 2007: 388). Status: valid genus and subgenus in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Brachyscelis Dejean, 1834. Note: nomen protectum (see Bouchard et al. 2007: 388).

Pachyscelodes Sénac, 189 [M]. Type Pachyscelishenoni Sénac, 1887, by subsequent designation (Gebien 1937a: 823). Status: junior synonym of Scelace Marseul, 1887 in Pimeliinae: Pimeliini. Synonymy: Reitter (1915: 2).

Pachysternoplax Skopin, 110, 154 [F]. Type Trigonoscelisarmeniaca Faldermann, 1837, by original designation. Status: valid subgenus of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini.

Pachystira Chen, 308 [F]. Type Pachystiraimpressipennis Chen, 1997, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Pachyurgus J.L. LeConte, 230 [M]. Type Iphthinusaereus Melsheimer, 1846, by original designation. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: J.L. LeConte (1873: 335, with Encyalesthus Motschulsky, 1860, a junior synonym of Derosphaerus J. Thomson, 1858). Note: the type locality for Iphthinusaereus Melsheimer, 1846 was incorrectly given as “Pennsylvania” originally (see Bousquet et al. 2018: 384).

Pactostoma J.L. LeConte, 19 [N]. Type Asidaanastomosis Say, 1824, by original designation. Status: junior synonym of Stenosides Solier, 1836 in Pimeliinae: Asidini. Synonymy: J.L. LeConte (1862: 222, with Ologlyptus Lacordaire, 1858, a junior synonym of Stenosides Solier, 1836).

Paita Fauvel, 173 [F]. Type Paitasetosella Fauvel, 1904, by monotypy. Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Kaszab (1982b: 56).

Paivaea Wollaston, 449 [F]. Type Tentyriahispida Brullé, 1839, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Paivea Scudder, 213 [F]. Type species [automatic]: Tentyriahispida Brullé, 1839, by monotypy. Status: junior synonym of Paivaea Wollaston, 1864 in Pimeliinae: Tentyriini. Note: unjustified emendation of Paivaea Wollaston, 1864, not in prevailing usage.

†Palaeobasanus Nabozhenko & Kirejtshuk, 25 [M]. Type Palaeobasanusneli Nabozhenko & Kirejtshuk, 2020, by original designation. Status: valid genus in Diaperinae: Scaphidemini. Note: described from Middle-Upper Paleocene deposits (France).

†Palaeosclerum Nabozhenko & Kirejtshuk, 308 [N]. Type Palaeosclerumpohli Nabozhenko & Kirejtshuk, 2017, by original designation. Status: valid genus in Blaptinae: Opatrini: Sclerina. Note: described from Middle-Upper Paleocene deposits (France).

Palembomimus Matthews & Lawrence, 539 [M]. Type Platydemadeplanata Champion, 1894, by original designation. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Palembus Casey, 65 [M]. Type Palembusocularis Casey, 1891, by monotypy. Status: junior synonym of Ulomoides Blackburn, 1888 in Diaperinae: Diaperini: Diaperina. Synonymy: Doyen et al. (1990: 237).

Palorinus Blair, 135 [M]. Type Palorushumeralis Gebien, 1914, by original designation. Status: valid genus in Tenebrioninae: Palorini.

Paloropsis Masumoto & Grimm, 127 [F]. Type Paloropsisirei Masumota & Grimm, 2004, by original designation. Status: valid genus in Tenebrioninae: Palorini.

Palorus Mulsant, 250 [M]. Type Hypophlaeusdepressus Fabricius, 1790, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Palpafrina Koch, 329 [F]. Type Afrinuswatsoni Koch, 1950, by original designation. Status: valid subgenus of Afrinus Fairmaire, 1888 in Pimeliinae: Tentyriini.

Palpichara Borchmann, 355 [F]. Type Palpicharaserricornis Borchmann, 1932, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Palpicula Novák, 168 [F]. Type Alleculafiliola Borchmann, 1925, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Palpomodes Koch, 223 [M]. Type Psammodesphysopterus Gebien, 1920, by monotypy. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Oxurina.

Pandarinus Mulsant & Rey, 50, 103 [M]. Type Pandarinustenellus Mulsant & Rey, 1854, by subsequent designation (Chatzimanolis and Löbl 2003: 260). Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Pandarus Dejean, 191 [M]. Type species [automatic]: Type Helopstristis Rossi, 1790, by subsequent designation Blanchard (1844: pl. 48). Status: junior synonym of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina. Note: unjustified emendation of Dendarus Dejean, 1821, not in prevailing usage (see Bousquet and Bouchard 2013a: 48); junior homonym of Pandarus Leach, 1816 [Crustacea].

Paniasis Champion, 208 [M]. Type Paniasisdilatipes Champion, 1886, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Paoligena Pic, 42 [F]. Type Praogenainhumeralis Pic, 1928, by monotypy. Status: valid genus in Tenebrioninae: Paoligenini. Note: originally described as a subgenus of Praeugena Laporte, 1840, elevated to the rank of genus by De Moor (1970: 15).

Papuamisolampus Kaszab, 288 [M]. Type Papuamisolampustoxopeusi Kaszab, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Parabantodemus Iwan, 345 [M]. Type Trigonopusspinipes Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Parabigopsis Español, 111, 114 [F]. Type Parabigopsispeyerimhoffi Español, 1946, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Parablops Rottenberg, 254 [M]. Type Parablopsaetnensis Rottenberg, 1871, by monotypy. Status: senior synonym of Gerandryus Rottenberg, 1873 in Alleculinae: Alleculini: Gonoderina. Note: junior homonym of Parablops Schönherr, 1839 [Coleoptera: Anthribidae].

Parabolitophagus Miyatake, 67, 70 [M]. Type Bolitophagusfelix Lewis, 1894, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Paracirta Schuster, 66 [F]. Type Paracirtaschatzmayri Schuster, 1930, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Paracistela Borchmann, 30 [F]. Type Paracistelavariabilis Borchmann, 1941, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Paracossyphus Viñolas & Cartagena, 29, 55 [M]. Type species [automatic]: Cossyphusminutissimus Laporte, 1840, by original designation. Status: junior synonym of Acontodactylus Desbrochers des Loges, 1894 in Lagriinae: Cossyphini. Note: unnecessary replacement name for Acontodactylus Desbrochers des Loges, 1894 (as “Acanthodactylus”).

Paradissonomus G.S. Medvedev, 229 [M]. Type Dissonomuslongulus Bogatchev & Kryzhanovsky, 1960, by original designation. Status: valid subgenus of Dissonomus Jacquelin du Val, 1861 in Tenebrioninae: Dissonomini.

Paradrus Jakobson, 242 [M]. Type species [automatic]: Pseudhadrusseriatus Kolbe, 1910, by subsequent designation (Gebien 1941: 332). Status: junior synonym of Pseudhadrus Kolbe, 1910 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Pseudhadrus Kolbe, 1910.

Paragena Bouchard & Bousquet, new subgenus [F]. Type Nesogenaviridicuprea Fairmaire, 1868, by present designation. Status: valid subgenus of Nesogena Mäklin, 1863 in Tenebrioninae: Praeugenini. Note: first proposed by Froussart (1961: 60) without type species designation; the subgenusParagena, which is currently used as valid, is therefore unavailable (ICZN 1999, Articles 13.3, 16.1); we hereby make the name available by selecting Nesogenaviridicuprea Fairmaire, 1868 as type species and referring to Froussart (1961: 60) for the character states that characterise and differentiate Paragena.

Paragonocnemis Kraatz, 118 [F]. Type Paragonocnemissculpticollis Kraatz, 1899 (= Gonocnemisfoveicollis Fairmaire, 1891), by subsequent designation (Gebien 1943: 918). Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Paraguania Marcuzzi, 31 [F]. Type Paraguaniarelicta Marcuzzi, 1953, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Parahymenorus Campbell, 100 [M]. Type Parahymenorusmetallicus Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Parahyocis Kaszab, 650 [M]. Type Hyocischampioni Fauvel, 1904, by original designation. Status: valid genus in Diaperinae: Hyociini: Hyociina.

Paraisomira Dubrovina, 138 [F]. Type Cistelaoculata Marseul, 1876, by original designation. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

†Parakeleusticus Haupt, 114, 126 [M]. Type Parakeleusticuspostumus Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Paraleptodes G.S. Medvedev, 354 [M]. Type Leptodeslindbergi Kaszab, 1959, by original designation. Status: valid subgenus of Leptodes Dejean, 1834 in Pimeliinae: Leptodini.

Paralitoborus Antoine, 190 [M]. Type Litoborussternalis Fairmaire, 1884, by original designation. Status: valid subgenus of Litoborus Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Paralorelopsis Marcuzzi, 117 [F]. Type Paralorelopsisbordoni Marcuzzi, 1994, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Paralyreus Grouvelle, 24 [M]. Type Paralyreusscotti Grouvelle, 1918, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Paramarygmus Quedenfeldt, 25 [M]. Type Paramarygmusnigroaeneus Quedenfeldt, 1885 (= Hybonotusfemoralis Imhoff, 1843), by monotypy. Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Paramellon C.O. Waterhouse, iv [N]. Type Paramellonsociale C.O. Waterhouse, 1882, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: Paramellonina.

Paramellops Andreae, 201 [M]. Type Cossyphodesbewicki Wollaston, 1861, by original designation. Status: valid genus in Pimeliinae: Cossyphodini: Cossyphodina.

Paramisolampidius Merkl & Masumoto in Löbl et al., 2020: 4 [M]. Type Paramisolampidiuskagoshimensis Nakane, 1968, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: name first proposed by Nakane (1968: 82) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Merkl and Masumoto (2007: 1) designated Paramisolampidiuskagoshimensis Nakane, 1968 as the type species of Nakane’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Paranemia Heyden, 103 [F]. Type Paranemiaschroederi Heyden, 1892, by monotypy. Status: valid genus in Diaperinae: Phaleriini.

Paranephodes Antoine, 208 [M]. Type Nephodescamusi Antoine, 1955, by monotypy. Status: valid subgenus of Nephodinus Gebien, 1943 in Tenebrioninae: Helopini: Helopina.

Paranopidium Dajoz, 429, 434 [N]. Type Paranopidiumafricanum Dajoz, 1974, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Parapachynotela Koch, 54 [F]. Type Parapachynotelabushmanica Koch, 1952, by original designation. Status: junior synonym of Horatoma Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Synonymy: Penrith and Endrödy-Younga (1994: 10).

Parapachyscelis Kwieton, 29 [F]. Type Pimeliavillosa Drapiez, 1820, by original designation. Status: valid subgenus of Pachyscelis Solier, 1836 in Pimeliinae: Pimeliini.

Paraphanes W.J. MacLeay, 308 [M]. Type Paraphanesnitidus W.J. MacLeay, 1887, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Paraphloeus Seidlitz, 553, 557 [M]. Type Hypophlaeuslongulus Gyllenhal, 1827, by subsequent designation (G.S. Medvedev 1990: 231). Status: junior synonym of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini. Synonymy: Bremer (1995: 3).

Paraplatyope Löbl, Bouchard, Merkl & Bousquet, 4 [F]. Type Leucolaephusarabicus Blair, 1931, by original designation. Status: valid genus in Pimeliinae: Pimeliini. Note: name first proposed by Gridelli (1953: 45) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Paraplatyscelis Kaszab, 912, 936 [F]. Type Platyscelissinuata Seidlitz, 1893, by original designation. Status: valid subgenus of Platyscelis Latreille, 1818 in Blaptinae: Platyscelidini.

Parapraocis Flores & Giraldo, 35 [M]. Type Praocisvagecostatus Fairmaire, 1902, by original designation. Status: valid genus in Pimeliinae: Praociini. Note: name originally proposed by Kulzer (1958a: 13, 58) as subgenus of Praocis Eschscholtz, 1829 without type species designation; Flores and Pizarro-Araya (2012: 4) designated Praocisvagecostatus Fairmaire, 1902 as type species, but did not explicitely propose the taxon as new (ICZN 1999, Article 16.1).

Paraprosodes Reitter, 119 [M]. Type Prosodestriangulipes Reitter, 1909, by original designation. Status: junior synonym of Prosodella Reitter, 1909 in Blaptinae: Blaptini: Prosodina. Synonymy: G.S. Medvedev (1997: 597).

Parapterocoma Skopin, 149 [F]. Type Pterocomavittata Frivaldszky, 1890, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Paraselinus Kamiński, 705, 711 [M]. Type Paraselinusiwani Kamiński, 2013, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Parasida Casey, 76, 126 [F]. Type Parasidalaciniata Casey, 1912, by original designation. Status: senior synonym of Pleisiasida Smith, 2013 in Pimeliinae: Asidini. Note: junior homonym of Parasida Daday, 1904 [Crustacea].

Parasternoplax Skopin, 110, 148 [F]. Type Pimeliadeplanata Krynicki, 1832, by original designation. Status: valid subgenus of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini.

Parastizopus Gebien, 93 [M]. Type Parastizopusdiehli Gebien, 1938, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Parastrongylium Kaszab, 10, 24 [N]. Type Strongyliumlorentzi Gebien, 1921, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Paratenetus Spinola, 116 [M]. Type Paratenetuspunctatus Spinola, 1845, by subsequent designation (R. Lucas 1920: 483). Status: valid genus in Lagriinae: Goniaderini.

Paratoxicum Champion, 380 [N]. Type Paratoxicumiridescens Champion, 1894, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Paravius Casey, 332 [M]. Type Emmenastusmarginatus Casey, 1890, by monotypy. Status: valid subgenus of Hylocrinus Casey, 1907 in Pimeliinae: Edrotini.

Parecatus Fairmaire, 245 [M]. Type Parecatusplicatulus Fairmaire, 1900, by subsequent designation (Gebien 1937a: 741). Status: junior synonym of Scotinesthes Fairmaire, 1895 in Pimeliinae: Asidini. Synonymy: Koch (1962a: 67).

Parepitragus Casey, 518 [M]. Type Parepitragussolieri Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Pareupezus Kolbe, 129 [M]. Type Pareupezusglaber Kolbe, 1889, by monotypy. Status: junior synonym of Paramarygmus Quedenfeldt, 1885 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1921b: 148).

Parimmedia Gebien, 220, 226 [F]. Type Parimmediaepipleuralis Gebien, 1928, by subsequent designation (Gebien 1943: 404). Status: valid genus in Stenochiinae: Cnodalonini.

Parmularia Koch, 35 [F]. Type Psammodescaffer Fåhraeus, 1870, by monotypy. Status: junior synonym of Psammodes W. Kirby, 1819 in Pimeliinae: Sepidiini: Molurina. Synonymy: Kamiński et al. (2019b: 31). Note: junior homonym of Parmularia MacGillivray, 1887 [Bryozoa].

Paroderus Mulsant & Rey, 111 [M]. Type Pandarinuselongatus Mulsant & Rey, 1854, by subsequent designation (Chatzimanolis and Löbl 2003: 260). Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Paroeatus Gebien, 169, 178 [M]. Type Paroeatusopacus Gebien, 1928, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Parogria Borchmann, 119 [F]. Type Lagriaatrata Borchmann, 1912, by original designation. Status: valid subgenus of Cerogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina.

Paromophlus Iablokoff-Khnzorian, 134, 139 [M]. Type Cistelapicipes Fabricius, 1792, by original designation. Status: junior synonym of Phibalus Gistel, 1856 in Alleculinae: Cteniopodini. Synonymy: Bousquet et al. (2015: 142). Note: the name Paromophlus was first introduced by Znojko in Ogloblin and Znojko (1950: 125) without designation of a type species and is therefore unavailable from that date.

†Paropiophorus Haupt, 114, 132 [M]. Type Paropiophorusnitidus Haupt, 1950, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Middle Eocene deposits (Germany); this genus was previously considered to be “definitely not a tenebrionid” by Watt (1975: 389) but is included here following Nabozhenko (2019: 8).

Partystona Watt, 47 [F]. Type Partystonametallica Watt, 1992, by original designation. Status: valid genus in Tenebrioninae: Titaenini.

Passalocharis Koch, 13 [F]. Type Chirocharisintermedius Gebien, 1911, by original designation. Status: valid genus in Lagriinae: Pycnocerini.

Patagonogenius Flores, 371, 390 [M]. Type Mitrageniusquadricollis Fairmaire, 1876, by original designation. Status: valid genus in Pimeliinae: Nycteliini.

Patagonopraocis Flores & Chani-Posse, 576 [M]. Type Patagonopraocismagellanicus Flores & Chani-Posse, 2005, by original designation. Status: valid genus in Pimeliinae: Praociini.

Paulianaria Bouchard & Bousquet, new genus [F]. Type Paulianariastrongylioides Ardoin, 1961, by present designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: Ardoin (1961d: 97) introduced the new genus name Paulianaria for five nominal species, but unfortunately did not designate a type species; the genus Paulianaria, which has been treated as valid since 1961, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Paulianariastrongylioides Ardoin, 1961 as type species and referring to Ardoin (1971: 362) for the character states that characterise and differentiate Paulianaria.

Paulianesthes Koch, 39, 57 [F]. Type Paulianesthesamplipennis Koch, 1962, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Paurodontomophlus Muche, 171 [M]. Type Omophluswittmeri Muche, 1979, by original designation. Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini.

Pechalius Casey, 379, 420 [M]. Type Pechaliussubvittatus Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Epitragini. Note: the First Reviser (Pechalius Casey, 1907 versus Epitragoma Casey, 1907) is Freude (1968: 61).

Pectinepitragus Pic, 44 [M]. Type Pectinepitraguspubescens Pic, 1927, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Pectphegoneus Freude, 90, 98 [M]. Type Schoenicuspectoralis Champion, 1884, by monotypy. Status: valid subgenus of Phegoneus Casey, 1907 in Pimeliinae: Epitragini.

Pedarasida Reitter, 11, 28 [F]. Type Asidacariosicollis Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Pedinopsis Gebien, 157 [F]. Type Pedinopsispilipes Gebien, 1910, by monotypy. Status: senior synonym of Loensus R. Lucas, 1920 in Blaptinae: Pedinini: Pedinina. Note: junior homonym of Pedinopsis Cotteau, 1863 [Echinoidea].

Pedinulus Seidlitz, 364, 373 [M]. Type Pedinusragusae Baudi di Selve, 1875, by subsequent designation (Iwan and Löbl 2008: 288). Status: junior synonym of Pedinus Latreille, 1797 in Blaptinae: Pedinini: Pedinina. Synonymy: Kamiński and Iwan (2017: 599).

Pedinus Latreille, 20 [M]. Type Tenebriofemoralis Linnaeus, 1767, by subsequent designation (Stephens 1835: 159). Status: valid genus and subgenus in Blaptinae: Pedinini: Pedinina. Note: originally proposed without included nominal species; Latreille (1802: 175), by listing Tenebriofemoralis Linnaeus, 1767 and Tenebriodermestoides Fabricius, 1787 in association with this genus name, was the first author to subsequently and expressly include nominal species in Pedinus (ICZN 1999, Article 67.2.2).

Pedionomus Haag-Rutenberg, 42 [M]. Type Metriopusfavosus Erichson, 1843, by subsequent designation (Gebien 1937a: 661). Status: senior synonym of Alogenius Gebien, 1910 in Pimeliinae: Adesmiini. Note: junior homonym of Pedionomus Gould, 1841 [Aves].

Pediris Motschulsky, 24 [F]. Type Pedirislongipes Motschulsky, 1872, by original designation. Status: junior synonym of Promethis Pascoe, 1869 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1921a: 289, with Setenis Motschulsky, 1872, a junior synonym of Promethis Pascoe, 1869).

Pedoeces Agassiz, 276 [M]. Type species [automatic]: Pedonoecesgalapagoensis G.R. Waterhouse, 1845, by subsequent designation (Aalbu and Triplehorn 1991: 170). Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Note: unjustified emendation of Pedonoeces G.R. Waterhouse, 1845, not in prevailing usage.

Pedonoeces G.R. Waterhouse, 32, 34 [M]. Type Pedonoecesgalapagoensis G.R. Waterhouse, 1845, by subsequent designation (Aalbu and Triplehorn 1991: 170). Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Aalbu and Triplehorn (1991: 170).

Pedostrongylium Pic, 11 [N]. Type Strongyliumluteonotatum Pic, 1916 (= Stenochiaxanthozona J. Thomson, 1858), by original designation. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Pelecypalpus Hinton, 91 [M]. Type Pelecypalpusmedon Hinton, 1947, by original designation. Status: junior synonym of Scotoderus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1973c: 258).

Pelecyphorus Solier, 406, 467 [M]. Type Pelecyphorusmexicanus Solier, 1836, by subsequent designation (Hope 1841: 110). Status: valid genus and subgenus in Pimeliinae: Asidini.

Pelleas Bates, 98 [M]. Type Tenebriocrotchii Wollaston, 1865, by original designation. Status: valid genus in Diaperinae: Diaperini: incertae sedis.

Pelops Gistel, 22 [M]. Type Helopsater Fabricius, 1775, by monotypy. Status: senior synonym of Prionychus Solier, 1835 in Alleculinae: Alleculini: Alleculina. Note: nomen oblitum (see Bousquet and Bouchard 2017: 132).

Pelorinus Vauloger de Beaupré, 675, 678 [M]. Type Helopsanthracinus Germar, 1813, by subsequent designation (Cherney 382). Status: valid subgenus of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina.

Pelorocnemis Solsky, 283 [F]. Type Pimeliapunctigera Ménétriés, 1849, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Peltadesmia Kuntzen, 149 [F]. Type Metriopusplatynotus Gerstaecker, 1854, by present designation. Status: junior synonym of Coeladesmia Reitter, 1916 in Pimeliinae: Adesmiini. Synonymy: Gebien (1937a: 656).

Peltarium Fischer von Waldheim, 106 [N]. Type Peltariummarginatum Fischer von Waldheim, 1844 (= Blaps baerii Fischer von Waldheim, 1842), by subsequent designation (G.S. Medvedev 1995b: 831). Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Peltasida Reitter, 39, 41 [F]. Type Asidafavieri Fairmaire, 1880, by monotypy. Status: valid subgenus of Asida Latreille, 1802 in Pimeliinae: Asidini.

Peltoides Laporte, 401 [M]. Type Peltoidessenegalensis Laporte, 1833, by subsequent designation (Lacordaire 1859a: 337). Status: valid genus and subgenus in Tenebrioninae: Alphitobiini. Note: bibliographic evidence indicates that Oopiestus Chevrolat, 1833 was published before Peltoides Laporte, 1833 and should be treated as the valid name for this genus and subgenus instead; an application to the ICZN is necessary to conserve usage of Peltoides Laporte, 1833.

Peltolobus Lacordaire, 71 [M]. Type species [automatic]: Megalophryspatagonica G.R. Waterhouse, 1845, by monotypy. Status: valid genus in Pimeliinae: Trilobocarini. Note: replacement name for Megalophrys G.R. Waterhouse, 1845.

Pemanoa Buck, 269 [F]. Type Pemanoamixta Buck, 1955, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Penadelium Matthews, 710, 754 [N]. Type Penadeliumaraucanum Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Penaus Freude, 130 [M]. Type Penauspenai Freude, 1968, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Peneta Lacordaire, 319 [F]. Type Penetalebasii Lacordaire, 1859, by subsequent designation (R. Lucas 1920: 492). Status: valid genus in Phrenapatinae: Penetini.

Pengalenganus Pic, 10 [M]. Type Pengalenganusinaequalis Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Penichrus Champion, 134 [M]. Type Penichrusblapstinoides Champion, 1885, by monotypy. Status: valid genus in Tenebrioninae: incertae sedis. Note: placed in Tenebrioninae incertae sedis by Johnston et al. (2020: 771).

Pentaphyllus Dejean, 68 [M]. Type Mycetophagustestaceus Hellwig, 1792, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Penthicinus Reitter, 161 [M]. Type Penthicinuskoltzei Reitter, 1896, by subsequent designation (Gebien 1939: 458). Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: Reitter (1896b) used two different spellings for this genus, including Penticinus (Reitter 1896b: 171); subsequently, Reitter (1904: 135, 170) used Penthicinus as the valid name of the genus and so acted as First Reviser (ICZN 1999: Article 24.2.4).

Penthicoides Fairmaire, 20 [M]. Type Penthicoidesseriatoporus Fairmaire, 1896, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Penthicomelanesthes Bogatchev, 631 [F]. Type Heliophilusgibbulus Faldermann, 1835, by original designation. Status: junior synonym of Myladion Reitter, 1887 in Blaptinae: Opatrini: Opatrina. Synonymy: G.S. Medvedev (1990: 186).

Penthicus Faldermann, 384 [M]. Type Penthicuspinguis Faldermann, 1836, by subsequent designation (Hope 1841: 126). Status: valid genus and subgenus in Blaptinae: Opatrini: Opatrina.

Pentholasius Reitter, 178 [M]. Type Halonomusvariolatus Allard, 1884, by monotypy. Status: junior synonym of Mesomorphus Miedel, 1880 in Blaptinae: Opatrini: Opatrina. Synonymy: Ferrer (2002: 376).

Penthomegus Reitter, 161 [M]. Type Penthicuscorpulentus Reitter, 1896, by subsequent designation (Iwan and Löbl 2007: 734). Status: junior synonym of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Synonymy: Gebien (1939: 459, with Lobodera Mulsant & Rey, 1859, a junior synonym of Penthicus Faldermann, 1836).

Perdicus Fairmaire, 386 [M]. Type Perdicusantrophilus Fairmaire, 1899, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Perdistretus Koch, 65 [M]. Type Distretusvilhenai Koch, 1953, by original designation. Status: valid subgenus of Distretus Haag-Rutenberg, 1871 in Pimeliinae: Sepidiini: Molurina.

Periatrum Sharp, 407 [N]. Type Periatrumhelmsi Sharp, 1886, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Periblaps Bauer, 32 [F]. Type none designated. Status: undetermined taxon in Blaptinae: Blaptini: Blaptina. Note: this genus was described before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Periblaps and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Perichilus Quedenfeldt, 18 [M]. Type Perichilusbrevicornis Quedenfeldt, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Periloma Gebien, 100 [N]. Type Perilomaalfkeni Gebien, 1938, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Peringueyia Koch, 44 [F]. Type Echinotusdispar Péringuey, 1899, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Periphanes Fairmaire, 245 [M]. Type Periphanesorichalceus Fairmaire, 1882, by monotypy. Status: senior synonym of Periphanodes Gebien, 1943 in Stenochiinae: Cnodalonini. Note: junior homonym of Periphanes Hübner, 1821 [Lepidoptera].

Periphanodes Gebien, 902 [M]. Type species [automatic]: Periphanesorichalceus Fairmaire, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Periphanes Fairmaire, 1882; new placement [OM], previously included in Tenebrioninae: Helopini.

Peristeptus Haag-Rutenberg, 3, 24 [M]. Type Pogonobasislaevigata Gerstaecker, 1854, by subsequent designation (R. Lucas 1920: 495). Status: junior synonym of Pogonobasis Solier, 1837 in Pimeliinae: Adelostomini. Synonymy: Koch (1952b: 17).

Perithrix Fairmaire, 193 [F]. Type Perithrixgranidorsis Fairmaire, 1879, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Pescennius Champion, 3 [M]. Type Pescenniusvillosus Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Petria Semenov, 611 [F]. Type Petriatachyptera Semenov, 1894, by subsequent designation (R. Lucas 1920: 496). Status: valid genus in Alleculinae: Cteniopodini.

Petrostetha Novák, 212 [F]. Type Petrostethatibialis Novák, 2008, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Peyerimhoffius Koch, 420 [M]. Type Phylaxplicatus P.H. Lucas, 1846, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Peyrierasia Dajoz, 113, 114 [F]. Type Peyrierasiasechellensis Dajoz, 1975, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Pezodontus Dejean, 203 [M]. Type species [automatic]: Odontopuscostatus Silbermann, 1833, by subsequent designation (Hope 1841: 126). Status: valid genus in Lagriinae: Pycnocerini. Note: replacement name for Odontopus Silbermann, 1833 (see Bousquet and Bouchard 2013a: 57).

Pezohelaeus Gebien, 282 [M]. Type Pterohelaeushirtus W.J. MacLeay, 1888, by monotypy. Status: junior synonym of Pterohelaeus Brême, 1842 in Tenebrioninae: Heleini: Heleina. Synonymy: Carter (1927: 230).

Pezomaia Kulzer, 752 [F]. Type Pezomaiafemoralis Kulzer, 1952, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pezophenus Gebien, 325, 339 [M]. Type Pezophenusrutilans Gebien, 1921 (= Prophanessubmetallicus W.J. MacLeay, 1887), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Phaedeucyrtus Pic, 14 [M]. Type Phaedeucyrtusobscuripes Pic, 1916, by monotypy. Status: junior synonym of Phaedis Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Ando (2016: 70).

Phaedis Pascoe, 474 [M]. Type Phaediselysius Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Phaedogria Borchmann, 16, 57 [F]. Type Lagriaionoptera Erichson, 1834, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Phaenogeton Bremer, 220 [M]. Type Amarygmusvaricolor Gebien, 1921, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Phaeostolus Fairmaire, cxlvi [M]. Type Phaeostolusgrandicornis Fairmaire, 1884, by monotypy. Status: valid genus in Tenebrioninae: Praeugenini. Note: redescribed as new by Fairmaire (1887b: 293).

Phaeotribon Kraatz, 81, 242 [M]. Type Phaeotribonpulchellus Kraatz, 1865, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Phaleria Latreille, 162 [F]. Type Tenebriocadaverinus Fabricius, 1792, by plenary powers (ICZN 1975, Opinion 1039). Status: valid genus and subgenus in Diaperinae: Phaleriini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1975, Opinion 1039).

Phaleriderma Koch, 25 [N]. Type Oncotusobscuricollis Solier, 1848, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Phalerisida Kulzer, 563 [F]. Type Phalerisidamaculata Kulzer, 1959, by original designation. Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Synonymy: Triplehorn (1991: 258).

Phaleromela Reitter, 4 [F]. Type Phaleriasubhumeralis Marseul, 1876, by monotypy. Status: valid genus in Diaperinae: Phaleriini.

Phallocentrion Koch, 166 [N]. Type Selinusedentatus Gebien, 1914, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Phanechloros Matthews & Bouchard, 300 [M]. Type species [automatic]: Platyphanespunctipennis Carter, 1911, by original designation. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: replacement name for Chlorophanes Matthews, 1992.

Phanerops Solier, 233 [M]. Type Phaneropselongatus Solier, 1851, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Phanerotoma Solier, 82, 126 [F]. Type Phanerotomaelongata Solier, 1843 (= Pimelialaevigata G.-A. Olivier, 1795), by original designation. Status: senior synonym of Ocnodes Fåhraeus, 1870 in Pimeliinae: Sepidiini: Molurina. Synonymy: Gebien (1937a: 759), Kamiński et al. (2019b: 55). Note: junior homonym of Phanerotoma Wesmael, 1838 [Hymenoptera].

Phanerotomea Koch, 58 [F]. Type species [automatic]: Phanerotomaelongata Solier, 1843 (= Pimelialaevigata G.-A. Olivier, 1795), by original designation. Status: junior synonym of Ocnodes Fåhraeus, 1870 in Pimeliinae: Sepidiini: Molurina. Note: replacement name for Phanerotoma Solier, 1843.

Phayllidius Gebien, 451 [M]. Type Phayllidiusdispar Gebien, 1922, by monotypy. Status: junior synonym of Ulomoides Blackburn, 1888 in Diaperinae: Diaperini: Diaperina. Synonymy: Grimm (2018: 103).

Phayllus Champion, 167 [M]. Type Phayllusminutus Champion, 1886, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Phediodes Campbell, 26 [M]. Type Phediodesapterus Campbell, 1976, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Phedius Champion, 447 [M]. Type Phediuschevrolati Champion, 1888, by subsequent designation (R. Lucas 1920: 500). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Phegoneus Casey, 380, 426 [M]. Type Epitragodesjulichi Casey, 1891, by original designation. Status: valid genus and subgenus in Pimeliinae: Epitragini.

Phellidius J.L. LeConte, 236 [M]. Type Bolitophaguscornutus Fabricius, 1801, by original designation. Status: junior synonym of Bolitotherus Candèze, 1861 in Tenebrioninae: Bolitophagini. Synonymy: J.L. LeConte (1866a: 62). Note: the younger species name Bolitophaguscornutus Fabricius, 1801 was given priority over the older synonym Opatrumbifurcum Fabricius, 1798 by the ICZN (2019, Opinion 2438).

Pheloneis Pascoe, 483 [M]. Type Adeliumharpaloides White, 1846 (= Adeliumamaroides Bates, 1874), by original designation. Status: valid genus in Lagriinae: Adeliini.

Phelopatrum Marseul, 100 [N]. Type Hadrusscaphoides Marseul, 1876, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Phenus Gebien, 324, 325 [M]. Type Phenuslatitarsis Gebien, 1921, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pheres Champion, 150 [M]. Type Pheresbatesi Champion, 1886, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Pheugonius Fairmaire, 313 [M]. Type Pheugoniusborneensis Fairmaire, 1899, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Phibalus Gistel, 384 [M]. Type Chrysomelapubescens Linnaeus, 1767, by monotypy. Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini.

Philhamellus Kaszab, 83, 84 [M]. Type Philhammusmyrmecophilus Kaszab, 1960, by original designation. Status: valid subgenus of Philhammus Fairmaire, 1871 in Pimeliinae: Cnemeplatiini: Cnemeplatiina.

Philhammus Fairmaire, 393 [M]. Type Philhammussericans Fairmaire, 1871, by monotypy. Status: valid genus and subgenus in Pimeliinae: Cnemeplatiini: Cnemeplatiina.

Philolithus Lacordaire in J.L. LeConte, 18 [M]. Type Pelecyphoruscarinatus J.L. LeConte, 1851, by subsequent designation (Casey 1912: 79). Status: valid genus and subgenus in Pimeliinae: Asidini.

Philorea Erichson, 242 [F]. Type Philoreapicipes Erichson, 1834, by monotypy. Status: valid genus in Pimeliinae: Physogasterini.

Phligra Laporte, 197 [F]. Type Sepidium degeeri Laporte, 1840 (= Tenebriocristatus DeGeer, 1778), by monotypy. Status: junior synonym of Cyrtoderes Dejean, 1834 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Lacordaire (1859a: 202).

Phobelius Blanchard, pl. 14 [M]. Type Phobeliuscrenatus Blanchard, 1842, by monotypy. Status: valid genus in Lagriinae: Goniaderini. Note: see Bouchard and Bousquet (2020a: 99) for comments about the date of publication of this genus.

Phrenapates Gray in Griffith and Pidgeon, pl. 50 [M]. Type Phrenapatesbennettii Gray, 1831, by monotypy. Status: valid genus in Phrenapatinae: Phrenapatini.

Phrynocarenum Gebien, 106 [N]. Type Phrynocarenumbruchianum Gebien, 1928 (= Emmalloderastrangulata Fairmaire, 1905), by monotypy. Status: valid genus in Pimeliinae: Phrynocarenini.

Phrynocolopsis Koch, 93 [F]. Type Phrynocolusfrondosus Gerstaecker, 1871, by original designation. Status: valid subgenus of Phrynocolus Lacordaire, 1859 in Pimeliinae: Sepidiini: Molurina.

Phrynocolus Lacordaire, 201 [M]. Type species [automatic]: Cryptogeniusdentatus Solier, 1843, by original designation. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Molurina. Note: replacement name for Cryptogenius Solier, 1843.

Phrynophanes Koch, 92 [M]. Type Molurisgredleri Haag-Rutenberg, 1877, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Phthora Champion, 531 [F]. Type species [automatic]: Phtoracrenata Mulsant, 1854 (= Clamorisinsurgens Gozis, 1886), by monotypy. Status: junior synonym of Clamoris Gozis, 1886 in Phrenapatinae: Penetini. Note: unjustified emendation of Phtora Mulsant, 1854, not in prevailing usage; junior homonym of Phthora Gemminger, 1870 [Coleoptera: Tenebrionidae: Diaperinae: Phaleriini].

Phthora Gemminger in Gemminger and Harold, 1959 [F]. Type species [automatic]: Phtoracrenata Germar, 1836, by monotypy. Status: junior synonym of Phtora Germar, 1836 in Diaperinae: Phaleriini. Note: unjustified emendation of Phtora Germar, 1836, not in prevailing usage.

Phtora Germar, pl. 11 [F]. Type Phtoracrenata Germar, 1836, by monotypy. Status: valid genus and subgenus in Diaperinae: Phaleriini. Note: the alternative original spelling Phthora, used by Germar (1836: explanation for pl. 11), was rejected by Neave (1940: 736) who acted as First Reviser.

Phtora Mulsant, 228 [F]. Type Phtoracrenata Mulsant, 1854 (= Clamorisinsurgens Gozis, 1886), by monotypy. Status: senior synonym of Clamoris Gozis, 1886 in Phrenapatinae: Penetini. Note: junior homonym of Phtora Germar, 1836 [Coleoptera: Tenebrionidae: Diaperinae: Phaleriini].

Phygoscotus Schulz, 134 [M]. Type species [automatic]: Spheniscuserotyloides W. Kirby, 1819, by monotypy. Status: junior synonym of Cuphotes Champion, 1887 in Stenochiinae: Stenochiini. Note: replacement name for Spheniscus W. Kirby, 1819.

Phylacastus Fairmaire, 116 [M]. Type Phylacastusstriolatus Fairmaire, 1897, by monotypy. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Phylacinus Fairmaire, 349 [M]. Type Phylacinusasperipennis Fairmaire, 1896, by monotypy. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Phylan Sturm, 23 [M]. Type Opatrumgibbum Fabricius, 1775, by subsequent monotypy (Steven 1828: 99). Status: valid genus and subgenus in Blaptinae: Dendarini: Dendarina. Note: originally proposed without included nominal species; Steven (1828: 99), by including the species Opatrumgibbum Fabricius, 1775 in association with this name, was the first author to subsequently and expressly include nominal species in Phylan (ICZN 1999, Article 67.2.2).

Phylanmania Ferrer, 38 [F]. Type Phylan ilerdensis Español & Viñolas, 1981, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Phylax Brullé, 209 [M]. Type species [automatic]: Opatrumgibbum Fabricius, 1775, by subsequent monotypy (Steven 1828: 99). Status: junior synonym of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina. Note: unnecessary replacement name of Phylan Sturm, 1826 (see Bouchard and Bousquet 2020a: 100).

Phylaximon Koch, 414 [M]. Type Opatrumvariolosum G.-A. Olivier, 1812, by original designation. Status: valid subgenus of Allophylax Bedel, 1906 in Blaptinae: Dendarini: Melambiina.

Phyletes Redtenbacher, 128 [M]. Type Phyletespopuli Chevrolat, 1847 (= Diaperisbifasciata Say, 1824), by subsequent monotypy (Chevrolat 1847b: 57). Status: junior synonym of Alphitophagus Stephens, 1832 in Diaperinae: Diaperini: Adelinina. Synonymy: Heyden et al. (1883: 133). Note: genus originally proposed without included nominal species; Chevrolat (1847b: 57) included the species “Phylethuspopuli Még.” as a synonym of Alphitophagusquadripustulatus Stephens, 1832 in the genus “Phylethus” and Redtenbacher (1848: 589) used “Phylethuspopuli” as valid subsequently; therefore, the author of the type species dates back to its first publication as a synonym (ICZN 1999, Article 11.6.1).

Phyllechus Bouchard & Bousquet, new genus [M]. Type Phyllechusboettcheri Kulzer, 1966, by present designation. Status: valid genus in Stenochiinae: Stenochiini. Note: Kulzer (1966: 374) introduced the new genus name Phyllechus for two new species, but unfortunately did not designate a type species; the genus Phyllechus, which has been treated as valid since 1966, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Phyllechusboettcheri Kulzer, 1966 as type species and referring to Kulzer (1966: 374) for the character states that characterise and differentiate Phyllechus.

Phyloradix Endrödy-Younga, 15, 16 [M]. Type Caenocrypticussoror Endrödy-Younga, 1996, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini.

Phymaeus Pascoe, 439 [M]. Type Phymaeuspustulosus Pascoe, 1883, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Phymatestes Pascoe, 142 [M]. Type species [automatic]: Lagriatuberculata Fabricius, 1787, by monotypy (see ICZN 1989, Opinion 1525). Status: valid genus in Lagriinae: Goniaderini. Note: replacement name for Phymatodes Dejean, 1834; placed on the Official List of Generic Names in Zoology by the ICZN (1989, Opinion 1525).

Phymatiotris Solier, 63 [F]. Type Tentyriaquadricollis Brullé, 1832, by subsequent designation (Hope 1841: 118). Status: valid genus in Pimeliinae: Pimeliini.

Phymatium Billberg, 31 [N]. Type Pimeliamaculata Fabricius, 1781, by present designation. Status: senior synonym of Cryptochile Latreille, 1828 in Pimeliinae: Cryptochilini: Calognathina. Synonymy: new synonym [PB]. Note: discovery of this forgotten name threatens the stability of the junior objective synonym Cryptochile Latreille, 1828; although Phymatium Billberg, 1820 has not been used as valid in the literature after 1899, we could not find usage of Cryptochile Latreille, 1828 in at least 25 works, published by at least ten authors in the immediately preceding 50 years and therefore reversal of precedence cannot be used to treat Phymatium Billberg, 1820 as a nomen oblitum; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to conserve usage of Cryptochile Latreille, 1828, the type genus of the valid tribe Cryptochilini and the subtribe Cryptochilina.

Phymatodes Dejean, 203 [M]. Type Lagriatuberculata Fabricius, 1787, by monotypy. Status: senior synonym of Phymatestes Pascoe, 1866 in Lagriinae: Goniaderini. Note: name suppressed for the purposes of the Principle of Homonymy and the Principle of Priority by the ICZN (1989, Opinion 1525).

Phymatoplata Koch, 269 [F]. Type Selinusasperulus Fairmaire, 1897, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Phymatosoma Laporte & Brullé, 332, 408 [N]. Type Phymatosomatuberculatum Laporte & Brullé, 1831, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: unjustified emendation of the original spelling Phymatisoma, introduced by Agassiz (1846b: 290), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Physadesmia Penrith, 7, 35 [F]. Type Adesmiaglobosa Haag-Rutenberg, 1875, by original designation. Status: valid genus in Pimeliinae: Adesmiini.

Physciolagria Pic, 7 [F]. Type Physciolagrialiturata Pic, 1930, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: we could not assign this genus to any particular group in Tenebrionidae based on the description.

Physignathus Gistel, 22 [M]. Type Helopsundatus Fabricius, 1792 (= Erotylusnebulosus Fabricius, 1781), by monotypy. Status: junior synonym of Cymatothes Dejean, 1834 Tenebrioninae: Amarygmini. Synonymy: Bousquet and Bouchard (2017: 132). Note: junior homonym of Physignathus Cuvier, 1829 [Reptilia].

Physocoelus Haldeman, 347 [M]. Type Psorodesinflata Solier, 1848 (= Helopscontractus Palisot de Beauvois, 1812), by monotypy. Status: junior synonym of Meracantha W. Kirby, 1837 in Tenebrioninae: Amarygmini. Synonymy: Schaum (1850: 181, through synonymy of the type species).

Physodera Solier, 78, 125 [F]. Type Pimeliagibba Fabricius, 1787, by original designation. Status: junior synonym of Moluris Latreille, 1802 in Pimeliinae: Sepidiini: Molurina. Synonymy: Lacordaire (1859a: 194). Note: junior homonym of Physodera Eschscholtz, 1829 [Coleoptera: Carabidae].

Physogaster Lacordaire, 276 [F]. Type Physogastermendocina Lacordaire, 1830, by monotypy. Status: valid genus in Pimeliinae: Physogasterini.

Physogasterinus Kaszab, 79 [M]. Type Physogasterinuslanuginosus Kaszab, 1981, by original designation. Status: valid genus in Pimeliinae: Physogasterini.

Physogria Borchmann, 48, 108 [F]. Type Lagriagibbosa Kolbe, 1901, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Physohelops Schuster, 50 [M]. Type Physohelopsfreyi Schuster, 1937, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Physolagria Fairmaire, 114 [F]. Type Physolagriamolleri Fairmaire, 1891, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Physophrynus Fairmaire, l [M]. Type Physophrynusburdoi Fairmaire, 1882, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Physosterna Dejean, 179 [F]. Type Pimeliaovata G.-A. Olivier, 1795 (= Tenebriotorulosus Pallas, 1781), by monotypy. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Phytolostoma Koch, 34 [N]. Type Phytolostomalimpopoana Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Phitophilus Guérin-Méneville, pl. 4 [M]. Type Phitophilushelopioides Guérin-Méneville, 1831, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Piccula Bousquet & Bouchard in Bousquet et al., 2015: 137 [F]. Type species [automatic]: Gerardiasublineata Pic, 1954, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: replacement name for Gerardia Pic, 1954.

Piciella Borchmann, 237, 435 [F]. Type Piciellahelopioides Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Picnotagalus Kaszab, 102 [M]. Type Picnotagalushorni Kaszab, 1939, by original designation. Status: junior synonym of Scolytocaulus Fairmaire, 1896 in Phrenapatinae: Penetini. Synonymy: Schawaller (1999b: 422).

Picocamaria Masumoto, 226, 232 [F]. Type Camariageniculata Pic, 1915, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Piesomera Solier, 77 [F]. Type Pimeliascabra Fabricius, 1775, by monotypy. Status: junior synonym of Psammodes W. Kirby, 1819 in Pimeliinae: Sepidiini: Molurina. Synonymy: Gebien (1937a: 759).

Piesterotarsa Sénac, 8 [F]. Type species [automatic]: Pimeliagigantea Fischer, 1820, by subsequent designation (Gebien 1937a: 840). Status: junior synonym of Pisterotarsa Motschulsky, 1860 in Pimeliinae: Pimeliini. Note: unjustified emendation of Pisterotarsa Motschulsky, 1860, not in prevailing usage.

Piestognathoides Kaszab, 305 [M]. Type Piestognathoidesbahrainicus Kaszab, 1981, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Piestognathus P.H. Lucas, clxxxviii [M]. Type Piestognathusdouei P.H. Lucas, 1858, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Pigeocaulinus Kaszab, 355, 388 [M]. Type Pigeocaulinussumatranus Kaszab, 1984 (= Leprocaulinuskrikkeni Kaszab, 1982), by original designation. Status: junior synonym of Leprocaulinus Kaszab, 1982 in Stenochiinae: Cnodalonini. Synonymy: Masumoto (2003: 60); see notes in the entry for Leprocaulinus Kaszab, 1982 regarding the validity of this older name.

Pigeostrongylium Kaszab, 355, 385 [N]. Type Pigeostrongyliumkedahense Kaszab, 1984, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pigeus Gebien, 28, 153 [M]. Type Camarimenanitidipes Fairmaire, 1893, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pilioloba Erichson in Agassiz, 144 [F]. Type Salaxlacordairii Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Salax Guérin-Méneville, 1834 in Pimeliinae: Trilobocarini. Synonymy: Erichson in Agassiz (1846a: 144). Note: the name Pilioloba was listed as synonym of Salax Guérin-Méneville, 1834 by Erichson in Agassiz (1846a: 144), it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Cazurro Ruiz 1894b: 451), Pilioloba was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Pilobalia Burmeister, 487 [F]. Type Nycteliadecorata Erichson, 1834, by subsequent designation (Rye 1877: 338). Status: valid genus in Pimeliinae: Nycteliini.

Pilobaloderes Kulzer, 192 [M]. Type Pilobaloderesgebieni Kulzer, 1958, by original designation. Status: valid genus in Pimeliinae: Praociini.

Pilosocasnonidea Pic, 31 [F]. Type Nemostirabipartita Pic, 1911, by monotypy. Status: valid subgenus of Casnonidea Fairmaire, 1882 in Lagriinae: Lagriini: Statirina.

Pilosoplonyx Bremer, 37 [M]. Type Plesiophthalmusbremeri Masumoto, 1999, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Piloxys Fairmaire, 23 [M]. Type Piloxysfoveatus Fairmaire, 1895, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pimalius Casey, 367 [M]. Type Trimytispulverea Horn, 1870, by original designation. Status: junior synonym of Trimytis J.L. LeConte, 1851 in Pimeliinae: Edrotini. Synonymy: MacLachlan and Olson (1990: 79).

Pimelia Fabricius, 251 [F]. Type Pimeliaangulata Fabricius, 1775, by subsequent designation (Hope 1841: 118). Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Pimeliocnera Reitter, 310 [F]. Type Pimeliocneragebieni Reitter, 1909, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Pimelionotus Ardoin, 86 [M]. Type Psorodeslugens Fåhraeus, 1870, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Pimeliopsis Champion, 477 [F]. Type Pimeliopsisgranulata Champion, 1892, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Pimelipachys Skopin, 232 [M]. Type Pachyscelislaevicollis Reitter, 1893, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Pimelosomus Burmeister, 488, 489 [M]. Type Pimelosomussphaericus Burmeister, 1875, by monotypy. Status: valid genus in Pimeliinae: Physogasterini.

Pimidia Rafinesque, 113 [F]. Type species [automatic]: Pimeliaangulata Fabricius, 1775, by subsequent designation (Hope 1841: 118). Status: junior synonym of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Note: unjustified emendation of Pimelia Fabricius, 1775, not in prevailing usage.

Pimplema Pascoe, 16 [F]. Type Pimplemaampliata Pascoe, 1887 (= Platydemahemisphaerica Laporte & Brullé, 1831), by monotypy. Status: valid genus in Diaperinae: Leiochrinini.

Piscicula Robiche, 736 [F]. Type Pisciculasprecherae Robiche, 2004, by monotypy. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Pisterotarsa Motschulsky, 532 [F]. Type Pimeliagigantea Fischer, 1820, by subsequent designation (Gebien 1937a: 840). Status: valid genus in Pimeliinae: Pimeliini.

Pitholaus Champion, 446 [M]. Type Pitholaushelopioides Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Pizura Novák, 436 [F]. Type Pizurabarbucha Novák, 2016, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Plamius Fairmaire, 30 [M]. Type Plamiustenuestriatus Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Planasida Escalera, 337 [F]. Type Asidapaulinoi Pérez Arcas, 1868, by subsequent designation (Viñolas and Cartagena 2005: 223). Status: junior synonym of Gracilasida Escalera, 1905 in Pimeliinae: Asidini. Synonymy: F. Soldati (2008: 137). Note: Planasida Escalera, 1907 was used as valid in the literature recently (e.g., F. Soldati 2008: 137); however, the name Gracilasida Escalera, 1905 is older and should be used as valid (e.g., F. Soldati 2020: 152–153).

Planibates Kaszab, 221 [M]. Type Planibatespapuanus Kaszab, 1939, by monotypy. Status: junior synonym of Bradymerus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Schawaller (2013b: 163).

Planirostrosis Penrith, 18, 172 [F]. Type Zophosishimba Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Planodes Mulsant & Rey, 94, 117 [M]. Type Planodesbyrroides Mulsant & Rey, 1859, by subsequent designation (R. Lucas 1920: 517). Status: senior synonym of Planostibes Gemminger, 1870 in Blaptinae: Opatrini: Stizopodina. Note: junior homonym of Planodes Newman, 1842 [Coleoptera: Cerambycidae].

Planoodescelis Egorov, 591 [F]. Type Oodesceliskansouensis Kaszab, 1940, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Planoplatyscelis Kaszab, 157, 206 [F]. Type Platyscelismargelanica Kraatz, 1882 (= Bioramixpamirensis Bates, 1879), by plenary powers (ICZN 1993a, Opinion 1728). Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1993a, Opinion 1728); the First Reviser (Planoplatyscelis Kaszab, 1940 versus Pleioplatyscelis Kaszab, 1940) is Egorov (2004: 601).

Planoprosodes G.S. Medvedev, 90 [M]. Type Prosodesreitteri Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Note: originally described as a section within a subgenus.

Planostibes Gemminger in Gemminger and Harold, 1926 [M]. Type species [automatic]: Planodesbyrroides Mulsant & Rey, 1859, by subsequent designation (R. Lucas 1920: 517). Status: valid genus in Blaptinae: Opatrini: Stizopodina. Note: replacement name for Planodes Mulsant & Rey, 1859.

Plastica C.O. Waterhouse, 563 [F]. Type Plasticapolita C.O. Waterhouse, 1903, by monotypy. Status: valid genus in Tenebrioninae: Apocryphini.

Platamodes Ménétriés, 233 [M]. Type Platamodesdentipes Ménétriés, 1849, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Platamodina.

Platasida Casey, 77, 182 [F]. Type Asidaembaphionides Horn, 1894, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Plateia Laporte, 215 [F]. Type Plateiaorientalis Laporte, 1840, by monotypy. Status: junior synonym of Catapiestus Perty, 1831 in Stenochiinae: Cnodalonini. Synonymy: Blanchard (1845: 16). Note: junior homonym of Plateia Hübner, 1820 [Lepidoptera].

Platesthes G.R Waterhouse, 317 [F]. Type Platesthessilphoides G.R. Waterhouse, 1845, by monotypy. Status: valid genus in Pimeliinae: Praociini.

Platolenes Gebien, 420 [M]. Type Platolenesrufipes Gebien, 1914, by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer (2001a: 69).

Platyallecula Blair, 266 [F]. Type Cistelabrunnea C.O. Waterhouse, 1876, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Platyblaps Motschulsky, 531 [F]. Type Blaps holconota Fischer von Waldheim, 1844, by subsequent designation (Nabozhenko 2008: 36). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860)

Platybolium Blair, 222 [N]. Type Platyboliumalvearium Blair, 1938, by original designation. Status: valid genus in Tenebrioninae: Triboliini.

Platyburak Iwan, 124 [M]. Type Notocoraxnervosus Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platyburmanicus Iwan, 715 [M]. Type Platyburmanicusignotus Iwan, 2003, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platycharlesus Iwan, 307 [M]. Type Trigonopusmorosus Mulsant & Rey, 1853, by original designation. Status: junior synonym of Schelodontes Koch, 1956 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan and Kamiński (2014: 171).

Platycilibe Carter, 205 [F]. Type Platycilibebrevis Carter, 1911, by monotypy. Status: junior synonym of Scolytocaulus Fairmaire, 1896 in Phrenapatinae: Penetini. Synonymy: Kaszab (1978a: 168).

Platycolpotus Iwan, 255, 266 [M]. Type Platydendarusdendaroides Kaszab, 1975, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platycotylus Olliff, 182 [M]. Type Platycotylusinusitatus Olliff, 1883 (= Ipsaphesnitidulus W.J. MacLaeay, 1873), by monotypy. Status: valid genus in Tenebrioninae: Palorini. Note: originally described in the family Cucujidae, transferred to Tenebrionidae by Crowson (1955: 103).

Platycrepis Lacordaire, 418 [F]. Type Platycrepisviolacea Lacordaire, 1859, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Platycrypticus Español, 9, 17 [M]. Type Crypticusviaticus Fairmaire, 1851, by original designation. Status: valid subgenus of Crypticus Latreille, 1816 in Diaperinae: Crypticini. Note: replacement name for Ulomoides Escalera, 1927. Note: Español (1952a: 117) introduced the replacement name Platycrypticus earlier; however, the name is unavailable from that date since the author did not fix a type species (ICZN 1999, Article 13.3.1).

†Platycteniopus Chang, Nabozhenko, Pu, Xu, Jia & Li, 291 [M]. Type Platycteniopusdiversoculatus Chang, Nabozhenko, Pu, Xu, Jia & Li, 2016, by original designation. Status: valid genus in Alleculinae: Cteniopodini. Note: described from Upper Cretaceous deposits (China).

Platydema Laporte & Brullé, 332, 350 [F]. Type Diaperisviolacea Fabricius, 1790, by subsequent designation (Westwood 1838: 32). Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: see Löbl and Smetana (2010: 34) for comments on the gender of this name.

Platydemoides Kaszab, 161 [M]. Type Platydemoidesbrincki Kaszab, 1980, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: Platydemoides was used earlier by Kaszab (1979a: 77) without a description, a definition or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Platydendarus Kaszab, 281, 312 [M]. Type Opatrumjavanum Wiedemann, 1819, by original designation. Status: junior synonym of Notocorax Dejean, 1834 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (1990: 123).

Platyesia Skopin, 335 [F]. Type Diesiakarelini Fischer von Waldheim, 1844, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Platyesthus Mäklin, 92 [M]. Type Platyesthuspallidipennis Mäklin, 1878, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Platyholmus Dejean, 180 [M]. Type Praocisdilaticollis Lacordaire, 1830, by subsequent designation (Solier 1841a: 270 [370]). Status: valid genus in Pimeliinae: Praociini.

Platykochius Iwan, 48, 88 [M]. Type Selinusplatessa Fairmaire, 1887, by original designation. Status: junior synonym of Anchophthalmops Koch, 1956 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 89).

Platylus Mulsant & Rey, 70 [M]. Type Blapsdilatata Fabricius, 1798, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Platymedvedevia Iwan & Banaszkiewicz, 725 [F]. Type Platymedvedeviademeyeri Iwan & Banaszkiewicz, 2007, by monotypy. Status: junior synonym of Angolositus Koch, 1955 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 90).

Platynoscelis Kraatz, 91 [F]. Type Platynoscelishelopioides Kraatz, 1882, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Platynosum Mulsant & Rey, 73, 81, 150 [N]. Type Platynosumpaulinae Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Sclerina. Note: the spelling Platysum (pp. 73, 81) was corrected to Platynosum in the “Errata” of the same work (p. 150), Platynosum is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Platynotoides Kaszab, 281, 296 [M]. Type Platynotusbellii Fairmaire, 1896, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platynotus Fabricius, 138 [M]. Type Blapsexcavata Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platyolmus Burmeister, 492 [M]. Type species [automatic]: Praocisdilaticollis Lacordaire, 1830, by subsequent designation (Solier 1841a: 270 [370]). Status: junior synonym of Platyholmus Dejean, 1834 in Pimeliinae: Praociini. Note: unjustified emendation of Platyholmus Dejean, 1834, not in prevailing usage.

Platyolus Mulsant & Rey, 153 [M]. Type Microsituslongulus Mulsant & Rey, 1854, by subsequent designation (Viñolas 1990: 65). Status: valid subgenus of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina.

Platyope Fischer, pl. 15 [F]. Type Tenebrioleucographus Pallas, 1781, by subsequent designation (Jacquelin du Val 261). Status: valid genus in Pimeliinae: Pimeliini. Note: Pallas (1773) used two different spellings for this type species, Tenebrioleucographus (p. 474) and Tenebrioleucogrammus (p. 719); subsequently, Pallas (1781: 54) used T.leucographus as the valid name of the species and so acted as First Reviser (ICZN 1999: Article 24.2.4).

Platyotus Gerstaecker, 62 [M]. Type Platyotusglabratus Gerstaecker, 1871, by monotypy. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Halstead (1967a: 72). Note: name also described as new in Gerstaecker (1873: 185).

Platyphanes Westwood, 206 [M]. Type Platyphanesgibbosus Westwood, 1849, by subsequent designation (Lacordaire 1859b: 410). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Platyphanus Koch, 26 [M]. Type Acestussimilis Haag-Rutenberg, 1878, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Platyprocnemis Español & Lindberg, 19, 21 [F]. Type Trichosternumgranulosum Wollaston, 1868, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Platyprosodes Reitter, 121 [M]. Type Nyctipatesrugulosus Gebler, 1841, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Skopin (1960a: 46).

Platypsorodes Ardoin, 307, 310 [M]. Type Acanthomerushelopioides Guérin-Méneville, 1834, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Platyscelis Latreille, 23 [F]. Type Tenebriohypolithus Pallas, 1781, by plenary powers (ICZN 1993b, Opinion 1729). Status: valid genus and subgenus in Blaptinae: Platyscelidini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1993b, Opinion 1729).

Platysemodes Strand, 302 [M]. Type species [automatic]: Platysemusbenguelensis Haag-Rutenberg, 1875, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Platysemus Haag-Rutenberg, 1875.

Platysemus Haag-Rutenberg, 4, 55 [M]. Type Platysemusbenguelensis Haag-Rutenberg, 1875, by monotypy. Status: senior synonym of Platysemodes Strand, 1935 in Pimeliinae: Adelostomini. Note: junior homonym of Platysemus Middendorff, 1848 [Mollusca].

Platystena Koch, 95 [F]. Type Mesostenablairi Koch, 1940, by monotypy. Status: valid subgenus of Mesostena Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: combined description of a new genus-group taxon and a single new species (ICZN 1999, Article 13.4).

Plegacerus Gebien, 142 [M]. Type Plegacerussensitivus Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Pleioplatyscelis Kaszab, 153, 220 [F]. Type Platynoscelislederi Seidlitz, 1893, by original designation. Status: junior synonym of Planoplatyscelis Kaszab, 1940 in Blaptinae: Platyscelidini. Synonymy: Egorov (2004: 601).

Pleiopleura Seidlitz, 342, 343 [F]. Type Platyscelisstriatus Motschulsky, 1859, by monotypy. Status: valid subgenus of Platyscelis Latreille, 1818 in Blaptinae: Platyscelidini.

Pleisiasida Smith, 610 [F]. Type species [automatic]: Parasidalaciniata Casey, 1912, by original designation. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini. Note: replacement name for Parasida Casey, 1912.

Plesia Klug, 25 [F]. Type Plesiamelanura Klug, 1833, by subsequent designation (Bousquet et al. 2015: 137). Status: senior synonym of Eubalia Laporte, 1840 in Alleculinae: Alleculini: Gonoderina. Synonymy: Borchmann (1909a: 713). Note: junior homonym of Plesia Jurine, 1807 [Hymenoptera].

Plesiamarygmus Masumoto, 314 [M]. Type Dietysusovoideus Fairmaire, 1882, by original designation. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer (2005: 205).

Plesioderes Mulsant & Rey, 126 [M]. Type Caediuscoriaceus Mulsant & Rey, 1859, by subsequent designation (Gebien 1939: 466). Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Plesiognaptor Chigray, Nabozhenko & Keskin, 1277 [M]. Type Gnaptorprolixus Fairmaire, 1866, by original designation. Status: valid subgenus of Gnaptor Brullé, 1831 in Blaptinae: Blaptini: Gnaptorina.

Plesiophthalmus Motschulsky, 34 [M]. Type Plesiophthalmusnigrocyaneus Motschulsky, 1857, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Pleuromophlus Reitter, 138, 146 [M]. Type Omophlusbaudueri Baudi, 1877, by subsequent designation (Bousquet et al. 2015: 141). Status: junior synonym of Odontomophlus Seidlitz, 1896 in Alleculinae: Cteniopodini. Synonymy: Bousquet et al. (2015: 141).

Pleurophorus Solier, 162 [M]. Type Pleurophorusquadricollis Solier, 1851, by monotypy. Status: senior synonym of Discopleurus Lacordaire, 1859 in Pimeliinae: Stenosini: Dichillina. Note: junior homonym of Pleurophorus Mulsant, 1842 [Coleoptera: Scarabaeidae].

Pleurostira Borchmann, 217, 230 [F]. Type Pleurostiraepipleuralis Borchmann, 1921, by original designation. Status: valid subgenus of Statira Lepeletier & Audinet-Serville, 1828 in Lagriinae: Lagriini: Statirina.

Pleuroxycara Koch, 582 [N]. Type Oxycaraproblematicum Koch, 1959, by original designation. Status: valid subgenus of Oxycara Solier, 1835 in Pimeliinae: Tentyriini.

Plicatocerus Pic, 11 [M]. Type Otocerusimpressipennis Champion, 1888, by monotypy. Status: valid subgenus of Oploptera Chevrolat, 1844 in Stenochiinae: Stenochiini.

Plinthochrous Fairmaire, 263 [M]. Type Plinthochrousgounellei Fairmaire, 1891, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Pocadiopsis Fairmaire, 17 [F]. Type Pocadiopsissimulator Fairmaire, 1896, by subsequent designation (Gebien 1939: 457). Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Podacamptus Ardoin, 849 [M]. Type Podacamptusruficolor Ardoin, 1964, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Podamarygmus Carter, 287 [M]. Type Podamarygmusalternatus Carter, 1928 (= Amarygmusviridipes Gebien, 1927), by monotypy. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Podhomala Solier, 9, 72 [F]. Type Podhomalasuturalis Solier, 1836, by monotypy. Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Podoces Péringuey, 122 [F]. Type Podocesgranosula Péringuey, 1886, by present designation. Status: senior synonym of Carchares Pascoe, 1887 in Tenebrioninae: Scaurini. Synonymy: Péringuey (1892b: [135]). Note: junior homonym of Podoces Fischer, 1821 [Aves].

Pododonta Agassiz, 300 [F]. Type species [automatic]: Cistelanigrita Fabricius, 1794, by subsequent designation (R. Lucas 1920: 527). Status: junior synonym of Podonta Solier, 1835 in Alleculinae: Cteniopodini. Note: unjustified emendation of Podonta Solier, 1835, not in prevailing usage.

Podomala Agassiz, 300 [F]. Type species [automatic]: Podhomalasuturalis Solier, 1836, by monotypy. Status: junior synonym of Podhomala Solier, 1836 in Pimeliinae: Pimeliini. Note: unjustified emendation of Podhomala Solier, 1836, not in prevailing usage.

Podonta Solier, 247 [F]. Type Cistelanigrita Fabricius, 1794, by subsequent designation (Cazurro Ruiz 1894b: 883). Status: valid genus in Alleculinae: Cteniopodini.

Podontinus Seidlitz, 197 [M]. Type Cteniopuspunctatissimus Kiesenwetter, 1861, by subsequent designation (R. Lucas 1920: 527). Status: valid genus in Alleculinae: Cteniopodini.

Poecilesthostrongylium Pic, 12 [N]. Type Strongyliumamazonicum Pic, 1918, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Poecilesthus Dejean, 207 [M]. Type Erotylusfasciatus Fabricius, 1781, by subsequent designation (Hope 1841: 133). Status: valid genus in Stenochiinae: Stenochiini. Note: Poecilesthus is an incorrect subsequent spelling of the original spelling Paecilesthus, first used by Dejean (1836: 229), and in prevailing usage; Poecilesthus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1), see Bousquet and Bouchard (2013a: 60).

Poecilocrypticus Gebien, 121 [M]. Type Poecilocrypticusformicophilus Gebien, 1928, by monotypy. Status: valid genus in Diaperinae: Crypticini.

Poeciltoides Fairmaire, 352 [M]. Type Poeciltoidesalternans Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pogonobasis Solier, 153, 161 [F]. Type Pogonobasisopatroides Solier, 1837 (= Eurychorarugosula Guérin-Méneville, 1831), by subsequent designation (Hope 1841: 121). Status: valid genus in Pimeliinae: Adelostomini.

Pogonocanta Koch, 22 [F]. Type Pogonobasislongipilis Fairmaire, 1894, by original designation. Status: valid genus in Pimeliinae: Adelostomini. Note: the alternative original spelling Pogonacanta, used by Koch (1952b: 123), was rejected by Koch (1953c: 3) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Pogonophloeus Bremer, 9 [M]. Type Hypophlaeusthoracicus Melsheimer, 1846, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Pogonoxenus Wasmann, 172 [M]. Type Pogonoxenuskraatzi Wasmann, 1899, by monotypy. Status: valid genus in Diaperinae: Hypophlaeini.

Pokryszkiella Iwan, 385, 414 [F]. Type Pokryszkiellacornuta Iwan, 1996, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Polasida Reitter, 39, 41 [F]. Type Opatrumsericeum G.-A. Olivier, 1795, by subsequent designation (Viñolas and Cartagena 2005: 188). Status: valid subgenus of Asida Latreille, 1802 in Pimeliinae: Asidini.

Polemiotus Casey, 379, 381 [M]. Type Epitragussubmetallicus J.L. LeConte, 1854, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Poliorcetes Champion, 70 [M]. Type Poliorcetesplatesthoides Champion, 1884, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Polopinus Casey, 326 [M]. Type Polypleurusnitidus J.L. LeConte, 1866, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Polpocara Solier, 46, 123 [N]. Type Polpocarapicipes Solier, 1843, by original designation. Status: junior synonym of Philorea Erichson, 1834 in Pimeliinae: Physogasterini. Synonymy: Erichson (1847a: 116).

Polpogenia Solier, 9, 70 [F]. Type Polpogeniaasidioides Solier, 1836, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Polposipus Solier, 154, 260 [M]. Type Polposipusherculeanus Solier, 1848, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Polycoelogastridion Reichardt, 85, 208 [N]. Type Sclerumsexcostatum Motschulsky, 1858, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Polyidus Champion, 441 [M]. Type Polyidusmeridionalis Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Polypleurus Eschscholtz, 10, 11 [M]. Type Polypleurusgeminatus Eschscholtz, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Polyscopus Waltl, 73 [M]. Type Polyscopuscostatus Waltl, 1835 (= Adelostomasulcatum Duponchel, 1827), by monotypy. Status: junior synonym of Adelostoma Duponchel, 1827 in Pimeliinae: Adelostomini. Synonymy: Erichson in Agassiz (1846a: 133).

Polytropus Kirsch, 201 [M]. Type Polytropuslaenoides Kirsch, 1866, by monotypy. Status: junior synonym of Chaetyllus Pascoe, 1860 in Lagriinae: Laenini. Synonymy: Kaszab (1983a: 138).

Ponapeida Kulzer, 242, 248 [F]. Type Ponapeidarufitarsis Kulzer, 1957, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pontianacus Fairmaire, 397 [M]. Type Pontianacusrubricrus Fairmaire, 1898, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Poopterocoma Skopin, 145 [F]. Type Pterocomazaidamica Skopin, 1974, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Porphyrhyba Fairmaire, 137 [F]. Type Porphyrhybaviolaceicolor Fairmaire, 1877, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Porphyrohyba Rye, 62 [F]. Type species [automatic]: Porphyrhybaviolaceicolor Fairmaire, 1877, by monotypy. Status: junior synonym of Porphyrhyba Fairmaire, 1877 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Porphyrhyba Fairmaire, 1877, not in prevailing usage.

Porrolagria Kolbe, 26 [F]. Type Porrolagrianuda Kolbe, 1883, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Posides Champion, 6 [M]. Type Posidesdissidens Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Postandrosus Kulzer, 490 [M]. Type Postandrosusmaculipennis Kulzer, 1951, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Postpraocis Flores & Pizarro-Araya, 60 [M]. Type Praocispentachorda Burmeister, 1875, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 12, 33) without type species designation.

Potocula Novák, 290 [F]. Type Potoculakubani Novák, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Praeugena Laporte, 241 [F]. Type Helopsmarginatus Fabricius, 1792, by subsequent designation (Hope 1841: 133). Status: valid genus in Tenebrioninae: Praeugenini. Note: nomen protectum (see Bouchard and Bousquet 2020b: 6).

†Praezolodinus Bao in Bao and Antunes-Carvalho, 2 [M]. Type Praezolodinuspilosus Bao, 2020, by original designation. Status: valid genus in Zolodininae. Note: described from Cretaceous Burmese amber (Myanmar).

Praocida Flores & Pizarro-Araya, 70 [F]. Type Praociszischkai Kulzer, 1958, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 13, 88) without type species designation.

Praocidia Fairmaire, 463 [F]. Type Praocisnervosus Fairmaire, 1902, by original designation. Status: valid genus in Pimeliinae: Praociini.

Praocis Eschscholtz, 6 [M]. Type Praocisrufipes Eschscholtz, 1829, by subsequent designation (Guérin-Méneville 1834: 8–9). Status: valid genus and subgenus in Pimeliinae: Praociini. Note: although Guérin-Méneville (1834: 8–9) did not specifically mention the scientific name of the species he considered the type of this genus, his comment “J’ai donné une nouvelle figure de l’espèce qui a servi de type à Eschscholtz, dans la pl. 4, fig. 1, du Voyage de la Coquille” [translated to “I gave a new figure of the species that served as the type by Eschscholtz, in pl. 4, fig. 1, of the Voyage de la Coquille”] leaves no ambiguity regarding the identity of the species, i.e., Praocisrufipes Eschscholtz, 1829, to which he was referring.

Praogena Agassiz, 306 [F]. Type species [automatic]: Helopsmarginatus Fabricius, 1792, by subsequent designation (Hope 1841: 133). Status: junior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Note: unjustified emendation of Praeugena Laporte, 1840, not in prevailing usage.

Praonoda Flores & Pizarro-Araya, 68 [F]. Type Praocisbicarinatus Burmeister, 1875, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 13, 66) without type species designation.

Praostetha Fairmaire, 138 [F]. Type Praostethaimpressifrons Fairmaire, 1897, by monotypy. Status: junior synonym of Amenophis J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1921b: 64).

Prateus J.L. LeConte, 238 [M]. Type Prateusfusculus J.L. LeConte, 1862, by original designation. Status: valid genus in Lagriinae: Goniaderini.

Predactylosis Penrith, 19, 243 [F]. Type Predactylosisholmi Penrith, 1977, by original designation. Status: junior synonym of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Synonymy: Penrith (1981a: 22).

Priocamaria Gebien, 28, 143 [F]. Type Priocamariamacilenta Gebien, 1919, by subsequent designation (Gebien 1942a: 323). Status: valid genus in Stenochiinae: Cnodalonini.

Prionalia Novák, 511 [F]. Type Gonoderaatronitens Fairmaire, 1892, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Prionotheca Dejean, 179 [F]. Type Pimeliacoronata G.-A. Olivier, 1795, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Prionotus Mulsant & Rey, 88 [M]. Type Opatrumdenticolle Blanchard, 1846, by monotypy. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Gemminger in Gemminger and Harold (1870: 1929, with Achora Pascoe, 1869, a junior synonym of Isopteron Hope, 1841). Note: junior homonym of Prionotus Lacepède, 1802 [Pisces].

Prionychus Solier, 231, 237 [M]. Type Helopsater Fabricius, 1775, by subsequent designation (R. Lucas 1920: 536). Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: nomen protectum (see Bousquet and Bouchard 2017: 132); see Löbl and Smetana (2011: 33) for comments on the gender of this genus-group name.

Prioproctus Kolbe, 165, 174 [M]. Type Prioproctusoertzeni Kolbe, 1903, by original designation. Status: valid genus in Lagriinae: Pycnocerini.

Prioscelida White, 11 [F]. Type Prioscelidatenebrionoides White, 1846, by monotypy. Status: junior synonym of Uloma Dejean, 1821 in Tenebrioninae: Ulomini. Synonymy: Broun (1880: 365).

Prioscelides Kolbe, 128 [M]. Type Prioscelidesrugosus Kolbe, 1889, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Prioscelis Hope, 128 [F]. Type Prioscelisfabricii Hope, 1841, by original designation. Status: valid genus in Lagriinae: Pycnocerini. Note: the alternative original spelling Priopus, used by Hope (1841: 73), was rejected by Westwood (1844: 211) who acted as First Reviser.

Priothorax Gebien, 318 [M]. Type species [automatic]: Opatrumdenticolle Blanchard, 1846, by monotypy. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Gebien (1910b: 318, as a valid synonym of Achora Pascoe, 1869), Blair (1919a: 529, as a synonym of Achora Pascoe, 1869, a junior synonym of Isopteron Hope, 1841). Note: replacement name for Prionotus Mulsant & Rey, 1859.

Pristophilus Kolbe, 165, 174 [M]. Type Chiroscelispassaloides Westwood, 1842, by original designation. Status: valid genus in Lagriinae: Pycnocerini.

Probaticus Seidlitz, 697, 704, 764 [M]. Type Helopsmori Brullé, 1832, by subsequent designation (Gebien 1943: 419). Status: junior synonym of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko et al. (2017: 496). Note: the alternative original spelling Probatius (pp. 697, 704, 765) was corrected to Probaticus in the “Amendments and corrections” of the same work (p. 849), Probaticus is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Prochoma Solier, 253, 393 [N]. Type Prochomaaudouini Solier, 1835, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Proctenius Reitter, 256 [M]. Type Cistelagranatensis Rosenhauer, 1856, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Proderops Fairmaire, 393 [M]. Type Proderopsforaminosus Fairmaire, 1873 (= Rhinandruselongatus Horn, 1867), by monotypy. Status: junior synonym of Rhinandrus J.L. LeConte, 1866 in Tenebrioninae: Tenebrionini. Synonymy: Kraatz (1880a: 132, with Exerestus Bates, 1870, a junior synonym of Rhinandrus J.L. LeConte, 1866).

Prodilamus Ardoin, 258 [M]. Type Dilamusbrevicollis Fairmaire, 1894, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Prohylithus Kaszab, 382 [M]. Type Prohylithuskulzeri Kaszab, 1964, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Prolabrus Fairmaire, 111 [M]. Type Prolabrusparallelus Fairmaire, 1897, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Prolaena Kaszab, 322 [F]. Type Laenaceylonica Motschulsky, 1858, by original designation. Status: valid genus in Lagriinae: Laenini. Note: Prolaena was used earlier by Kaszab (1979a: 107) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore not available from that date.

Proleptodes G.S. Medvedev, 354 [M]. Type Leptodessulcicollis Reitter, 1889, by original designation. Status: valid subgenus of Leptodes Dejean, 1834 in Pimeliinae: Leptodini.

Promethis Pascoe, 148 [M]. Type Upisangulata Erichson, 1842, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Prometopion Casey, 366, 370 [N]. Type Prometopionamplipenne Casey, 1907 (= Chilometoponhelopioides Horn, 1874), by original designation. Status: junior synonym of Chilometopon Horn, 1874 in Pimeliinae: Edrotini. Synonymy: MacLachlan and Olson (1990: 72).

Promorphostenophanes Kaszab, 277 [M]. Type Promorphostenophanesatavus Kaszab, 1960, by original designation. Status: junior synonym of Morphostenophanes Pic, 1925 in Stenochiinae: Cnodalonini. Synonymy: Masumoto and Bečvář (2008: 206).

Promus J.L. LeConte, 226 [M]. Type Blapsopaca Say, 1824, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Propemicrosis Penrith, 127, 152 [F]. Type Microsistransbechuana Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Prophanes Westwood, 203 [M]. Type Prophanesaculeatus Westwood, 1849, by subsequent designation (Lacordaire 1859b: 411). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Propterocoma Skopin, 144 [F]. Type Pterocomatibialis Bates, 1879, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Prorhytinota Bouchard & Bousquet, new subgenus [F]. Type Rhytinotaoxyoma Fairmaire, 1884, by present designation. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: Koch (1943b: 761, 795) introduced the new subgenus name Prorhytinota for several nominal species, but unfortunately did not designate a type species; the subgenusProrhytinota, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Rhytinotaoxyoma Fairmaire, 1884 as type species and referring to Koch (1943b: 761, 795) for the character states that characterise and differentiate Prorhytinota.

Proscheimus Desbrochers des Loges, 127 [M]. Type Proscheimusarabicus Desbrochers des Loges, 1881, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Proscorus Fairmaire, 188 [M]. Type Proscoruscyaneostriatus Fairmaire, 1901, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Proselytus Fåhraeus, 302 [M]. Type Proselytuscaffer Fåhraeus, 1870 (= Alphitobiusdiaperinus Panzer, 1796), by monotypy. Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Ferrer (1999: 266; through synonymy of the type species).

Prosoblapsia Skopin & Kaszab, 208 [F]. Type Leptocolenaallardiana Reitter, 1889, by original designation. Status: junior synonym of Genoblaps Bauer, 1921 in Blaptinae: Blaptini: Blaptina. Synonymy: new synonym [YB]. Note: Genoblaps Bauer, 1921 has been forgotten in the literature; its type species is currently included in the subgenusProsoblapsia Skopin & Kaszab, 1978 and for that reason Skopin & Kaszab’s name is considered a junior synonym of Genoblaps.

Prosodella Reitter, 120 [F]. Type Prosodesbactriana Semenov, 1894, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Note: the First Reviser (Prosodella Reitter, 1909 versus Paraprosodes Reitter, 1909) is G.S. Medvedev (1997: 597).

Prosodes Eschscholtz, 9 [F]. Type Blapsattenuata Fischer, 1820 (= Blaps obtusa Fabricius, 1798), by subsequent designation (Gebien 1937a: 846). Status: valid genus and subgenus in Blaptinae: Blaptini: Prosodina. Note: the gender of Prosodes is feminine by indication of the author himself, who “von προσώδης, foetida”, the latter word being Latin and explaining the Greek meaning (stinking); the word foetida is feminine and therefore Prosodes is feminine also; note, however, that all other genus-group names in this list with the ending –prosodes are masculine.

Prosodestes Reitter, 114 [M]. Type Prosodesgrandicollis Kraatz, 1883, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosodidius Fairmaire, 69 [M]. Type Prosodidiusperrieri Fairmaire, 1903, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Prosodila Reitter, 121 [F]. Type Prosodesstrigiventris Reitter, 1893, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Löbl et al. (2008c: 219).

Prosodinia Reitter, 115 [F]. Type Prosodescalcarata Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosodopria Reitter, 116 [F]. Type Blapsangustata Zubkov, 1833, by monotypy. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosodoscelis Reitter, 117 [F]. Type Prosodessolskyi Faust, 1875, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosodura Reitter, 118 [F]. Type Prosodessemenowi Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosomenes Blanchard, 10 [M]. Type Diceroderesmexicanus Solier, 1841, by subsequent monotypy (Chevrolat 1847b: 562). Status: junior synonym of Diceroderes Solier, 1841 in Tenebrioninae: Toxicini: Dysantina. Synonym: Lacordaire (1859a: 356, as “Prosomenes Dejean”), Bousquet et al. (2018: 219). Note: the name Prosomenes was originally introduced as a synonym of “Dicérodères” by Blanchard (1845: 10); however, because the name was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Chevrolat 1847b: 562) we treat it as available and dating from its first publication as a synonym (ICZN 1999, Article 11.6.1); originally proposed without included nominal species; Chevrolat (1847b: 562), by including the species “Pros.mexicanus” in association with this name, was the first author to subsequently and expressly include nominal species in Prosomenes (ICZN 1999, Article 67.2.2).

Prostenus Klug, 5 [M]. Type Prostenusperiscelis Perty, 1830, by subsequent designation (Bousquet et al. 2015: 139, but see Note). Status: valid genus in Alleculinae: Alleculini: Xystropodina. Note: as pointed out by Bousquet et al. (2015: 139) this genus name had been attributed to other authors in the literature (e.g., Latreille, 377, Berthold, 369) but it was first made available by Klug (1829: 5) who described two new species in it, Prostenusfemoratus and Prostenuspilosus, neither species being included in Prostenus in its currently accepted sense; Bousquet et al. (2015: 139) suggested using Prostenusperiscelis Perty, 1830 as the type species of Prostenus to promote nomenclatural stability; however, an application to the Commission is needed to confirm the type species of Prostenus.

†Proteleates Wickham,1914a: 267 [M]. Type Proteleatescentralis Wickham, 1914, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini. Note: described from Upper Eocene deposits (USA).

Prothraustocola Kaszab, 293 [F]. Type Ibnsaudiabelutschistanica Kaszab, 1957, by original designation. Status: valid subgenus of Thraustocolus Kraatz, 1866 in Pimeliinae: Tentyriini.

Protoblaps Bauer, 230, 231 [F]. Type none designated. Status: undetermined taxon in Blaptinae: Blaptini: Blaptina. Note: this genus was described before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Protoblaps and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Protoblaps G.S. Medvedev, 200 [F]. Type Protoblapskashkarovi G.S. Medvedev, 1998, by original designation. Status: senior synonym of Medvedevoblaps Bouchard & Bousquet, nom. nov. in Blaptinae: Blaptini: Blaptina. Note: junior homonym of Protoblaps Bauer, 1921 [Coleoptera: Tenebrionidae: Blaptinae: Blaptini: Blaptina].

Protocalosis Penrith, 21, 116 [F]. Type Zophosisbalti Penrith, 1977, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Protodactylus Koch, 83 [M]. Type Protodactylusopticus Koch, 1952, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Protomachlasida Escalera, 137 [F]. Type Machlasidaliouvillei Escalera, 1925, by original designation. Status: junior synonym of Machlasida Escalera, 1907 in Pimeliinae: Asidini. Synonymy: Gebien (1937a: 717).

†Protoplatycera Wickham, 484 [F]. Type Protoplatyceralaticornis Wickham, 1914, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Upper Eocene deposits (USA).

Prototyrtaeus Spiessberger & Ivie, 669 [M]. Type Prototyrtaeusdarlingtoni Spiessberger & Ivie, 2020, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Prunaspila Koch, 67 [F]. Type species [automatic]: Aspilabicostata Fåhraeus, 1870, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Aspila Fåhraeus, 1870.

Przewalskia Semenov, 262 [F]. Type Platyopedilatata Reitter, 1887, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Psammestus Reichardt, 194, 216 [M]. Type Ammobiusdilatatus Reitter, 1893, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Psammetichus Latreille, 578 [M]. Type Psammetichuscostatus Guérin-Méneville, 1831, by subsequent monotypy (Guérin-Méneville 1831b: pl. 28bis). Status: valid genus in Pimeliinae: Elenophorini: Megelenophorina. Note: originally proposed without included nominal species; Guérin-Méneville (1831b: pl. 28bis), by including the new species “Psammeticuscostatus Guérin-Méneville, 1831” in association with this name, was the first author to subsequently and expressly include nominal species in Psammetichus (ICZN 1999, Article 67.2.2).

Psammoardoinellus Leo, 34 [M]. Type Isocerussardiniensis Ardoin, 1972, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Psammocryptus Kraatz, 81, 239 [M]. Type Tentyriaminuta Tauscher, 1812, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Psammodes W. Kirby, 412 [M]. Type Psammodeslongicornis W. Kirby, 1819, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Psammodophysis Péringuey, 296 [F]. Type Psammodophysisprobes Péringuey, 1899, by subsequent designation (Kamiński et al. 2019b: 31). Status: junior synonym of Psammodes W. Kirby, 1819 in Pimeliinae: Sepidiini: Molurina. Synonymy: Gebien (1910a: 154).

Psammoeca Agassiz, 311 [F]. Type species [automatic]: Microderalucida Solier, 1835, by subsequent designation (Gebien 1937a: 611). Status: junior synonym of Psammoica Solier, 1835 in Pimeliinae: Tentyriini. Note: unjustified emendation of Psammoica Solier, 1835, not in prevailing usage.

Psammogaster Koch, 237 [F]. Type Psammogastermalani Koch, 1953, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Psammoica Solier, 307 [F]. Type Microderalucida Solier, 1835, by subsequent designation (Gebien 1937a: 611). Status: valid genus in Pimeliinae: Tentyriini.

Psammolophus Koch, 154 [M]. Type Psammodesacuticosta Fairmaire, 1884, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psammophanes Lesne, 690 [M]. Type Moluriscatenata Reiche, 1850, by original designation. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Molurina.

Psammophrynopsis Koch, 157 [F]. Type Phrynocolusfrommi Wilke, 1921, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psammophrynus Koch, 146 [M]. Type Psammophanesjokli Koch, 1953, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psammoryssus Kolbe, 289 [M]. Type Psammoryssustitanus Kolbe, 1886, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Psammostretus Koch, 145 [M]. Type Psammodesbisbicostatus Gebien, 1910, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psammotopulus Endrödy-Younga, 15, 29 [M]. Type Caenocrypticuspenrithae Endrödy-Younga, 1996, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini.

Psammotyria Koch, 137 [F]. Type Psammodesertli Kolbe, 1904, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Psammotyriopsis Koch, 144 [F]. Type Psammophanesbredoi Koch, 1953, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psaryphis Erichson, 241 [F]. Type Psaryphisnana Erichson, 1843, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Psaryphulum Koch, 32 [N]. Type Adelostomaabbreviatum Haag-Rutenberg, 1875, by original designation. Status: valid subgenus of Adelostoma Duponchel, 1827 in Pimeliinae: Adelostomini.

Psectes Hesse, 572 [M]. Type Psectesbechuanus Hesse, 1935, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Psectrapus Solier, 153, 213 [M]. Type Psectrapusbipartitus Solier, 1848, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Psectrascelis Solier, 307, 311 [F]. Type Nycteliapilipes Guérin-Méneville, 1834, by subsequent designation (Gebien 1937a: 750). Status: valid genus in Pimeliinae: Nycteliini. Note: the First Reviser (Psectrascelis Solier, 1836 versus Cerostena Solier, 1836) is Fairmaire (1876: 356).

Psectropus Gemminger in Gemminger and Harold, 1911 [M]. Type species [automatic]: Psectrapusbibartitus Solier, 1848, by original designation. Status: junior synonym of Psectrapus Solier, 1848 in Blaptinae: Platynotini: Eurynotina. Note: unjustified emendation of Psectrapus Solier, 1848, not in prevailing usage.

Pselaphidion Gebien, 29, 229 [N]. Type Platydemamacularia Gemminger, 1870, by subsequent designation (Gebien 1940: 405). Status: junior synonym of Stomylus Fåhraeus, 1870 in Diaperinae: Diaperini: Diaperina. Synonymy: Koch (1953g: 23). Note: the alternative original spelling Pselaphidium, used by Gebien (1921b: 28, 29), was rejected by Gebien (1940: 405) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Pseudabax Kraatz, 107 [M]. Type Pseudabaxformosus Kraatz, 1880, by subsequent designation (Gebien 1942a: 308). Status: valid genus in Stenochiinae: Cnodalonini.

Pseudadelium Kaszab, 234 [N]. Type Adeliumpustulosum Fauvel, 1904, by original designation. Status: junior synonym of Neoadelium Carter, 1908 in Lagriinae: Adeliini. Synonymy: Matthews (1998: 781).

Pseudadrus Fairmaire, 39 [M]. Type Hadrusscaphoides Marseul, 1876, by monotypy. Status: junior synonym of Phelopatrum Marseul, 1876 in Blaptinae: Opatrini: Opatrina. Synonymy: Reichardt (1936: 90).

Pseudalymon Ardoin, 543, 544 [M]. Type Asthenochirusfoveolatus Péringuey, 1899, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Pseudamarsenes Ardoin, 142 [M]. Type Amarsenesviridistriatus Fairmaire, 1894, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudamarygmus Pic, 9 [M]. Type Pseudamarygmustestaceipes Pic, 1915, by subsequent designation (Gebien 1948: 509). Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer (2001a: 67).

Pseudamblyptera Pierre, 295 [F]. Type Pimeliafrigida Escalera, 1925, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Pseudamenophis Pic, 13 [M]. Type Pseudamenophisaeneus Pic, 1916 (= Amenophisepipleuralis Gebien, 1904), by monotypy. Status: junior synonym of Amenophis J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1962a: 64).

Pseudanaedus Gebien, 107, 111 [M]. Type Pseudanaedusbiangulatus Gebien, 1921, by subsequent designation (Gebien 1941: 820). Status: valid genus in Lagriinae: Goniaderini.

Pseudandrosus Kulzer, 486 [M]. Type Chariothecaneomidina Fairmaire, 1881, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudanemia Wollaston, 492 [F]. Type Pseudanemiabrevicollis Wollaston, 1864, by monotypy. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini.

Pseudanopidium Dajoz, 431, 434 [N]. Type Pseudanopidiumpunctatum Dajoz, 1974, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Pseudapocrypha Champion, 260 [F]. Type Pseudapocryphalacordairii Champion, 1886, by monotypy. Status: valid genus in Tenebrioninae: Apocryphini.

Pseudapsida Kulzer, 219 [F]. Type Pseudapsidabrasiliensis Kulzer, 1961, by original designation. Status: junior synonym of Paniasis Champion, 1886 in Diaperinae: Diaperini: Diaperina. Synonymy: Ferrer and Ødegaard (2005: 637).

Pseudasida Fairmaire, 444 [F]. Type Pseudasidaobesa Fairmaire, 1895, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Pseudeba Blackburn, 119 [F]. Type Pseudebanovica Blackburn, 1903, by monotypy. Status: valid genus in Tenebrioninae: Palorini. Note: transferred from “Tenebrionoidea: Colydiidae” by Carter and Zeck (1937: 194).

Pseudeleodes Blaisdell, 146 [M]. Type Eleodesgranosa J.L. LeConte, 1866, by monotypy. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Pseudemmallus Koch, 355 [M]. Type Pseudemmallusaspericollis Koch, 1956, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Pseudephalus Casey, 333 [M]. Type Pseudephalusbrevicornis Casey, 1924, by original designation. Status: junior synonym of Ephalus J.L. LeConte, 1862 in Blaptinae: Opatrini: Opatrina. Synonymy: Lumen et al. (2020: 344).

Pseudesarcus Champion, 115 [M]. Type Pseudesarcusvillosus Champion, 1913, by original designation. Status: valid genus in Lagriinae: incertae sedis. Note: originally described in Mycetophagidae, transferred to Tenebrionidae: Lagriinae by Lawrence and Newton (1995: 886).

Pseudethas Fairmaire, 57 [M]. Type Pseudethasquadraticeps Fairmaire, 1896, by monotypy. Status: valid genus and subgenus in Pimeliinae: Stenosini: Dichillina.

Pseudethas Fairmaire, 477 [M]. Type Pseudethaslongiceps Fairmaire, 1898, by monotypy. Status: senior synonym of Anethas Jakobson, 1924 in Pimeliinae: Stenosini: Stenosina. Note: junior homonym of Pseudethas Fairmaire, 1896 [Coleoptera: Tenebrionidae: Pimeliinae: Stenosini: Dichillina].

Pseudeucyrtus Pic, 14 [M]. Type Pseudeucyrtusniasensis Pic, 1916 (= Cleomisviolaceipes Fairmaire, 1892), by subsequent designation (Gebien 1942a: 308). Status: junior synonym of Cleomis Fairmaire, 1892 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983b: 382). Note: we act as First Revisers and reject the alternative original spelling Pseudocyrtus, used by Pic (1916e: 14), in order to avoid homonymy with Pseudocyrtus Salmon, 1956 [Collembola].

Pseudeumolpus Kraatz, 111 [M]. Type Pseudeumolpusbicolor Kraatz, 1880, by subsequent designation (Löbl et al. 2008b: 345). Status: junior synonym of Phaedis Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1914d: 71).

Pseudeuthriptera Bogatchev & Kryzhanovsky, 240 [F]. Type Trigonoscelisuzboica Bogatchev & Kryzhanovsky, 1955, by monotypy. Status: junior synonym of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini. Synonymy: Skopin (1964: 395).

Pseudeutrapela Pic, 110 [F]. Type Pseudeutrapelanigra Pic, 1952, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Pseudhadrus Kolbe, 31 [M]. Type Pseudhadrusseriatus Kolbe, 1910, by subsequent designation (Gebien 1941: 332). Status: valid genus in Stenochiinae: Cnodalonini.

Pseudhelops Guérin-Méneville, 124 [M]. Type Pseudhelopstuberculatus Guérin-Méneville, 1841, by monotypy. Status: valid genus in Tenebrioninae: Heleini: incertae sedis. Note: transferred from Titaenini by Matthews and Lawrence (2019: 628).

Pseudimmedia Kulzer, 213 [F]. Type Pseudimmediafreyi Kulzer, 1958, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudisopus Kulzer, 254 [M]. Type Pseudisopusgressitti Kulzer, 1957, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudobasides Pic, 2 [M]. Type Pseudobasidescornutus Pic, 1916, by subsequent designation (Gebien 1940: 419). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Pseudobates Fairmaire, 231 [M]. Type Nyctobatescoracinus Fairmaire, 1882 (= Nyctobatessubrobustus Motschulsky, 1872), by monotypy. Status: junior synonym of Promethis Pascoe, 1869 in Stenochiinae: Cnodalonini. Synonymy: Kolbe (1900: 74, with Setenis Motschulsky, 1872, a junior synonym of Promethis Pascoe, 1869).

Pseudoblapida Pic, 18 [F]. Type Blapidaboliviensis Pic, 1912, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Pseudoblaps Guérin-Méneville, 28 [F]. Type Pseudoblapssubstriata Guérin-Méneville, 1834, by subsequent designation (Hope 1841: 124). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Pseudobolbophanes Kulzer, 29 [M]. Type Pseudobolbophanesmalaisei Kulzer, 1954, by original designation. Status: junior synonym of Bolbophanes Carter, 1913 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 490).

Pseudobradymerus Pic, 16 [M]. Type Bradymerussimplicithorax Pic, 1926, by monotypy. Status: junior synonym of Bradymerus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1939: 751).

Pseudobyrsax Kaszab, 214 [M]. Type Pseudobyrsaxcurtus Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Adeliini.

Pseudocaedius Blackburn, 91 [M]. Type Pseudocaediussquamosus Blackburn, 1890 (= Trigonotarsusaustralis Hope, 1843), by monotypy. Status: junior synonym of Sobas Pascoe, 1863 in Blaptinae: Opatrini: Opatrina. Synonymy: Carter (1921: 307).

Pseudocaedius G.S. Medvedev, 98 [M]. Type Pseudocaediuskiseritzkii G.S. Medvedev, 1966, by original designation. Status: senior synonym of Asiocaedius G.S. Medvedev & Nepesova, 1985 in Blaptinae: Opatrini: Ammobiina. Note: junior homonym of Pseudocaedius Blackburn, 1890 [Coleoptera: Tenebrionidae: Blaptinae: Opatrini: Ammobiina].

Pseudocaelophus Pic, 28 [M]. Type Strongyliumdifforme Pic, 1922, by monotypy. Status: junior synonym of Leprocaulus Fairmaire, 1896 in Stenochiinae: Cnodalonini. Synonymy: Bečvář and Purchart (2008: 39).

Pseudocamaria Bates, 287 [F]. Type Camariaalternata Fairmaire, 1875, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudocamarimena Pic, 21 [F]. Type Pseudocamarimenastriata Pic, 1923, by monotypy. Status: junior synonym of Pigeus Gebien, 1919 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1984: 363).

Pseudocasnonidea Borchmann, 240, 489 [F]. Type Pseudocasnonideaceylanica Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Pseudochariotheca Pic, 32 [F]. Type Pseudochariothecaminutissima Pic, 1934, by original designation. Status: junior synonym of Steneucyrtus Fairmaire, 1896 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 136).

Pseudochillus Fouquè, 226, 240 [M]. Type Indochillusbangaloreanus Kaszab, 1981, by original designation. Status: valid genus and subgenus in Pimeliinae: Stenosini: Dichillina.

Pseudochrysomela Pic, 7 [F]. Type Pseudeumolpusseriatoporus Fairmaire, 1888, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: the older name Pseudochrysomela Voet, 1806 [Coleoptera: Erotylidae] was published in a work that did not include consistent application of binominal nomenclature and is therefore unavailable (ICZN 1999, Article 11.4).

Pseudocilibe Kaszab, 231 [F]. Type Celibeasidaeformis Fauvel, 1904, by original designation. Status: valid genus in Lagriinae: Adeliini. Note: we act as First Revisers and reject the alternative original spelling Pseudocylibe, used by Kaszab (1982b: 291).

Pseudocistela Blackburn, 316 [F]. Type Pseudocistelaovalis Blackburn, 1891, by monotypy. Status: senior synonym of Neocistela Borchmann, 1909 in Alleculinae: Alleculini: Alleculina. Note: junior homonym of Pseudocistela Crotch, 1874 [Coleoptera: Tenebrionidae: Alleculinae: Alleculini: Gonoderina].

Pseudocistela Crotch, 108 [F]. Type Cistelabrevis Say, 1824, by subsequent designation (Novák and Pettersson 2008: 327). Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Pseudocistelopsis Novák, 176 [M]. Type Pseudocistelopsisjakli Novák, 2018, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Pseudocoelus Casey, 152 [M]. Type Coeluspacificus Fall, 1897, by subsequent designation (Doyen 1976: 608). Status: junior synonym of Coelus Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Blaisdell (1919: 322).

Pseudocolparthrum Borchmann, 230, 236 [N]. Type Colparthrumcalcaratum Champion, 1889, by subsequent designation (Borchmann 1936: 452). Status: valid subgenus of Colparthrum Kirsch, 1866 in Lagriinae: Lagriini: Statirina.

Pseudoderiles Gebien, 169, 181 [M]. Type Pseudoderilesdentipennis Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudoderosphaerus Pic, 24 [M]. Type Leprocaulusrotundicollis Pic, 1922, by monotypy. Status: junior synonym of Leprocaulus Fairmaire, 1896 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983c: 177).

Pseudodiaphanidus Bogatchev, 234 [M]. Type Diaphanidusrobustus Bogatchev, 1950, by monotypy. Status: valid subgenus of Diaphanidus Reitter, 1900 in Pimeliinae: Erodiini.

Pseudoelongasida Escalera, 173 [F]. Type Asidasilvestrei Escalera, 1922, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Elongasida Escalera, 1906 in Pimeliinae: Asidini. Synonymy: Pérez-Vera et al. (2017: 3).

Pseudoenanea Pic, 25 [F]. Type Pseudoenanearobusta Pic, 1924, by monotypy. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Pseudogena Fairmaire, 539 [F]. Type Pseudogenapolyzona Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Pseudognaptorina Kaszab, 250 [F]. Type Pseudognaptorinanepalica Kaszab, 1977, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

†Pseudohelops Haupt, 130 [M]. Type Pseudohelopsgroenlandicus Haupt, 1950, by monotypy. Status: valid genus in Stenochiinae: incertae sedis. Note: combined description of new genus-group taxon and new species (ICZN 1999, Article 13.4); described from Middle Paleocene deposits (Greenland).

Pseudohymenalia Novák, 213 [F]. Type Pseudohymenaliayunnanica Novák, 2008, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Pseudolagria Champion, 218 [F]. Type Pseudolagriamutabilis Champion, 1917, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Pseudolamus Fairmaire, 388 [M]. Type Pseudolamusseriatoporus Fairmaire, 1874, by monotypy. Status: valid genus in Blaptinae: Opatrini: Neopachypterina.

Pseudoleichenum Ardoin, 205 [N]. Type Pseudoleichenumbenoiti Ardoin, 1972, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Pseudolyprops Fairmaire, 236 [M]. Type Pseudolypropsdilaticollis Fairmaire, 1882, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Pseudomachla Wilke, 260 [F]. Type species [automatic]: Opatrumvillosum G.-A. Olivier, 1795, by subsequent designation (R. Lucas 1920: 386). Status: junior synonym of Machla Herbst, 1799 in Pimeliinae: Asidini. Note: unnecessary replacement name for Machla Herbst, 1799.

Pseudomorocaulus Pic, 6 [M]. Type Pseudomorocaulusrufonotatus Pic, 1915, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Pseudonautes Fairmaire, 52 [M]. Type Pseudonautesvagevittatus Fairmaire, 1892, by subsequent designation (R. Lucas 1920: 551). Status: valid genus in Stenochiinae: Cnodalonini.

Pseudonomus Fairmaire, 510 [M]. Type Pseudonomusdermestiformis Fairmaire, 1884, by monotypy. Status: junior synonym of Ammodonus Mulsant, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1939: 470).

Pseudonotocorax Iwan, 255, 269 [M]. Type Pseudonotocoraxmroczkowskii Iwan, 1997, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Pseudoogeton Masumoto, 304 [M]. Type Plesiophthalmusamplipennis Fairmaire, 1897, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Pseudopachyscelis Skopin, 99 [F]. Type Trigonoscelispygmaea Ménétriés, 1849, by original designation. Status: valid genus in Pimeliinae: Pimeliini. Note: first proposed by Skopin (1962: 225) without a type species originally designated.

Pseudoparablops Heyden, 132 [M]. Type Parablopssardiniensis Allard, 1877, by monotypy. Status: junior synonym of Allardius Ragusa, 1898 in Tenebrioninae: Helopini: Helopina. Synonymy: Reitter (1922b: 167).

Pseudopatrum Sharp, 406 [N]. Type Pseudopatrumsordidum Sharp, 1886, by subsequent designation (Watt 1992: 30). Status: valid genus in Lagriinae: Adeliini.

Pseudopedinus Ardoin, 178 [M]. Type Pseudopedinuslaosensis Ardoin, 1969, by original designation. Status: valid subgenus of Loensus R. Lucas, 1920 in Blaptinae: Pedinini: Pedinina.

Pseudopenthicinus Bogatchev, 628 [M]. Type Penthicusmedvedevi Bogatchev, 1972, by original designation. Status: valid subgenus of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina.

Pseudoperichilus Pic, 21 [M]. Type Pseudoperichilusolivaceus Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudophthora Kaszab, 267 [F]. Type Pseudophthoralaeana Kaszab, 1970, by original designation. Status: valid genus and subgenus in Phrenapatinae: Penetini.

Pseudopigeus Kaszab, 355, 359 [M]. Type Pseudopigeusunidentatus Kaszab, 1984, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudopimelia Gebler, 473 [F]. Type Lasiostolavariolaris Gebler, 1841, by monotypy. Status: junior synonym of Lasiostola Dejean, 1834 in Pimeliinae: Pimeliini. Synonymy: new synonym [YB]. Note: Pseudopimelia Gebler, 1859 has been forgotten in the literature; its type species is currently included in the nominotypical subgenus of Lasiostola Dejean, 1834 and for that reason Gebler’s name is considered a new junior synonym of Lasiostola.

Pseudopimelia Motschulsky, 536 [F]. Type Pterocomatuberculata Motschulsky, 1845, by subsequent designation (Skopin 1974b: 159). Status: senior synonym of Subpterocoma Bouchard & Bousquet, nom. nov. in Pimeliinae: Pimeliini. Note: junior homonym of Pseudopimelia Gebler, 1859 [Coleoptera: Tenebrionidae: Pimeliinae: Pimeliini].

Pseudoplanasida Escalera, 360 [F]. Type Asidapygmaea Rosenhauer, 1856, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Gracilasida Escalera, 1905 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 192, with Planasida Escalera, 1907, a junior synonym of Gracilasida Escalera, 1905).

Pseudoplatyope Pierre, 867, 873 [F]. Type Storthocnemisantoinei Español, 1951, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Pseudopodhomala Schuster, 88 [F]. Type Pseudopodhomalagabrieli Schuster, 1938, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Pseudopodhomalina Kaszab, 22 [F]. Type Diesiacostifera C.O. Waterhouse, 1889, by original designation. Status: junior synonym of Pseudopodhomala Schuster, 1938 in Pimeliinae: Pimeliini. Synonymy: Kwieton (1982: 35).

Pseudopraeugena De Moor, 8,40 [F]. Type Pseudopraeugenarufa De Moor, 1970, by original designation. Status: valid genus in Tenebrioninae: Praeugenini.

Pseudoprobaticus Nabozhenko, 513 [M]. Type Helopsgranipennis Allard, 1876, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Pseudoprosodes Reitter, 120 [M]. Type Prosodestransfuga Reitter, 1893, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Skopin (1960a: 46).

Pseudopterocoma Skopin, 145 [F]. Type Pterocomatrapezicollis Skopin, 1974, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Pseudorozonia Bouchard & Bousquet, new subgenus [F]. Type Rozoniaconophthalma Koch, 1944, by present designation. Status: valid subgenus of Rozonia Fairmaire, 1888 in Pimeliinae: Tentyriini. Koch (1944a: 162) introduced the new subgenus name Pseudorozonia for three nominal species, but unfortunately did not designate a type species; the subgenusPseudorozonia, which has been treated as valid since 1944, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Rozoniaconophthalma Koch, 1944 as type species and referring to Koch (1944a: 162) for the character states that characterise and differentiate Anemiadena.

Pseudortheolus Freude, 110 [M]. Type Epitragusminutissimus Fairmaire, 1892, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Pseudoscaphidema Pic, 2 [F]. Type Pseudoscaphidemarufonotata Pic, 1926, by monotypy. Status: valid genus in Diaperinae: Scaphidemini.

Pseudoscotobius Kulzer, 384, 393 [M]. Type Emmalloderastrangulata Fairmaire, 1905, by original designation. Status: junior synonym of Phrynocarenum Gebien, 1928 in Tenebrioninae: Phrynocarenini. Synonymy: Marcuzzi (1976: 117).

Pseudoselinus Iwan, 48, 94 [M]. Type Eurynotuspunctatostriatus Gerstaecker, 1854, by original designation. Status: valid subgenus of Upembarus Koch, 1956 in Blaptinae: Platynotini: Platynotina.

Pseudoseriscius Español, 125 [M]. Type Pedinuspruinosus Dufour, 1820, by original designation. Status: valid genus and subgenus in Diaperinae: Crypticini.

Pseudostene Wollaston, 247 [F]. Type Pseudosteneangusta Wollaston, 1861, by subsequent designation (Löbl et al. 2008b: 317). Status: junior synonym of Phtora Germar, 1836 in Diaperinae: Phaleriini. Synonymy: Escalera (1914: 347).

Pseudosternoplax Skopin, 110, 141 [F]. Type Trigonoscelislacerta Bates, 1879, by original designation. Status: valid subgenus of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini.

Pseudostira Fairmaire, 213 [F]. Type Pseudostiralaevipennis Fairmaire, 1903, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Pseudostorthocnemis Gridelli, 83 [F]. Type Storthocnemispatrizii Gridelli, 1933, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Pseudostrongylium Kraatz, 114 [N]. Type Pseudostrongyliumsemperi Kraatz, 1880, by subsequent designation (Kaszab 1977b: 11). Status: junior synonym of Lophocnemis Mäklin, 1867 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 543).

Pseudotalpophila Reitter, 95 [F]. Type Thalpophilaplicifrons Wollaston, 1864, by subsequent designation (Gebien 1937a: 641). Status: valid subgenus of Hegeter Latreille, 1802 in Pimeliinae: Tentyriini.

Pseudothinobatis Bouchard & Bousquet, new genus [F]. Type Thinobatisohausi Kulzer, 1956, by present designation. Status: valid genus in Pimeliinae: Epitragini. Note: Freude (1960b: 32) introduced the new genus name Pseudothinobatis for two nominal species, but unfortunately did not designate a type species; the genus Pseudothinobatis, which has been treated as valid since 1960, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Thinobatisohausi Kulzer, 1956 as type species and referring to Freude (1960b: 32) for the character states that characterise and differentiate Pseudothinobatis.

Pseudothryoneus Pic, 12 [M]. Type Pseudothryoneusbicoloripes Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudotocerus Champion, 383 [M]. Type Stenochialongipes P.H. Lucas, 1859, by subsequent designation (Gebien 1948: 542). Status: valid genus in Stenochiinae: Stenochiini.

Pseudotrichoplatyscelis Kaszab, 83, 111 [F]. Type Platynoscelisbadakschanica Kaszab, 1960 (= Bioramixlapidicola Kaszab, 1940), by original designation. Status: junior synonym of Trichoplatyscelis Reinig, 1931 in Blaptinae: Platyscelidini. Synonymy: Egorov (1990: 402). Note: we act as First Revisers and reject the alternative original spellings Pseudotrichoplatynoscelis and Pseudotrichoplatycelis, used by Kaszab (1960a: 82, 83).

Pseuduloma Fairmaire, 27 [N]. Type Pseudulomacribricolle Fairmaire, 1893 (= Ulomimusindicus Bates, 1873), by monotypy. Status: junior synonym of Ulomimus Bates, 1873 in Tenebrioninae: Ulomini. Synonymy: Gebien (1940: 770).

Pseuduroplatopsis Pic, 16 [F]. Type Borchmanniajavana Pic, 1913, by present designation. Status: valid subgenus of Borchmannia Pic, 1912 in Lagriinae: Lagriini: Statirina.

Psilachnopus Reitter, 161 [M]. Type Psilachnopuscribratellus Reitter, 1901, by monotypy. Status: junior synonym of Philhammus Fairmaire, 1871 in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Synonymy: Gebien (1938a: 420), Löbl and Smetana (2010: 30).

Psilocastus Ardoin, 149 [M]. Type Psilocastusletestui Ardoin, 1963, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: Psilocastus was used earlier by Ardoin (1962b: 970) without designation of a type species and is therefore unavailable from that date.

Psilolaena Heller, 70 [F]. Type Psilolaenaschusteri Heller, 1923, by monotypy. Status: junior synonym of Laena Dejean, 1821 in Lagriinae: Laenini. Synonymy: Schawaller (2001b: 34).

Psilomera Motschulsky, 400 [F]. Type Pelecyphorusangulatus J.L. LeConte, 1851, by monotypy. Status: junior synonym of Stenomorpha Solier, 1836 in Pimeliinae: Asidini. Synonymy: Bousquet in Bousquet et al. (2018: 74).

Psilonesogena Bates, 305 [F]. Type Psilonesogenahybrida Bates, 1879, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Psis Novák, 71 [M]. Type Psisnanensis Novák, 2019, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Psoroderes Ardoin, 969, 1017 [M]. Type Psorodeshottentottus Péringuey, 1899, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Psorodes Dejean, 189 [M]. Type species [automatic]: Pimeliadentipes Fabricius, 1787, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: replacement name for Acanthomera Latreille, 1828.

Psorophodes Ardoin, 83 [F]. Type Pimeliaarmata Herbst, 1799, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: Psorophodes was used earlier by Ardoin (1962b: 969) without designation of a type species and is therefore unavailable from that date.

Psydocamaria Pic, 17 [F]. Type Psydocamariarobusta Pic, 1923, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Psydomorphus Pic, 24 [M]. Type Psydomorphusdiversipes Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Psydus Pascoe, xii [M]. Type Psydusplantaris Pascoe, 1868, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pteraulus Solier, 152, 200 [M]. Type Pteraulussulcatipennis Solier, 1848, by present designation. Status: junior synonym of Helopinus Solier, 1848 in Blaptinae: Pedinini: Helopinina. Synonymy: Koch (1958: 149).

Pterelaeus Gemminger in Gemminger and Harold, 1968 [M]. Type species [automatic]: Pterohelaeuswalkerii Brême, 1842, by subsequent designation (Gebien 1940: 1069). Status: junior synonym of Pterohelaeus Brême, 1842 in Tenebrioninae: Heleini: Heleina. Note: unjustified emendation of Pterohelaeus Brême, 1842, not in prevailing usage.

Pterna Reitter, 249 [F]. Type Ernocharisauricoma Reitter, 1884, by monotypy. Status: valid subgenus of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: as mentioned by Bousquet et al. (2015: 138) this name was originally proposed as a synonym of Mycetochares Latreille, 1829 but was used as valid subsequently and adopted as the name of a taxon (e.g., Seidlitz 1896: 132, 137) before 1961; therefore, it is available and dates from its publication as a synonym (ICZN 1999, Article 11.6.1).

Pterocoma Dejean, 178 [F]. Type Pimeliapiligera Gebler, 1830, by monotypy. Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Pterocomodes Reitter, 159 [M]. Type Pterocomodesacutus Reitter, 1901, by monotypy. Status: junior synonym of Podhomala Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Skopin (1962: 247).

Pteroctenus Kirsch, 193 [M]. Type Pteroctenuspexus Kirsch, 1866, by monotypy. Status: valid genus in Pimeliinae: Falsomycterini.

Pterodes Ardoin, 97 [M]. Type Hoplonyxasper Péringuey, 1899, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: Pterodes was used earlier by Ardoin (1962: 970) without a type species designation and is therefore unavailable from that date.

Pteroglymmius Gebien, 219, 223 [M]. Type Pteroglymmiuserotyloides Gebien, 1928, by monotypy. Status: junior synonym of Isaminas Champion, 1887 in Stenochiinae: Cnodalonini. Synonymy: Doyen (1988: 301).

Pterohelaeus Brême, 17, 27 [M]. Type Pterohelaeuswalkerii Brême, 1842, by subsequent designation (Gebien 1940: 1069). Status: valid genus in Tenebrioninae: Heleini: Heleina.

Pterolasia Solier, 9, 66 [F]. Type Pterolasiasqualida Solier, 1836, by subsequent designation (Hope 1841: 118). Status: valid genus in Pimeliinae: Pimeliini.

Pteroselinus Kamiński, 92, 94 [M]. Type Opatrinusinsularis Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Pterostichula Koch, 224 [F]. Type Pterostichulacalathoides Koch, 1952, by original designation. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Oxurina.

Ptilonix Allard, 62 [M]. Type Helopsclavicrus Marseul, 1876, by subsequent designation (Löbl et al. 2008a: 42). Status: junior synonym of Misolampidius Solsky, 1876 in Stenochiinae: Cnodalonini. Synonymy: Lewis (1894: 476).

Pubamarygmus Pic, 8 [M]. Type Pubamarygmusviridipennis Pic, 1915, by subsequent designation (Gebien 1948: 510). Status: valid genus in Tenebrioninae: Amarygmini.

Pubeirosoma Pic, 257 [N]. Type Isomiradiscoglabrata Pic, 1954, by monotypy. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Pulposipes Gemminger in Gemminger and Harold, 1974 [M]. Type species [automatic]: Polposipusherculeanus Solier, 1848, by original designation. Status: junior synonym of Polposipus Solier, 1848 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Polposipus Solier, 1848, not in prevailing usage.

Pumiliofossorum Silvestro & Giraldo-Mendoza in Silvestro et al., 2015: 462, 470 [N]. Type Pumiliofossorummoche Silvestro & Flores, 2015, by original designation. Status: valid genus in Tenebrioninae: Scotobiini.

Punctacula Campbell, 112 [F]. Type Punctaculahowdeni Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Pushtunillus G.S. Medvedev, 855, 865 [M]. Type Dichillusskopini G.S. Medvedev & Kabakov, 1995, by original designation. Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Pyanirygmus Pic, 9 [M]. Type Pyanirygmuscorinthius Pic, 1915, by monotypy. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Pyanisia Laporte, 235 [F]. Type Helopsundatus Fabricius, 1792 (= Erotylusnebulosus Fabricius, 1781), by subsequent designation (Lacordaire 1859b: 476). Status: junior synonym of Cymatothes Dejean, 1834 in Tenebrioninae: Amarygmini. Synonymy: Chevrolat (1847b: 643).

Pycna Fairmaire, 68 [F]. Type Pycnaaphodina Fairmaire, 1894, by monotypy. Status: senior synonym of Madagassa Koch, 1950 in Phrenapatinae: Penetini. Note: redescribed as new by Fairmaire (1894b: 141); junior homonym of Pycna Amyot & Audinet-Serville, 1843 [Hemiptera].

Pycnocerus Westwood, 67 [M]. Type species [automatic]: Pachyloceruswestermanni Hope, 1841, by monotypy. Status: valid genus and subgenus in Lagriinae: Pycnocerini. Note: replacement name for Pachylocerus Hope, 1841.

Pycnochilus C.O. Waterhouse, 263 [M]. Type Pycnochilusadvenus C.O. Waterhouse, 1879, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Pycnomorpha Motschulsky, 398 [F]. Type Pycnomorphacalifornica Motschulsky, 1870, by monotypy. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Pycnonotida Casey, 75, 89 [F]. Type Microschatiainaequalis J.L. LeConte, 1851, by original designation. Status: junior synonym of Microschatia Solier, 1836 in Pimeliinae: Asidini. Synonymy: K.W. Brown and Doyen (1992: 546).

Pycnuloma Fairmaire, 99 [N]. Type Pycnulomaraffrayi Fairmaire, 1896, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Pygidiphorus Mulsant, [1, supplement] [M]. Type Pygidiphoruscaroli Mulsant, 1856, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: included in a list of “Tenebrionidaenomina dubia” by Iwan et al. (2020: 475).

Pygmaeodes Koch, 223 [M]. Type Namibomodesrudebecki Koch, 1952, by monotypy. Status: valid subgenus of Palpomodes Koch, 1952 in Pimeliinae: Sepidiini: Oxurina.

Pyres Champion, 100 [M]. Type Pyresmetallicus Champion, 1885 (= Centronopusspeciosus Pascoe, 1883), by subsequent designation (Gebien 1941: 336). Status: junior synonym of Menechides Motschulsky, 1872 in Tenebrioninae: Centronopini. Synonymy: Spilman (1962b: 3).

†Pyrochalcaspis Haupt, 113, 115 [F]. Type Pyrochalcaspisgiseltalensis Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Pystelops Gozis, 103 [M]. Type Helopsmeridianus Mulsant, 1854, by original designation. Status: valid subgenus of Stenomax Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina.

Pythiopus Koch, 245 [M]. Type Pythiopuscornutipectus Koch, 1953, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Pythonissus Gistel, 21 [M]. Type Helopsmorio Fabricius, 1777 (= Tenebrioatratus Fabricius, 1775), by subsequent designation (Bousquet and Bouchard 2017: 132). Status: junior synonym of Zophobas Dejean, 1834 in Tenebrioninae: Tenebrionini. Synonymy: Bousquet and Bouchard (2017: 132).

Quadrideres Koch, 189 [M]. Type Anchophthalmusscutatus Gerstaecker, 1871, by original designation. Status: junior synonym of Glyptopteryx Gebien, 1910 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 91).

Quadroncotus Koch, 40 [M]. Type Oncotusirrepertus Koch, 1954, by original designation. Status: valid subgenus of Oncotus Blanchard, 1845 in Blaptinae: Platynotini: Eurynotina.

Raiboscelis Allard, 5 [F]. Type Helopscorvinus Küster, 1850, by subsequent designation (Gebien 1943: 415). Status: valid genus in Tenebrioninae: Helopini: Helopina. Note: nomen protectum (see Nabozhenko and Löbl 2009: 194); Raiboscelis is an incorrect subsequent spelling of the original spelling Raibosceles and is in prevailing usage; Raiboscelis is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1), see Nabozhenko and Löbl (2009: 189).

Raptor Gistel, ix, xi [M]. Type species [automatic]: Melaphorusreichii Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Melaphorus Guérin-Méneville, 1834 in Pimeliinae: Evaniosomini. Note: unnecessary replacement name for Melaphorus Guérin-Méneville, 1834.

Rasphytus Kulzer, 639 [M]. Type Rasphytusfreyi Kulzer, 1956, by original designation. Status: valid subgenus of Apentanodes Reitter, 1914 in Pimeliinae: Erodiini.

Raynalius Chatanay, 297 [M]. Type Raynaliushispidus Chatanay, 1912, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Rehumius Fairmaire, 32 [M]. Type Rehumiusamplithorax Fairmaire, 1893, by subsequent designation (Gebien 1942a: 312). Status: valid genus in Stenochiinae: Cnodalonini.

Reichardtiella Kaszab, 18 [F]. Type Reichardtiellaarmata Kaszab, 1942, by original designation. Status: senior synonym of Reichardtiellina Kaszab, 1982 in Blaptinae: Opatrini: Opatrina. Note: junior homonym of Reichardtiella Filipjev, 1928 [Lepidoptera].

Reichardtiellina Kaszab, 79 [F]. Type species [automatic]: Reichardtiellaarmata Kaszab, 1942, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: replacement name for Reichardtiella Kaszab, 1942 (as “Reichardtia Kaszab, 1940”; see Iwan and Löbl 2007: 735).

Reichenspergeria Wasmann, 18, 19 [F]. Type Reichenspergeriaaurocincta Wasmann, 1921, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Reitterella Semenov, 362 [F]. Type Reitterellafusiformis Semenov, 1891, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Reitterellus Escalera, 91 [M]. Type Dendarusdepressus Reitter, 1915, by monotypy. Status: junior synonym of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina. Synonymy: Español (1961a: 44).

Reitterohelops Skopin, 308, 309 [M]. Type Zophohelopslazarus Reitter, 1922, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Reminius Casey, 321 [M]. Type Reminiusocularis Casey, 1924 (= Tenebrioterminatus Say, 1824), by original designation. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Spilman (1959: 63).

Remipedella Semenov-Tjan-Shansky, 257 [F]. Type Remipedelladeserti Semenov-Tjan-Shansky, 1907, by monotypy. Status: valid genus in Blaptinae: Blaptini: Remipedellina.

Renatiella Koch, 155 [F]. Type Macropodareticulata Gerstaecker, 1854, by monotypy. Status: valid genus in Pimeliinae: Adesmiini. Note: the First Reviser (Renatiella Koch, 1944 versus Spongesmima Koch, 1944) is Penrith (1979: 27).

Renefouqueosis Aalbu, Smith, Kanda & Bouchard, 314 [F]. Type Renefouqueosisperuviensis Aalbu, Smith, Kanda & Bouchard, 2017, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Rhacius Champion, 120 [M]. Type Rhaciussulcatulus Champion, 1885, by subsequent designation (Gebien 1941: 805). Status: junior synonym of Adelonia Laporte, 1840 in Lagriinae: Belopini. Synonymy: Spilman (1961: 50).

Rhacolaena Kaszab, 299 [F]. Type Rhacolaenatarsalis Kaszab, 1979, by original designation. Status: valid genus in Lagriinae: Laenini.

Rhaebosceles Rye, 69 [F]. Type species [automatic]: Helopscorvinus Küster, 1850, by subsequent designation (Gebien 1943: 415). Status: junior synonym of Raiboscelis Allard, 1876 in Tenebrioninae: Helopini: Helopina. Note: unjustified emendation of Raiboscelis Allard, 1876, not in prevailing usage.

Rhagostira Borchmann, 238, 468 [F]. Type Rhagostiracollarti Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Rhaibodera Borchmann, 217, 219 [F]. Type Rhaiboderapachycera Borchmann, 1921 (= Statiraeurydera Champion, 1917), by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Rhaibogria Borchmann, 17, 144 [F]. Type Lagriaampla Fairmaire, 1887, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Rhammatodes Haag-Rutenberg, 83 [M]. Type Rhammatodeslongicornis Haag-Rutenberg, 1876, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: the First Reviser (Rhammatodes Haag-Rutenberg, 1876 versus Euleantus Haag-Rutenberg, 1876) is Koch (1952a: 133).

Rhicnodus Fairmaire, 87 [M]. Type Rhicnodusasper Fairmaire, 1892, by subsequent designation (R. Lucas 1920: 568). Status: junior synonym of Chaetyllus Pascoe, 1860 in Lagriinae: Laenini. Synonymy: Ardoin (1969b: 126), Kaszab (1983a: 132).

Rhinandrus J.L. LeConte, 119 [M]. Type Rhinandrusgracilis J.L. LeConte, 1866, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Rhinobarus Reitter, 131 [M]. Type Cistelasulphuripes Germar, 1823, by subsequent designation (Iablokoff-Khnzorian 1983: 133). Status: valid subgenus of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini.

†Rhinohelaeites Haupt, 115, 140 [M]. Type Rhinohelaeiteslongipes Haupt, 1950, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Middle Eocene deposits (Germany); this genus was previously considered to be “definitely not a tenebrionid” by Watt (1975: 389) but is included here following Nabozhenko (2019: 8).

Rhipidandrus J.L. LeConte, 236 [M]. Type Xylotinusflabellicornis Sturm, 1826 (= Melolonthaparadoxa Palisot de Beauvois, 1818), by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Rhipidonyx Reitter, 304 [M]. Type Rhipidonyxadustus Reitter, 1876, by monotypy. Status: valid genus in Alleculinae: incertae sedis. Note: taxon originally described in Mycetophagidae, transferred to Tenebrionidae by Lawrence et al. (2014: 209).

Rhizalemus Reitter, 79 [M]. Type Dendarusreitteri Seidlitz, 1893, by subsequent designation (Chatzimanolis and Löbl 2003: 260). Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Rhizalus Gebien, 303 [M]. Type species [automatic]: Opatrumpiceum G.-A. Olivier, 1812, by monotypy. Status: junior synonym of Rizalus Mulsant & Rey, 1854 in Blaptinae: Dendarini: Dendarina. Note: unjustified emendation of Rizalus Mulsant & Rey, 1854, not in prevailing usage.

Rhizoblaps Motschulsky, 532 [F]. Type Blapspruinosa Faldermann, 1833, by subsequent designation (Nabozhenko 2008: 36). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860)

Rhomaleus Chatanay, 64 [M]. Type Rhomaleusscauroides Chatanay, 1915, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Rhopalobates Fairmaire, 230 [M]. Type Rhopalobatesvillardi Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Rhophobas Motschulsky, 36 [M]. Type Rhophobasasperatus Motschulsky, 1872, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Rophobas, used by Motschulsky (1872: 26).

Rhosaces Champion, 73 [M]. Type Rhosacesclavipes Champion, 1889, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Rhostax Fischer von Waldheim, 67 [M]. Type Rhostaxmenetriesii Fischer von Waldheim, 1844 (= Microderagracilis Eschscholtz, 1831), by subsequent designation (G.S. Medvedev 1990: 109). Status: junior synonym of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Kraatz (1865: 184).

Rhydimorpha Koch, 768, 863 [F]. Type Tentyriasubfossulata Solier, 1835, by monotypy. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: we act as First Revisers and reject the alternative original spelling Rhytimorpha, used by Koch (1943b: 888), since Rhytimorpha Szépligeti, 1901 is available in Hymenoptera.

Rhypasma Pascoe, 325 [N]. Type Rhypasmapusillum Pascoe, 1862, by monotypy. Status: valid genus in Lagriinae: Belopini.

Rhysodina Wasmann, 16 [F]. Type species [automatic]: Rhyzodinamniszechii Chevrolat, 1873, by monotypy. Status: junior synonym of Rhyzodina Chevrolat, 1873 in Tenebrioninae: Rhysopaussini. Note: unjustified emendation of Rhyzodina Chevrolat, 1873, not in prevailing usage.

Rhysopaussus Wasmann, 616 [M]. Type Rhysopaussusdohertyi Wasmann, 1896, by monotypy. Status: valid genus in Tenebrioninae: Rhysopaussini.

Rhytidonota Agassiz, 327 [F]. Type species [automatic]: Rhytinotascabriuscula Eschscholtz, 1831, by subsequent designation (Chevrolat 1848: 279, as “Rytinota”). Status: junior synonym of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: unjustified emendation of Rhytinota Eschscholtz, 1831 (as “Rytinota”), not in prevailing usage.

Rhytinopsis Bouchard & Bousquet, new subgenus [F]. Type Rhytinotafossulata Kraatz, 1880, by present designation. Status: valid subgenus of Thalpophilodes Strand, 1942, in Pimeliinae: Tentyriini. Note: Koch (1943b: 774, 870) introduced the new subgenus name Rhytinopsis for several nominal species, but unfortunately did not designate a type species; the subgenusRhytinopsis, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Rhytinotafossulata Kraatz, 1880 as type species and referring to Koch (1943b: 774, 870) for the character states that characterise and differentiate Rhytinopsis.

Rhytinota Eschscholtz, 5, 7 [F]. Type Rhytinotascabriuscula Eschscholtz, 1831, by subsequent designation (Chevrolat 1848: 279). Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: this name is an unjustified emendation of the original spelling Rytinota, introduced by Chevrolat (1848: 279), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Rhytistena Bouchard & Bousquet, new subgenus [F]. Type Rhytistenagridelli Koch, 1943, by present designation. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: Koch (1943b: 769) introduced the new subgenus name Rhytistena (also spelled Rhytisten on page 888 of the same work) for several nominal species, but unfortunately did not designate a type species; the subgenusRhytistena, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Rhytistenagridelli Koch, 1943 as type species and referring to Koch (1943b: 769) for the character states that characterise and differentiate Rhytistena.

Rhyzodina Chevrolat, 208 [F]. Type Rhyzodinamniszechii Chevrolat, 1873, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Rhysopaussini.

Ripicolodes Koch, 225, 230 [M]. Type Pterostichulamisanthropa Koch, 1952, by original designation. Status: valid subgenus of Pterostichula Koch, 1952 in Pimeliinae: Sepidiini: Oxurina.

Rizalus Mulsant & Rey, 104 [M]. Type Opatrumpiceum G.-A. Olivier, 1812, by monotypy. Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Robustocamaria Pic, 25 [F]. Type Camariafortipes Pic, 1922, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Robustosora Pic, 232 [F]. Type Robustosorafreynei Pic, 1954, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Rondoniella Kaszab, 112 [F]. Type Rondoniellacostata Kaszab, 1970, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Rondoniellina.

Rouyerus Pic, 3 [M]. Type Rouyerusbimaculatus Pic, 1911, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Rozonia Fairmaire, 184 [F]. Type Rozoniastrigicollis Fairmaire, 1888, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Ruandania Pic, 182 [F]. Type Ruandaniarufescens Pic, 1955, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Rues Casey, 66 [M]. Type Helopsovipennis Casey, 1890, by monotypy. Status: junior synonym of Adelium W. Kirby, 1819 in Lagriinae: Adeliini. Synonymy: Spilman (1959: 63). Note: as pointed out by Spilman (1959: 63) the type species, which was erroneously described from the United States of America, is in fact Australiana.

Rugasida Escalera, 65 [F]. Type Asidareticulata Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 291).

Rugoplatynotus Kaszab, 281, 288 [M]. Type Pseudoblapsandrewesii Fairmaire, 1896, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Rugosiheliofugus Freude, 125, 128 [M]. Type Heliophygussulcatulus Gemminger, 1870, by original designation. Status: valid subgenus of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini.

Rhyssochiton Gray in Griffith and Pidgeon, pl. 50 [M]. Type Rhyssochitonpolitus Gray, 1831, by monotypy. Status: junior synonym of Blapida Perty, 1830 in Stenochiinae: Cnodalonini. Synonymy: Lacordaire (1859b: 425). Note: two spellings originally appeared in plates of the same work, Ryssocheton (Gray in Griffith and Pidgeon, pl. 50) and Rhyssochiton (Gray in Griffith and Pidgeon, pl. 69); the third spelling Ryssochiton was also used in the “Index for plates” of Volume 15 of the same work (Gray in Griffith and Pidgeon, 793); we act as First Revisers and select the alternative original spelling Rhyssochiton.

Sabularius Escalera, 353 [M]. Type Helopsfossor Escalera, 1914, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Sabulophosis Penrith, 19, 227 [F]. Type Zophosisgaerdesi Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Saccophorella Strand, 303 [F]. Type species [automatic]: Saccophoruscrenulatus Haag-Rutenberg, 1872, by monotypy. Status: junior synonym of Horatoma Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Note: replacement name for Saccophorus Haag-Rutenberg, 1872.

Saccophorus Haag-Rutenberg, 274, 303 [M]. Type Saccophoruscrenulatus Haag-Rutenberg, 1872, by monotypy. Status: junior synonym of Horatoma Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Synonymy: Penrith and Endrödy-Younga (1994: 10). Note: junior homonym of Saccophorus Kuhl, 1820 [Mammalia].

Sadanaria Ando & Ichiyanagi, 80 [F]. Type Sadanariasakaii Ando & Ichiyanagi, 2009, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Saeculum Kamiński, Kanda & Smith, 196 [N]. Type Saeculumzoologicum Kamiński, Kanda & Smith, 2021, by original designation. Status: valid genus in Pimeliinae: Asidini.

Saerangodes Sturm, 163 [M]. Type Helopsinterpunctatus Germar, 1823, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Blanchard (1845: 33, with Stenochia W. Kirby, 1819, a synonym of Styrongylium W. Kirby, 1819).

Saharoplarion Koch, 431 [N]. Type Microsituscompactus Fairmaire, 1880, by original designation. Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Saitostrongylium Masumoto, 34 [N]. Type Saitostrongyliumacco Masumoto, 1996, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Sakaiomenimus Ando, 135 [M]. Type Sakaiomenimustodai Ando, 2003, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Salax Guérin-Méneville, 11 [M]. Type Salaxlacordairii Guérin-Méneville, 1834, by monotypy. Status: valid genus in Pimeliinae: Trilobocarini.

Saptine Champion, 180 [F]. Type Saptineovata Champion, 1886, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Sarachus Gistel, viii [M]. Type species [automatic]: Adesmialongipes Fischer, 1822 (= Pimeliaanomala Fischer, 1820), by monotypy. Status: junior synonym of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: unnecessary replacement name for Adesmia Fischer, 1822.

Saragella Carter, 136 [F]. Type Saragellapalpalis Carter, 1937, by monotypy. Status: junior synonym of Dysarchus Pascoe, 1866 in Tenebrioninae: Heleini: Heleina. Synonymy: Matthews (1993: 1059).

Saragodinus Bates, 269 [M]. Type Saragodinusduboulayi Bates, 1872, by subsequent designation (R. Lucas 1920: 579). Status: junior synonym of Dysarchus Pascoe, 1866 in Tenebrioninae: Heleini: Heleina. Synonymy: Carter (1921: 307).

Saragus Erichson, 171 [M]. Type fixed herein (ICZN 1999, Article 70.3) as Celibecostata Solier, 1848, misidentified as Silphalaevicollis Fabricius, 1775 in the original designation by monotypy in Erichson (1842a). Status: valid genus in Tenebrioninae: Heleini: Heleina. Note: the type species “Silphalaevicollis Fabricius” was first established by monotypy; Schaum (1850: 183) first noted that Silphalaevicollis Fabricius of Erichson (1842a) was misidentified and corresponded to Celibecostata Solier, 1848 (as “Cilibecostatus”); we follow currently accepted concepts (e.g., Matthews and Bouchard 2008: 310) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Silphalaevicollis Fabricius, 1775 is a valid species in the genus Boreosaragus Matthews, 1993 [Coleoptera: Tenebrionidae].

Sarandonyx Gozis, 151 [M]. Type species [automatic]: Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Westwood 1838: 32). Status: junior synonym of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: unnecessary replacement name for Cteniopus Solier, 1835.

Sarothropus Kraatz, 268 [M]. Type Akisdepressa Zoubkoff, 1837, by monotypy. Status: valid genus in Pimeliinae: Akidini.

Satanocalcar Pic, 9 [N]. Type Satanocalcarcornutum Pic, 1925, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Saxistena Löbl & Merkl in Löbl et al., 2020: 3 [F]. Type Mesostenalongicornis Kraatz, 1865, by original designation. Status: valid subgenus of Mesostena Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: name first proposed by Koch (1940c: 65) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 252) designated Mesostenalongicornis Kraatz, 1865 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Saziches Champion, 261 [M]. Type Sazichessubcaudatus Champion, 1886, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Scaletomerus Blackburn, 330 [M]. Type Scaletomerusharpaloides Blackburn, 1891 (= Cistelapolitus W.J. MacLeay, 1872), by subsequent designation (R. Lucas 1920: 580). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Scaphidema Redtenbacher, 591 [F]. Type Scaphidiumbicolor Fabricius, 1798, by monotypy. Status: valid genus in Diaperinae: Scaphidemini. Note: see Kerzhner (2003: 322) and Löbl and Smetana (2010: 34) for comments on the gender of this name.

Scaphinion Matthews, 2 [M]. Type Scaphinionclavatus Matthews, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Scaptes Champion, 222 [M]. Type Scaptessquamulatus Champion, 1886 (= Asidatropica Kirsch, 1866), by subsequent designation (Bousquet et al. 2018: 194). Status: junior synonym of Ammodonus Mulsant, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Fall (1912: 48).

Scauris Rafinesque, 113 [F]. Type species [automatic]: Scaurusatratus Fabricius, 1775, by monotypy. Status: junior synonym of Scaurus Fabricius, 1775 in Tenebrioninae: Scaurini. Note: unjustified emendation of Scaurus Fabricius, 1775, not in prevailing usage.

Scaurus Fabricius, 253 [M]. Type Scaurusatratus Fabricius, 1775, by monotypy. Status: valid genus in Tenebrioninae: Scaurini.

Scelace Marseul, 325 [F]. Type Pimeliatuberculifera P.H. Lucas, 1858, by subsequent designation (Bouchard and Bousquet 2020b: 7). Status: valid genus in Pimeliinae: Pimeliini. Note: Scelace was published by Marseul on June 9, 1887 while its junior synonym Pachyscelodes was published by Sénac (1887: 189) on August 15 of the same year.

Sceleodis Gemminger in Gemminger and Harold, 1836 [M]. Type species [automatic]: Cratopuscastaneus Eschscholtz, 1831, by monotypy. Status: junior synonym of Scelosodis Solier, 1835 in Pimeliinae: Tentyriini. Note: unjustified emendation of Scelosodis Solier, 1835, not in prevailing usage.

Sceleoides Agassiz, 333 [M]. Type species [automatic]: Cratopuscastaneus Eschscholtz, 1831, by monotypy. Status: junior synonym of Scelosodis Solier, 1835 in Pimeliinae: Tentyriini. Note: unjustified emendation of Scelosodis Solier, 1835, not in prevailing usage.

Scelidospecta Kulzer, 204 [F]. Type Entomodereslobatus Burmeister, 1875, by original designation. Status: valid genus in Pimeliinae: Nycteliini. Note: Scelidospecta is an incorrect subsequent spelling of the original name Scelidopsecta in prevailing usage and attributed to the publication of the original spelling; Scelidospecta is treated as the correct original spelling (ICZN 1999, Article 33.3.1).

Scelocolpis Reitter, 105 [M]. Type Colposcelisdamone Reitter, 1900, by subsequent designation (G.S. Medvedev 1990: 64). Status: valid subgenus of Colposcelis Dejean, 1834 in Pimeliinae: Tentyriini.

Scelosodis Solier, 253, 283 [M]. Type species [automatic]: Cratopuscastaneus Eschscholtz, 1831, by monotypy. Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Cratopus Eschscholtz, 1831.

Schedarosus Reitter, 42 [M]. Type Schedarosuscucujiformis Reitter, 1876 (= Pythopallidus Say, 1823), by subsequent designation (Löbl et al. 2008a: 42). Status: junior synonym of Adelina Dejean, 1835 in Diaperinae: Diaperini: Adelinina. Synonymy: Champion (1886: 157, with Doliema Pascoe, 1860, a junior synonym of Adelina Dejean, 1835).

Schelodontes Koch, 81 [M]. Type Trigonopusimmundus Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Schevodera Borchmann, 16, 59 [F]. Type Lagriahirticollis Borchmann, 1911, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Schevogria Borchmann, 20, 164 [F]. Type Schevogriamethneri Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Schizaraeus Kulzer, 479 [M]. Type Schizaraeusacuticosta Kulzer, 1955, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Schizillus Horn, 33 [M]. Type Schizilluslaticeps Horn, 1874, by monotypy. Status: valid genus in Pimeliinae: Cryptoglossini.

Schizillus Wasmann, 166 [M]. Type Schizillusrogersi Wasmann, 1899, by monotypy. Status: junior synonym of Pseudethas Fairmaire, 1896 in Pimeliinae: Stenosini: Dichillina. Synonymy: Gebien (1937a: 679). Note: junior homonym of Schizillus Horn, 1874 [Coleoptera: Tenebrionidae: Pimeliinae: Cryptoglossini].

Schizomma Gebien, 325, 392 [N]. Type Schizommacucumericola Gebien, 1921, by original designation. Status: senior synonym of Diachoriops Ando, 2020 in Stenochiinae: Cnodalonini. Note: junior homonym of Schizomma Ehrenberg, 1861 [Protista].

Schizophthalmotribolium Kaszab, 173 [N]. Type Schizophthalmotriboliumaustraliae Kaszab, 1940, by original designation. Status: junior synonym of Paratoxicum Champion, 1894 in Tenebrioninae: Tenebrionini. Synonymy: Matthews (2004: 261).

Schlinkus R. Lucas, 584 [M]. Type species [automatic]: Cyphonotusdromedarius Guérin-Méneville, 1831, by monotypy. Status: junior synonym of Homocyrtus Dejean, 1834 in Tenebrionidae: incertae sedis. Synonymy: Gebien (1948: 514). Note: replacement name for Cyphonotus Guérin-Méneville, 1831.

Schoenicus J.L. LeConte, 109 [M]. Type Schoenicuspuberulus J.L. LeConte, 1866, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Schoeniphegoneus Freude, 99 [M]. Type Epitragussemicastaneus Curtis, 1845, by original designation. Status: valid subgenus of Phegoneus Casey, 1907 in Pimeliinae: Epitragini.

Schusteriella Koch, 745, 746 [F]. Type Stenosisruficornis Reitter, 1886, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Schweinfurthia Andres in Kneucker, 26 [F]. Type Schweinfurthiasinaitica Andres, 1922, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Schyzoschelus Koch, 72 [M]. Type Schyzoscheluskaszabi Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Sciaca Solier, 408 [F]. Type Hylithusdisctinctus Solier, 1835, by present designation. Status: junior synonym of Hylithus Guérin-Méneville, 1834 in Pimeliinae: Edrotini. Synonymy: Chevrolat (1848: 423). Note: the name Sciaca was originally listed as synonym of Hylithus Guérin-Méneville, 1834; it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Dejean 1836: 204), Sciaca was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Sciophagus Sharp in Blackburn and Sharp, 167 [M]. Type Ulomapandanicola Boisduval, 1835, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Scleroides Fairmaire, 32 [M]. Type Scleroidespluricostatus Fairmaire, 1883, by monotypy. Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Scleron Hope, 111 [N]. Type Opatrumorientale Fabricius, 1775, by original designation. Status: junior synonym of Sclerum Dejean, 1834 in Blaptinae: Opatrini: Sclerina. Synonymy: Erichson (1842b: 237); Bouchard et al. (2005: 511).

Scleronimon Reitter, 127 [M]. Type Eurycaulusgranulatus Reitter, 1904, by subsequent designation (Löbl and Merkl 2003: 249). Status: junior synonym of Eurycaulus Fairmaire, 1868 in Blaptinae: Opatrini: Sclerina. Synonymy: Grimm (2005: 9).

Scleronopsis Koch, 86, 87 [F]. Type Scleronhirsutus Miller, 1861, by monotypy. Status: junior synonym of Eurycaulus Fairmaire, 1868 in Blaptinae: Opatrini: Sclerina. Synonymy: Grimm (2005: 9).

Scleropatroides Löbl & Merkl, 250 [M]. Type Opatrumhirtulum Baudi di Selve, 1876, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Scleropatrum Reitter, 388 [N]. Type Scleropatrumtuberculatum Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Sclerum Dejean, 193 [N]. Type Opatrumorientale Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: valid genus in Blaptinae: Opatrini: Sclerina.

Scolytocaulus Fairmaire, 98 [M]. Type Scolytocaulusbouchardi Fairmaire, 1896, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Scopulophosis Penrith, 19, 215 [F]. Type Zophosiscacozela Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Scotaeus Hope, 14 [M]. Type Scotaeuscorallipes Hope, 1834, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Scotera Motschulsky, 365 [F]. Type Scoteragibbosa Motschulsky, 1845, by monotypy. Status: junior synonym of Cibdelis Mannerheim, 1843 in Stenochiinae: Cnodalonini. Synonymy: Motschulsky (1845b: 365). Note: as pointed out by Bousquet et al. (2018: 296) the generic name Scotera and the specific name gibbosa, proposed as synonyms of Cibdelisblaschkii by Motschulsky (1845b: 365, 366), are available from Motschulsky’s work (ICZN 1999, Article 11.6.1) because they were treated as valid names before 1961 (e.g., Chevrolat 1848: 454).

Scotinesthes Fairmaire, 19 [F]. Type Scotinesthesacuticosta Fairmaire, 1895, by subsequent designation (Gebien 1937a: 741). Status: valid genus in Pimeliinae: Asidini.

Scotinus W. Kirby, 415 [M]. Type Scotinuscrenicollis W. Kirby, 1819, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Scotobaenus J.L. LeConte, 87 [M]. Type Scotobaenusparallelus J.L. LeConte, 1859, by monotypy. Status: valid genus in Tenebrioninae: Centronopini.

Scotobates Rye, 341 [M]. Type Helopscalcaratus Fabricius, 1798, by subsequent designation (R. Lucas 1920: 587). Status: junior synonym of Menechides Motschulsky, 1872 in Tenebrioninae: Centronopini. Synonymy: R. Lucas (1920: 587). Note: see Spilman (1962b: 2–3) regarding availability of this genus name; Scotobates was used earlier by Horn (1876a: 151) without a description, a definition, or an indication (ICZN 1999, Article 12.1) and is therefore unavailable from that date.

Scotobiopsis Brèthes, 207 [F]. Type Scotobiopsisbreyeri Brèthes, 1910, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Scotobius Germar, 135 [M]. Type Scotobiuspilularius Germar, 1823, by subsequent designation (Lacordaire 1830a: 282). Status: valid genus in Tenebrioninae: Scotobiini.

Scotochares Boheman, 95 [M]. Type Scotocharesinsularis Boheman, 1858, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Scotoderus Perroud & Montrouzier, 114 [M]. Type Scotoderuscancellatus Perroud & Montrouzier, 1865 (= Tenebriocancellatus Montrouzier, 1860), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Scutopiloxys Pic, 13 [M]. Type Scutopiloxysperroti Pic, 1924, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Scymena Pascoe, 455 [F]. Type Scymenavariabilis Pascoe, 1866, by original designation. Status: valid genus in Blaptinae: Opatrini: Heterotarsina.

Scythis Schaum in Kraatz, 80, 102 [F]. Type Tentyriamacrocephala Tauscher, 1812, by subsequent designation (Gebien 1937a: 622). Status: valid genus in Pimeliinae: Tentyriini.

Scythodonta Reitter, 300 [F]. Type Scythishumeridens Reitter, 1887, by monotypy. Status: junior synonym of Scytosoma Reitter, 1895 in Pimeliinae: Tentyriini. Synonymy: Skopin (1979: 171).

Scytosoma Reitter, 281 [N]. Type Scytosomaarcibasis Reitter, 1895 (= Tentyriapygmaea Gebler, 1832), by subsequent designation (Gebien 1937a: 622). Status: valid genus in Pimeliinae: Tentyriini. Note: redescribed as new by Reitter (1897a: 299).

Sebastianus Iwan, 385, 416 [M]. Type Melanocratusmajor Fairmaire, 1899, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Sechuranus Flores & Giraldo-Mendoza in Giraldo-Mendoza and Flores, 93 [M]. Type Prohylithusbarbatus Kaszab, 1964, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Seidlitzellus Reitter, 221 [M]. Type species [automatic]: Anisocerustristis Faldermann, 1837, by monotypy. Status: junior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Note: unnecessary replacement name for Anisocerus Faldermann, 1837, a senior synonym of Ceratanisus Gemminger, 1870.

Seirotrana Pascoe, 483 [F]. Type Adeliumcatenulatum Boisduval, 1835, by original designation. Status: valid genus in Lagriinae: Adeliini.

Selenomma Dejean, 203 [F]. Type species [automatic]: Ammophorusperuvianus Guérin-Méneville, 1831, by monotypy. Status: junior synonym of Ammophorus Guérin-Méneville, 1831 in Tenebrioninae: Scotobiini. Note: unnecessary replacement name for Ammophorus Guérin-Méneville, 1831.

Selenepistoma Dejean, 190 [N]. Type Opatrumacutum Wiedemann, 1823, by subsequent designation (Bousquet and Bouchard 2013a: 50). Status: valid genus in Blaptinae: Dendarini: Melambiina.

Selinopodus Koch, 79 [M]. Type Selinopodusgiganteus Koch, 1956, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Selinus Mulsant & Rey, 70, 97 [M]. Type Opatrumplanum Fabricius, 1792, by subsequent designation (Cazurro Ruiz 1896: 963). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Sellio Mulsant & Rey, 70, 105 [M]. Type Blaps tibidens Quensel, 1806, by subsequent designation (Gebien 1938a: 407). Status: junior synonym of Diastolinus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Blapstinina. Synonymy: Ivie and Hart (2016: 468).

Semenovonymus Bogatchev, 391 [M]. Type Semenovonymustenuis Bogatchev, 1946, by monotypy. Status: junior synonym of Scythis Kraatz, 1865 in Pimeliinae: Tentyriini. Synonymy: G.S. Medvedev (1990: 101).

Semieutochia Kaszab, 187 [F]. Type Semieutochiaooidea Kaszab, 1980, by original designation. Status: valid genus in Tenebrioninae: Ulomini. Note: Semieutochia was used earlier by Kaszab (1979a: 91) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Seorsophloeus Bremer, 10, 12 [M]. Type Corticeusbirmanicus Blair, 1921, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Seorsoplonyx Bremer, 144, 156 [M]. Type Seorsoplonyxantennatus Bremer, 2010, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Sepedonastes Gistel, 382 [M]. Type Tenebriobimaculatus Herbst, 1799 (= Dytiscusbimaculatus Linnaeus, 1767), by subsequent designation (Bouchard et al. 2005: 501). Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Synonymy: Bouchard et al. (2005: 513).

Sepidiacis Fairmaire, cxlvi [F]. Type Sepidiaciscompressa Fairmaire, 1884, by subsequent designation (W.F. Kirby 1885b: 83). Status: junior synonym of Sepidiostenus Fairmaire, 1884 in Pimeliinae: Sepidiini: Sepidiina. Synonymy: Gestro (1892: 775). Note: redescribed as new by Fairmaire (1887a: 185).

Sepidiopsis Gestro, 771 [F]. Type Sepidiopsiscornigera Gestro, 1892, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Sepidiostenus Fairmaire, lxxv [M]. Type Sepidiostenuserinaceus Fairmaire, 1884, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina. Note: redescribed as new, under the spelling Sepidostenus, by Fairmaire (1887a: 184).

Sepidium Fabricius, 250 [N]. Type Sepidium tricuspidatum Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Sepilokus Iwan & Raś, 776 [M]. Type Sepilokustenenbaumi Iwan & Raś, 2020, by original designation. Status: valid genus in Phrenapatinae: Archaeoglenini.

Septentriophosis Penrith, 167 [F]. Type Erodiusplanus Fabricius, 1775, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Seriscius Motschulsky, 77 [M]. Type Serisciuspubescens Motschulsky, 1845 (= Crypticusrufipes Gebler, 1830), by monotypy. Status: valid subgenus of Crypticus Latreille, 1816 in Diaperinae: Crypticini.

Serrania Garrido, 50 [F]. Type Diaperisviridula Zayas, 1988 (= Platydemavirens Laporte & Brullé, 1831), by original designation. Status: junior synonym of Nesocyrtosoma Marcuzzi, 1976 in Stenochiinae: Cnodalonini. Synonymy: Hopp and Ivie (2009: 13).

Serrichora Koch, 15 [F]. Type Eurychoracrenata Solier, 1837, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Serridenos Koch, 325 [M]. Type Zadenossolenopistoma Koch, 1956, by original designation. Status: valid subgenus of Selenepistoma Dejean, 1834 in Blaptinae: Dendarini: Melambiina.

Setenis Motschulsky, 24 [M]. Type Tenebriovalgus Wiedemann, 1823, by original designation. Status: junior synonym of Promethis Pascoe, 1869 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1988: 70).

Seydelicistela Pic, 260 [F]. Type Seydelicistelarubrithorax Pic, 1954, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Sicharbas Champion, 67 [M]. Type Sicharbaslobatus Champion, 1884, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Sicinus Champion, 146 [M]. Type Sicinusguatemalensis Champion, 1886, by subsequent designation (Löbl et al. 2008a: 43). Status: junior synonym of Gnatocerus Thunberg, 1814 in Diaperinae: Diaperini: Adelinina. Synonymy: Leng (1920: 233).

Silvestriellum Koch, 362 [N]. Type Silvestriellumalatum Koch, 1956, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Simalura Gebien, 71 [F]. Type Simalurajacobsoni Gebien, 1914, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Simarus Borchmann, 713 [M]. Type species [automatic]: Ismarusgodeffroyi Haag-Rutenberg, 1878, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: replacement name for Ismarus Haag-Rutenberg, 1878.

Similepitragus Freude, 148, 167 [M]. Type Epitragussimilis Steinheil, 1872, by original designation. Status: valid subgenus of Epitragus Latreille, 1802 in Pimeliinae: Epitragini.

Singapura Gebien, 325 [F]. Type Singapuraquadrihamata Gebien, 1925, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Sinocatomus Nabozhenko, 852 [M]. Type Catomussolitarius Nabozhenko, 2006, by original designation. Status: valid subgenus of Catomus Allard, 1876 in Tenebrioninae: Helopini: Helopina.

†Sinocistela Zhang, 145 [F]. Type Sinocistelasiphla Zhang, 1989, by original designation. Status: valid genus in Alleculinae: Cteniopodini. Note: described from Lower Miocene deposits (China).

Sinoecia Chatanay, 220 [F]. Type Sinoeciapuncticollis Chatanay, 1914, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Sinomenimus G.S. Medvedev, 675 [M]. Type Menimuskabaki G.S. Medvedev, 2007, by original designation. Status: valid subgenus of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina.

Sinopium Pascoe, 487 [N]. Type Strongyliumvariabile Walker, 1858, by original designation. Status: junior synonym of Camarimena Motschulsky, 1863 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1919: 155).

Sinorus Mulsant & Revelière, 153 [M]. Type Sinorusciliaris Mulsant & Revelière, 1861 (= Opatrumcolliardi Fairmaire, 1860), by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Sintagona G.S. Medvedev, 585 [F]. Type Sintagonamiranda G.S. Medvedev, 1998, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Sipirocus Fairmaire, 103 [M]. Type Sipirocusritsemae Fairmaire, 1896, by monotypy. Status: junior synonym of Aediatorix Bates, 1868 in Lagriinae: Pycnocerini. Synonymy: Gebien (1905: 258).

Sipolisia Fairmaire, xlix [F]. Type Sipolisiaserricornis Fairmaire, 1889, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Sitophagus Mulsant, 264 [M]. Type Sitophagussolieri Mulsant, 1854 (= Ulomahololeptoides Laporte, 1840), by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Sloanea Carter, 209 [F]. Type Sloaneacostata Carter, 1916, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Asphalina.

Smiliophanus Koch, 66 [M]. Type species [automatic]: Smiliotussteiroides Haag-Rutenberg, 1875, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Smiliotus Haag-Rutenberg, 1875.

Smiliotus Haag-Rutenberg, 4, 52 [M]. Type Smiliotussteiroides Haag-Rutenberg, 1875, by monotypy. Status: senior synonym of Smiliophanus Koch, 1950 in Pimeliinae: Adelostomini. Note: junior homonym of Smiliotus Loew, 1857 [Diptera].

Sobas Pascoe, 45 [F]. Type species [automatic]: Trigonotarsusaustralis Hope, 1843, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: replacement name for Trigonotarsus Hope, 1843. Note: moved from the subtribe Ammobiina to Opatrina by Kamiński et al. (2021b: 151).

Socotralia Novák, 321 [F]. Type Socotraliamajor Novák, 2007, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Socotraphanes Nabozhenko in Nabozhenko and Purchart, 150 [M]. Type Socotraphaneskrali Nabozhenko, 2019, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Socotropatrum Koch, 109 [N]. Type Opatrumcostiferum C.O. Waterhouse, 1881, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Soemias Champion, 4 [F]. Type Soemiasminuta Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Solenomerus Fåhraeus, 306 [M]. Type Solenomeruslongipes Fåhraeus, 1870, by monotypy. Status: junior synonym of Micrantereus Solier, 1848 in Blaptinae: Pedinini: Helopinina. Synonymy: Fairmaire (1897f: 131).

Solskyia Solsky, 48 [F]. Type Solskyiaperegrina Solsky, 1881, by monotypy. Status: valid genus in Pimeliinae: Akidini.

Somaladesmia Koch, 147 [F]. Type Adesmiaconsimilis Gahan, 1896, by monotypy. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Somalammodes Koch, 500, 510 [M]. Type Somalammodesdelaruei Koch, 1943, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Somalarabes Koch, 155 [M]. Type Psammodesgracilentus Fairmaire, 1882, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Somaticus Hope, 117 [M]. Type Sepidiumrugosum Fabricius, 1781, by original designation. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Trachynotina. Note: Somaticus, introduced by Lacordaire (1859a: 197), is an incorrect subsequent spelling of the original name Somaticum in prevailing usage and attributed to the publication of the original spelling; we follow Kamiński et al. (2019b: 86) and treat Somaticus as the correct original spelling (ICZN 1999, Article 33.3.1); the First Reviser (Somaticus Hope, 1841 versus Tracheloeum Hope, 1841) is Koch (1955a: 53).

Somocoelia Heyden & Kraatz, 331 [F]. Type Somocoeliapinguis Heyden & Kraatz, 1882, by monotypy. Status: valid genus in Blaptinae: Platyscelidini.

Somocoeloplatys Skopin, 82 [M]. Type Platynoscelisboroldaica Skopin, 1965, by original designation. Status: valid genus in Blaptinae: Platyscelidini.

Sophrobates Fairmaire, xxxvi [M]. Type Sophrobatesarcadii Fairmaire, 1889, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Sora Walker, 259 [F]. Type Soramarginata Walker, 1859, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Soradeus Rafinesque, 114 [M]. Type species [automatic]: Heleaperforata Latreille, 1816, by subsequent monotypy (Latreille 1816b: 261). Status: junior synonym of Helea Latreille, 1804 in Tenebrioninae: Heleini: Heleina. Note: unnecessary replacement name for Helea Latreille, 1804 (as “Heleus Latr.”).

Spathulipezus Gebien, 398, 458 [M]. Type Spathulipezusmiritaris Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Spectrocnera Kwieton, 402 [F]. Type Spectrocneraanguliceps Kwieton, 1981, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Spelaebiosis Bousquet & Bouchard in Bousquet et al., 2018: 223 [F]. Type species [automatic]: Orghidaniatorrei Ardoin, 1977, by monotypy. Status: valid genus in Tenebrioninae: Triboliini. Note: replacement name for Ardoinia Özdikmen, 2005.

Sphaeriontis Casey, 56, 75 [F]. Type Eusattusmuricatus J.L. LeConte, 1851, by original designation. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: La Rivers (1949: 180).

Sphaerocaulus Fairmaire, 235 [M]. Type Sphaerocaulusgraniger Fairmaire, 1869, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Sphaerognathium Dajoz, 112 [N]. Type Sphaerognathiumglobosum Dajoz, 1975, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Sphaeromatris Fairmaire, 535 [F]. Type Sphaeromatrisaurovittata Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Sphaerostibes Koch, 64 [M]. Type Sphaerostibessabulicola Koch, 1963, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 13.4).

Sphaerotidius Kaszab, 3, 38 [M]. Type Sphaerotidiusduplicatus Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Spaerotidius, used by Kaszab (1941a: 39, 40).

Sphaerotus W. Kirby, 416 [M]. Type Sphaerotuscurvipes W. Kirby, 1819, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Sphargeris Pascoe, 122 [M]. Type Sphargerisphysodes Pascoe, 1860, by monotypy. Status: valid genus in Lagriinae: Chaerodini.

Sphenaria Ménétriés, 240 [F]. Type Sphenariaelongata Ménétriés, 1849, by subsequent designation (Gebien 1937a: 576). Status: valid genus in Pimeliinae: Tentyriini.

Sphenariopsis Kraatz, 80, 175 [F]. Type Sphenariopsistristis Kraatz, 1865, by monotypy. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Spheneuphloeus Kaszab, 5, 35 [M]. Type Osdarametallica Pic, 1931, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Spheniscus W. Kirby, 421 [M]. Type Spheniscuserotyloides W. Kirby, 1819, by monotypy. Status: senior synonym of Cuphotes Champion, 1887 in Stenochiinae: Stenochiini. Note: junior homonym of Spheniscus Moehring, 1758 [Aves].

Sphenogenius Solier, 154, 351 [M]. Type Sphenogeniusclathratus Solier, 1848 (= Nyctozoilusobesus Guérin-Méneville, 1831), by original designation. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Lacordaire (1859a: 350).

Sphenolampidius Kaszab, 4, 40 [M]. Type Sphenolampidiushemisphaericus Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Sphenosdara Kaszab, 2, 28 [F]. Type Sphenosdarasachtlebeni Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Sphenosoma Dejean, 212 [N]. Type species [automatic]: Acropteronrufipes Perty, 1832, by subsequent designation (Hope 1841: 133). Status: junior synonym of Acropteryx Gistel, 1831 in Tenebrioninae: Acropteronini. Note: unnecessary replacement name for Acropteron Perty, 1832.

Sphenothorax Gebien, 232 [M]. Type Tenebrionitidulus Fabricius, 1801, by monotypy. Status: junior synonym of Zophophilus Fairmaire, 1881 in Stenochiinae: Cnodalonini. Synonymy: Carter (1930: 547).

Sphinctoderus Fairmaire, 301 [M]. Type Sphinctoderusstrangulatus Fairmaire, 1903, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Sphingocorse Gebien, 110, 111 [F]. Type Sphingocorseangulicollis Gebien, 1921, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Sphragidophorus Champion, 61 [M]. Type Statiracyanipennis Mäklin, 1863, by subsequent designation (R. Lucas 1920: 603). Status: valid genus in Lagriinae: Lagriini: Statirina.

Spiloscapha Bates, 202 [F]. Type Spiloscaphacrassicornis Bates, 1873 (= Platydemathallioides Pascoe, 1869), by monotypy. Status: valid genus in Diaperinae: Scaphidemini.

Spinadaenus Pic, 18 [M]. Type Spinadaenussingularis Pic, 1921, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Spinamarygmus Pic, 7 [M]. Type Spinamarygmusindicus Pic, 1915, by monotypy. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Spinanemia Löbl, Bouchard, Merkl & Bousquet, 2 [F]. Type Anemiacornuta Pic, 1898, by original designation. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini. Note: subgenus first proposed by Ardoin (1971: 362, 378) without type species originally designated.

Spinecula Novák, 437 [F]. Type Spineculahouaphanica Novák, 2019, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Spinepicalla Pic, 21 [F]. Type Spinepicallaarmata Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Spinoderosphaerus Pic, 26 [M]. Type Spinoderosphaerusbrevicornis Pic, 1922, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: the First Reviser (Spinoderosphaerus Pic, 1922 versus Spinogauromaia Pic, 1922) is Kaszab (1983a: 132).

Spinodietysus Pic, 21 [M]. Type Cyriogetonconvexipennis Pic, 1927, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Spinogauromaia Pic, 23 [F]. Type Spinogauromaiarufescens Pic, 1922 (= Spinoderosphaerusbrevicornis Pic, 1922), by monotypy. Status: junior synonym of Spinoderosphaerus Pic, 1922 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 132).

Spinolagriella Pic, 183 [F]. Type Spinolagriellaminutissima Pic, 1955, by original designation. Status: valid genus in Lagriinae: Lupropini.

Spinolyprops Pic, 12 [M]. Type Spinolypropsrufithorax Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Spinoodescelis Kaszab, 938, 966 [F]. Type Platyscelissomocoeloides Seidlitz, 1893, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Spinophrynus Koch, 90 [M]. Type Phrynocolusspinipennis Gebien, 1910, by original designation. Status: valid subgenus of Phrynocolus Lacordaire, 1859 in Pimeliinae: Sepidiini: Molurina.

Spinorhacus Kaszab, 262 [M]. Type Spinorhacusbaloghi Kaszab, 1969, by original designation. Status: junior synonym of Spinolagriella Pic, 1955 in Lagriinae: Lupropini. Synonymy: Kaszab (1976: 453).

Spinosdara Bouchard & Bousquet, new subgenus [F]. Type Osdarabiroi Kaszab, 1939, by present designation. Status: valid subgenus of Osdara Walker, 1858 in Stenochiinae: Cnodalonini. Note: Kaszab (1941a: 33) introduced the new subgenus name Spinosdara for three nominal species, but unfortunately did not designate a type species; the subgenusSpinosdara, which has been treated as valid since 1941, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Osdarabiroi Kaszab, 1939 as type species and referring to Kaszab (1941a: 33) for the character states that characterise and differentiate Spinosdara.

Spinostatira Pic, 22 [F]. Type Statiraspinipes Pic, 1918, by subsequent designation (Borchmann 1936: 247). Status: valid subgenus of Statira Lepeletier & Audinet-Serville, 1828 in Lagriinae: Lagriini: Statirina.

Splenoodescelis Egorov, 593 [F]. Type Platyscelisturkestanica Seidlitz, 1893, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Splichalia Reitter, 664 [F]. Type Splichaliatigrinella Reitter, 1913, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Spongesmia Bouchard & Bousquet, new subgenus [F]. Type Adesmianassata Erichson, 1843, by present designation. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: Koch (1944b: 149) introduced the new subgenus name Spongesmia for four nominal species, but unfortunately did not designate a type species; the subgenusSpongesmia, which has been treated as valid since 1944, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Adesmianassata Erichson, 1843 as type species and referring to Koch (1944b: 149) for the character states that characterise and differentiate Spongesmia.

Spongesmima Koch, 157 [F]. Type Adesmiascrobipennis Haag-Rutenberg, 1875, by monotypy. Status: junior synonym of Renatiella Koch, 1944 in Pimeliinae: Adesmiini. Synonymy: Penrith (1979: 27).

Spyrathus Kraatz, 6, 9 [M]. Type Spyrathusindicus Kraatz, 1865, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Srilanka Kaszab, 319 [F]. Type Srilankamirabilis Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: Srilanka was used earlier by Kaszab (1979a: 107) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Staius Fairmaire, 359 [M]. Type Staiusmiricornis Fairmaire, 1896, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Stalagmoptera Solsky, 286 [F]. Type Stalagmopteratuberculatocostata Solsky, 1876, by subsequent designation (Gebien 1937a: 821). Status: valid genus in Pimeliinae: Pimeliini.

Statira Lepeletier & Audinet-Serville, 479 [F]. Type Statiraagroides Lepeletier & Audinet-Serville, 1828, by subsequent designation (Blanchard 1844: pl. 53bis). Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Statiropsis Borchmann, 389 [F]. Type Statiropsisaenea Borchmann, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Stegastopsis Kraatz, 80, 176 [F]. Type Stegastopsisbabylonica Kraatz, 1865, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Steira Westwood, pl. 176 [F]. Type Steiracostata Westwood, 1837, by monotypy. Status: senior synonym of Stips Koch, 1950 in Pimeliinae: Adelostomini. Note: junior homonym of Steira Eschscholtz, 1825 [Mollusca].

Stemmatoderus Agassiz, 351 [M]. Type species [automatic]: Stemmoderussingularis Spinola, 1842, by monotypy. Status: junior synonym of Stemmoderus Spinola, 1842 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Stemmoderus Spinola, 1842, not in prevailing usage.

Stemmoderus Spinola, pl. 91 (p. 1) [M]. Type Stemmoderussingularis Spinola, 1842, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Stenadelium Watt, 32 [N]. Type Stenadeliumstriatum Watt, 1992, by original designation. Status: valid genus in Lagriinae: Adeliini.

Stene Stephens, 19 [F]. Type fixed herein (ICZN 1999, Article 70.3) as Colydiumcastaneum Herbst, 1797, misidentified as Tenebrioferrugineus Fabricius, 1781 in the original designation by monotypy in Stephens (1829). Status: junior synonym of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini. Synonymy: Shuckard (1840: vii). Note: the type species “Tenebrioferrugineus Fabricius” was first established by monotypy; as noted by C.O. Waterhouse (1896: 230) and Blair (1913: 223) the Tenebrioferrugineus Fabricius of authors, including Stephens (1829: 19, as “ferruginea, Oliv.”), was misidentified; Blair (1913: 223) noted that the species the authors referred to is in fact Colydiumcastaneum Herbst, 1797; we follow currently accepted concepts (e.g., Bousquet et al. 2018: 224) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Tenebrioferrugineus Fabricius, 1781 is a valid species in the genus Tribolioides Blair, 1913 [Coleoptera: Cucujidae].

Steneleodes Blaisdell, 409 [M]. Type Eleodeslongicollis J.L. LeConte, 1851, by subsequent designation (Bousquet et al. 2018: 166). Status: valid subgenus in Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Stenerophlina Reitter, 130 [F]. Type Omophlinahauseri Reitter, 1894, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Stenerula Fairmaire, 42 [F]. Type Stenerulasubopaca Fairmaire, 1875, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Steneryx Reitter, 256 [M]. Type Cisteladejeanii Faldermann, 1836, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Stenethmus Gebien, 41 [M]. Type Psammodestentyriniformis Hesse, 1935, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Steneucyrtus Fairmaire, 31 [M]. Type Steneucyrtuspexicollis Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stenholma Solier, 253, 412 [F]. Type Stenholmatentyrioides Solier, 1835, by monotypy. Status: junior synonym of Melaphorus Guérin-Méneville, 1834 in Pimeliinae: Evaniosomini. Synonymy: Solier (1835b: 412).

Stenillus Blair, 245 [M]. Type Stenillusmonticola Blair, 1927, by monotypy. Status: valid subgenus of Pseudethas Fairmaire, 1896 in Pimeliinae: Stenosini: Dichillina.

Stenocara Solier, 512, 553 [N]. Type Pimelialongipes Fabricius, 1775, by subsequent designation (Desmarest 141). Status: valid genus and subgenus in Pimeliinae: Adesmiini.

Stenocephalus Agassiz, 71, 351 [M]. Type species [automatic]: Cephalostenusdejeanii Solier, 1838 (= Scauruselegans Brullé, 1832), by subsequent designation (Hope 1841: 115). Status: junior synonym of Cephalostenus Solier, 1838 in Tenebrioninae: Scaurini. Note: unjustified emendation of Cephalostenus Solier, 1838; junior homonym of Stenocephalus Tschudi, 1838 [Amphibia].

Stenocera Agassiz, 74, 351 [F]. Type species [automatic]: Cerostenadeplanata Solier, 1836, by present designation. Status: junior synonym of Psectrascelis Solier, 1836 in Pimeliinae: Nycteliini. Note: unjustified emendation of Cerostena Solier, 1836, not in prevailing usage; junior homonym of Stenocera Brullé, 1834 [Coleoptera: Carabidae].

Stenochia W. Kirby, 423 [F]. Type Stenochiarufipes W. Kirby, 1819, by subsequent designation (Hope 1841: 133). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Latreille (1829b: 683).

Stenochidus J.L. LeConte, 244 [M]. Type Stenochiagracilis J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 608). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Stenochinus Motschulsky, 102 [M]. Type Stenochinusreticulatus Motschulsky, 1860, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stenodesia Reitter, 4 [F]. Type Stenocaraglobulum Haag-Rutenberg, 1875, by monotypy. Status: valid genus in Pimeliinae: Adesmiini.

Stenogena Fairmaire, 33 [F]. Type Stenogenamadecassa Fairmaire, 1895, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: placed in the subfamily Alleculinae by Chatanay (1915a: 526).

Stenogenomorpha Pic, 5 [F]. Type Stenogenomorphaimpressa Pic, 1919, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Stenogonopus Gebien, 91 [M]. Type Tenebrioplumosus Thunberg, 1787, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Stenohelops Reitter, 22 [M]. Type Isopedusplicatulus Kraatz, 1880, by subsequent designation (Gebien 1943: 423). Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina. Note: the First Revisers (Stenohelops Reitter, 1922 versus Gunarellus Reitter, 1922) are Nabozhenko et al. (2020b: 297).

Stenolagria Merkl, 124, 157 [F]. Type Stenolagriamatthewsi Merkl, 1987, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Stenolamus Gebien, 107 [M]. Type Stenolamussulciceps Gebien, 1920, by subsequent designation (Gebien 1938a: 397). Status: valid genus in Blaptinae: incertae sedis. Note: this taxon is placed in “Blaptinae incertae sedis” based on comments in Kamiński et al. (2021b: 150).

†Stenolassus Nabozhenko, Chigray & Bukejs, 519 [M]. Type Nalassusklebsi Nabozhenko, Perkovsky & Chernei, 2016, by original designation. Status: valid subgenus of Stenohelops Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Note: described from Eocene Baltic amber.

Stenomacidius Seidlitz, 791 [M]. Type Hedyphanesacutangulus Seidlitz, 1895, by subsequent designation (G.S. Medvedev 1990: 242). Status: junior synonym of Cylindrinotus Faldermann, 1837 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: Nabozhenko (2006: 799).

Stenomaleis Español, 21, 34 [M]. Type Stenohelopsardoini Español, 1957 (= Helopsprotensulus Seidlitz, 1896), by original designation. Status: junior synonym of Helopelius Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko et al. (2020b: 293).

Stenomax Allard, 4 [M]. Type Tenebriolanipes Linnaeus, 1771 (= Tenebrioaeneus Scopoli, 1763), by subsequent designation (Español and Comas 1989: 166, 170). Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina.

Stenomorpha Solier, 407, 487 [F]. Type Stenomorphablapsoides Solier, 1836, by subsequent designation (Desmarest 150). Status: valid genus and subgenus in Pimeliinae: Asidini.

Stenopalorus Blair, 135 [M]. Type Palorushypophloeoides Blair, 1930, by monotypy. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Halstead (1967a: 114, 115).

Stenophanes Solsky, 294 [M]. Type Hedyphanesmesostenus Solsky, 1871, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Stenophloeus Blair, 1 [M]. Type Hypophlaeusfilum Fairmaire, 1893, by subsequent designation (Löbl et al. 2008b: 312). Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Stenopsis Rafinesque, 113 [F]. Type species [automatic]: Pimeliareflexa Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Akis Herbst, 1799 in Pimeliinae: Akidini. Note: unnecessary replacement name for Akis Herbst, 1799.

Stenoscapha Bates, 237 [F]. Type Stenoscaphatibialis Bates, 1873, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Stenoscapha Fairmaire, 234 [F]. Type Stenoscaphaspissicornis Fairmaire, 1885, by monotypy. Status: senior synonym of Brachypophlaeus Fairmaire, 1897 in Tenebrioninae: Ulomini. Note: junior homonym of Stenoscapha Bates, 1873 [Coleoptera: Tenebrionidae: Diaperinae: Diaperini: Diaperina].

Stenosethas Kaszab, 11 [M]. Type Stenosethascarinipennis Kaszab, 1975, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Stenosida Solier, 253, 281 [F]. Type Stenosidatenuicollis Solier, 1835 (= Tageniastriatopunctata Wiedemann, 1821), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Stenosides Solier, 406, 484 [M]. Type Stenosidesgraciliformis Solier, 1836, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Stenosidops Koch, 733 [M]. Type Tageniasabulosa Guérin-Méneville, 1849, by monotypy. Status: valid subgenus of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina.

Stenosis Herbst, 160 [F]. Type fixed herein (ICZN 1999, Article 70.3) as Tageniaintermedia Solier, 1838, misidentified as Pimeliaangustata Fabricius, 1775 in the subsequent designation in Lacordaire (1859a). Status: valid genus and subgenus in Pimeliinae: Stenosini: Stenosina. Note: the type species “Pimeliaangustata Fabricius” (as angustata Herbst) was selected by Lacordaire (1859a: 102) as the type species; Baudi de Selve (1875: 67) first noted that Pimeliaangustata Fabricius of Herbst (1799) was misidentified and corresponded to Tageniaintermedia Solier, 1838; we follow currently accepted concepts (e.g., Löbl et al. 2008b: 179) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Pimeliaangustata Fabricius, 1775 is a valid species in the genus Mesostena Eschscholtz, 1831 [Coleoptera: Tenebrionidae].

Stenothesilea Kulzer, 131, 134 [F]. Type Stenothesileacylindriformis Kulzer, 1951, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stenotrichus J.L. LeConte, 239 [M]. Type Amphidorarufipes J.L. LeConte, 1851, by original designation. Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Aalbu et al. (2002: 496).

Steptochora Koch, 10 [F]. Type Eurychorasetosula Fairmaire, 1887, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Sterces Champion, 640 [M]. Type Stercesviolaceipennis Champion, 1891, by present designation. Status: junior synonym of Alcyonotus Pascoe, 1882 in Stenochiinae: Cnodalonini. Synonymy: Champion (1894b: lxiii).

Steriphanides Casey, 515 [M]. Type Emmenastusstolidus Champion, 1892, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Steriphanus Casey, 289 [M]. Type Emmenastusconicicollis Casey, 1890, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Sternocnera Skopin, 394 [F]. Type Sternocneralindti Skopin, 1964, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Sternodes Fischer von Waldheim, 10 [M]. Type Sternodeskarelini Fischer von Waldheim, 1837 (= Tenebriocaspicus Pallas, 1781), by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Sternomaia Kulzer, 731 [F]. Type Sternomaiacoeruleovirens Kulzer, 1952, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Sternoplax Frivaldszky, 207 [F]. Type Trigonoscelisszechenyii Frivaldszky, 1890, by monotypy. Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Sternotrigon Skopin, 109, 110 [M]. Type Trigonoscelissetosa Bates, 1879, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Stethasida Casey, 78, 203 [F]. Type Pelecyphorusmuricatulus J.L. LeConte, 1851, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Stethotrypes Gebien, 26 [M]. Type Stethotrypesbicornutus Gebien, 1914, by subsequent designation (Gebien 1940: 431). Status: valid genus in Diaperinae: Leiochrinini.

Sthenoboea Champion, 112 [F]. Type Sthenoboeaapicalis Champion, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stibia Horn, 258, 260 [F]. Type Stibiapuncticollis Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Stictodera Casey, 289, 352 [F]. Type Emmenastuspinguis J.L. LeConte, 1866, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Stictodere Gebien, 101 [F]. Type Stictoderesubseriata Gebien, 1928, by subsequent designation (Gebien 1937a: 571). Status: valid genus in Pimeliinae: Epitragini.

Stictoderia Gebien, 571 [F]. Type species [automatic]: Stictoderesubseriata Gebien, 1928, by subsequent designation (Gebien 1937a: 571). Status: junior synonym of Stictodere Gebien, 1928 in Pimeliinae: Epitragini. Note: unnecessary replacement name for Stictodere Gebien, 1928, which is not a homonym of Stictodera Casey, 1907 [Coleoptera: Tenebrionidae: Pimeliinae: Edrotini].

Stierlinius Forel, 167 [M]. Type species [automatic]: Dolichoderusacuminatus Klug, 1833, by monotypy. Status: junior synonym of Macellocerus Solier, 1848 in Tenebrioninae: Toxicini: Nycteropina. Note: replacement name for Dolichoderus Klug, 1833.

Stigmatoma J.L. LeConte, 244 [F]. Type Cistelafraterna Say, 1824, by monotypy. Status: junior synonym of Ernocharis C.G. Thomson, 1859 in Alleculinae: Alleculini: Mycetocharina. Synonymy: Seidlitz (1896: 147).

Stilbocistela Borchmann, 319 [F]. Type Stilbocistelaluzonica Borchmann, 1932, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: genus described as new again by Borchmann (1932b: 125).

Stips Koch, 66 [F]. Type species [automatic]: Steiracostata Westwood, 1837, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Steira Westwood, 1837.

Stipsostoma Koch, 34, 105 [N]. Type Steirasculpta Gebien, 1920, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Stira Agassiz, 350, 353 [F]. Type species [automatic]: Steiracostata Westwood, 1837, by monotypy. Status: senior synonym of Stips Koch, 1950 in Pimeliinae: Adelostomini. Note: unjustified emendation of Steira Westwood, 1837 (and Steira Eschscholtz, 1825 in Mollusca), not in prevailing usage; we act as First Revisers and treat Stira Agassiz, 1846 [Coleoptera] as a junior homonym of Stira Agassiz, 1846 [Mollusca].

Stizopus Erichson, 245 [M]. Type Stizopuslaticollis Erichson, 1843, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Stomion G.R. Waterhouse, 27 [N]. Type Stomiongalapagoense G.R. Waterhouse, 1845, by subsequent designation (Gebien 1937a: 590). Status: valid genus in Pimeliinae: Edrotini.

Stomium Agassiz, 354 [N]. Type species [automatic]: Stomiongalapagoense G.R. Waterhouse, 1845, by subsequent designation (Gebien 1937a: 590). Status: junior synonym of Stomion G.R. Waterhouse, 1845 in Pimeliinae: Edrotini. Note: unjustified emendation of Stomion G.R. Waterhouse, 1845, not in prevailing usage.

Stomylus Fåhraeus, 300 [M]. Type Stomylusbicolor Fåhraeus, 1870, by subsequent designation (R. Lucas 1920: 616). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Stonavus Reitter, 161 [M]. Type Penthicusalaiensis Reitter, 1896, by subsequent designation (Iwan and Löbl 2007: 734). Status: junior synonym of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Synonymy: Iwan and Löbl (2007: 734).

Storthephora Mäklin, 658 [F]. Type Storthephoradenticollis Mäklin, 1875, by subsequent designation (Bousquet and Bouchard 2014: 26). Status: junior synonym of Paratenetus Spinola, 1845 in Lagriinae: Goniaderini. Synonymy: Champion (1893b: 47).

Storthocnemis Karsch, 47 [F]. Type Storthocnemissteckeri Karsch, 1881, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Stratodemus Gebien, 98 [M]. Type Stratodemusheraldicus Gebien, 1921, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Strepsius Fairmaire, 351 [M]. Type Strepsiusspretulus Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stridigula Koch, 60 [F]. Type Stridigulaarguta Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Stridulomus Koch, 37 [M]. Type Psammodessulcicollis Péringuey, 1885, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Strongyallecula Pic, 186 [F]. Type Strongyalleculabasilewskyi Pic, 1955, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Strongylacanthus Brèthes, 13 [M]. Type Strongylacanthusperuvianus Brèthes, 1925, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Strongylagria Pic, 4 [F]. Type Strongylagriametallica Pic, 1915, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Strongyliastrum Fairmaire, 39 [N]. Type Strongyliastrumbraetii Fairmaire, 1894 (= Strongyliumrufipenne Kollar & Redtenbacher, 1844), by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Strongylium W. Kirby, 417 [N]. Type Strongyliumchalconatum W. Kirby, 1819, by monotypy. Status: valid genus and subgenus in Stenochiinae: Stenochiini. Note: nomen protectum (see Bouchard et al. 2005: 501); the First Reviser (Strongylium W. Kirby, 1819 versus Stenochia W. Kirby, 1819) is Latreille (1829b: 683); we recognize two valid subgenera in the diverse genus Strongylium W. Kirby, 1819, i.e., the recently described Afrostrongylium Robiche, 2019 and the nominotypical subgenus; however, much taxonomic research is needed to evaluate the possible validity of the many synonyms of the subgenusStrongylium W. Kirby, 1819.

Strophia Robiche, 130 [F]. Type Oncosomaertli Gebien, 1910, by original designation. Status: senior synonym of Strophiamixa Robiche, 2005 in Blaptinae: Pedinini: Helopinina. Note: junior homonym of Strophia Meigen, 1832 [Lepidoptera] and Strophia Albers, 1850 [Mollusca].

Strophiamixa Robiche, 358 [F]. Type species [automatic]: Oncosomaertli Gebien, 1910, by original designation. Status: valid subgenus of Amatodes Dejean, 1834 in Blaptinae: Pedinini: Helopinina. Note: replacement name for Strophia Robiche, 2004.

Stygohelops Leo & Liberto, 299 [M]. Type Probaticuskalavriticus Schawaller, 2001, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Styphacus Fairmaire, 71 [M]. Type Styphacusdecorsii Fairmaire, 1901, by subsequent designation (R. Lucas 1920: 619). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Styphloeus Kaszab, 4, 36 [M]. Type Styphloeusindicus Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Styrax Westwood, 227 [M]. Type Styraxtricondyloides Westwood, 1875, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Styrus Bates, 348 [M]. Type Styruselongatulus Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Suarezius Fairmaire, 22 [M]. Type Suareziusgibbosulus Fairmaire, 1895, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Subalphasida Escalera, 136 [F]. Type Asidaluctuosa Boisduval, 1835, by original designation. Status: junior synonym of Betasida Reitter, 1917 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Subpterocoma Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Pterocomatuberculata Motschulsky, 1845, by subsequent designation (Skopin 1974b: 159). Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini. Note: replacement name for Pseudopimelia Motschulsky, 1860.

Sulcipectus Louw, 99, 109 [N]. Type Sulcipectusleve Louw, 1979, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Sulcolagria Pic, 181 [F]. Type Sulcolagriasemiopaca Pic, 1955, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Sulcosis Penrith, 19, 220 [F]. Type Zophosisangolensis Erichson, 1843, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Sulpius Fairmaire, 273 [M]. Type Sulpiuspunctostriatus Fairmaire, 1906, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Sulpiusoma Ferrer, 81 [N]. Type Sulpiusomamedquisti Ferrer, 2006, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Sulpiosoma, used by Ferrer (2006c: 79).

Sumbawia Gebien, 441 [F]. Type Sumbawiatetrops Gebien, 1925, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Sundon Pic, 18 [N]. Type Sundonatricorne Pic, 1923, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Syachis Bates, 467 [M]. Type Syachishimalaicus Bates, 1879, by subsequent designation (Gebien 1937a: 594). Status: valid genus in Pimeliinae: Tentyriini. Note: redescribed as new by Bates (1890: 55).

Sycophantes Kirsch, 198 [M]. Type Sycophantesdentipes Kirsch, 1866, by subsequent designation (Gebien 1928: 203). Status: valid genus in Stenochiinae: Cnodalonini.

Sycophantomorphus Pic, 13 [M]. Type Sycophantomorphusater Pic, 1924, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Syggona Fåhraeus, 330 [F]. Type Syggonaconcinna Fåhraeus, 1870, by monotypy. Status: junior synonym of Luprops Hope, 1833 in Lagriinae: Lupropini. Synonymy: Ferrer (1995b: 62).

Sylvanoplonyx Bremer, 156, 157 [M]. Type Sylvanoplonyxfemoralis Bremer, 2010, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Sympetes Pascoe, 464 [M]. Type Sympetesmacleayi Pascoe, 1866 (= Emcephalustricostellus White, 1841), by subsequent designation (Gebien 1940: 1075). Status: valid genus in Tenebrioninae: Heleini: Heleina.

Symphochora Koch, 12 [F]. Type Eurychorahumerifera Gebien, 1937, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Symphoxycara Koch, 577 [N]. Type Oxycarabreviusculum Fairmaire, 1892, by monotypy. Status: valid subgenus of Oxycara Solier, 1835 in Pimeliinae: Tentyriini.

Sympiezocnemis Solsky, 290 [F]. Type Sympiezocnemiskessleri Solsky, 1876, by subsequent designation (Löbl et al. 2008b: 164). Status: junior synonym of Pisterotarsa Motschulsky, 1860 in Pimeliinae: Pimeliini. Synonymy: Heyden (1882: 140).

Synallecula Kolbe, 25 [F]. Type Alleculalivida Sahlberg, 1823, by subsequent designation (R. Lucas 1920: 621). Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: raised from a subgenus of Alogista Fåhraeus, 1870 to valid genus by Novák (2017b: 130).

Synatractus W.J. MacLeay, 312 [M]. Type Synatractusvariabilis W.J. MacLeay, 1887, by monotypy. Status: junior synonym of Casnonidea Fairmaire, 1882 in Lagriinae: Lagriini: Statirina. Synonymy: Carter (1920b: 199), Merkl (1986: 188).

Syncolydium Kolbe, 110 [N]. Type Syncolydiumglabratum Kolbe, 1897, by monotypy. Status: junior synonym of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini. Synonymy: Bremer (1995: 3). Note: genus originally described in Tenebrionoidea: Colydiidae.

Syngona Rye, 293 [F]. Type species [automatic]: Syggonaconcinna Fåhraeus, 1870, by monotypy. Status: junior synonym of Luprops Hope, 1833 in Lagriinae: Lupropini. Note: unjustified emendation of Syggona Fåhraeus, 1870, not in prevailing usage.

Synhimba Koch, 216 [N]. Type Psammodescordiformis Haag-Rutenberg, 1871, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Synquadrideres Iwan, 182 [M]. Type Synquadrideresnaivashaensis Iwan, 2003, by original designation. Status: junior synonym of Glyptopteryx Gebien, 1910 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 91).

Syntyphlus Koch, 243 [M]. Type Syntyphlussubterraneus Koch, 1953, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Szentivanya Kaszab, 1 [F]. Type Szentivanyametasternalis Kaszab, 1958, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Tabarus Gebien, 317 [M]. Type Tabarusinfernalis Gebien, 1921, by subsequent designation (Gebien 1940: 1089). Status: valid genus in Stenochiinae: Cnodalonini.

Taclamacanius Ferrer & Yvinec, 121 [M]. Type species [automatic]: Taklamakanialepetzi Ferrer & Yvinec, 2004, by monotypy. Status: junior synonym of Paranemia Heyden, 1892 in Diaperinae: Phaleriini. Synonymy: G.S. Medvedev (2006: 557). Note: replacement name for Taklamakania Ferrer & Yvinec, 2004.

Tactoderus Fairmaire, 112 [M]. Type Tactoderussubopacus Fairmaire, 1892 (= Praogenagagatina Mäklin, 1863), by monotypy. Status: junior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Synonymy: Gebien (1948: 544).

Tadzhikistania Bogatchev, 35 [F]. Type Tadzhikistaniamystacea Bogatchev, 1960, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Taenobates Motschulsky, 25 [M]. Type Tenebriosaperdoides G.-A. Olivier, 1795, by original designation. Status: junior synonym of Xylopinus J.L. LeConte, 1862 in Stenochiinae: Cnodalonini. Synonymy: C.O. Waterhouse (1876: 288). Note: we act as First Revisers and reject the alternative original spelling Taeniobates, used by Motschulsky (1872: 32).

Tagalinus Kaszab, 301, 333 [M]. Type Ulomalifuanum Montrouzier, 1860, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Tagalopsis Kaszab, 471, 475 [F]. Type Tagalopsisszekessyi Kaszab, 1955, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Tagalus Gebien, 388 [M]. Type Tagalusimpressicollis Gebien, 1914, by subsequent designation (Gebien 1940: 756). Status: junior synonym of Dioedus J.L. LeConte, 1862 in Phrenapatinae: Penetini. Synonymy: Kaszab (1977a: 314).

Tagenesthes Koch, 276, 282 [F]. Type Asphaltesthesafrogermanica Koch, 1941, by monotypy. Status: valid subgenus of Asphaltesthes Kraatz, 1865 in Pimeliinae: Tentyriini.

Tagenia Latreille, 170 [F]. Type Tenebriofiliformis Fabricius, 1792, by monotypy. Status: junior synonym of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina. Synonymy: G.-A. Olivier (1803: 355, through synonymy of the type species).

Tagenodes Haag-Rutenberg, 87 [M]. Type Tagenodesmoufleti Haag-Rutenberg, 1876, by monotypy. Status: junior synonym of Rhammatodes Haag-Rutenberg, 1876 in Pimeliinae: Tentyriini. Synonymy: Koch (1952a: 133).

†Tagenopsis Heer, 377 [F]. Type Tagenopsisbrevicornis Heer, 1864, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6); described from Middle Miocene deposits (Germany).

Tagenostola Reitter, 138, 151 [F]. Type Stenosisturkestanica Reitter, 1886, by subsequent designation (Gebien 1937a: 684). Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Tagona Fischer, pl. 16 [F]. Type Tagonaacuminata Fischer, 1820, by subsequent designation (Hope 1841: 124). Status: valid genus in Blaptinae: Blaptini: Prosodina.

Tagonoides Fairmaire, 343 [F]. Type Tagonoidesdelavayi Fairmaire, 1886, by subsequent designation (Blair 1923a: 283). Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Taichius Ando, 196, 197 [M]. Type Platycrepishemiceroides Blair, 1929, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Taiwanocryphaeus Masumoto, 67 [M]. Type Taiwanocryphaeusrhinoceros Masumoto, 1996, by original designation. Status: valid genus in Tenebrioninae: Toxicini: Toxicina.

Taiwanolagria Masumoto, 41 [F]. Type Taiwanolagriamerkli Masumoto, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Taiwanomenephilus Masumoto, 61 [M]. Type Taiwanomenephiluschui Masumoto, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Taiwanomenimus Masumoto, Akita & Lee, 161 [M]. Type Taiwanomenimusnakasatoi Masumoto, Akita & Lee, 2019, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Taiwanotagalus Masumoto, 143 [M]. Type Taiwanotagalusklapperichi Masumoto, 1982, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Taiwanotrachyscelis Masumoto, Akita & Lee, 29 [F]. Type Taiwanotrachyscelischengi Masumoto, Akita & Lee, 2012, by original designation. Status: valid genus in Diaperinae: Trachyscelini.

Taklamakania Ferrer & Yvinec, 42 [F]. Type Taklamakanialepetzi Ferrer & Yvinec, 2004, by monotypy. Status: junior synonym of Paranemia Heyden, 1892 in Diaperinae: Phaleriini. Synonymy: G.S. Medvedev (2006: 557). Note: junior homonym of Taklamakania Zhang, 1979 [Trilobita].

Talanus Jacquelin du Val, 156 [M]. Type Talanuscribrarius Jacquelin du Val, 1857, by monotypy. Status: valid genus in Stenochiinae: Talanini.

Tamatasida Koch, 90, 145 [F]. Type Scotinesthestuberculosa Fairmaire, 1895, by original designation. Status: valid genus in Pimeliinae: Asidini.

Tamdaous Pic, 11 [M]. Type Tamdaousimpressus Pic, 1923, by monotypy. Status: valid subgenus of Oedemutes Pascoe, 1860 in Stenochiinae: Cnodalonini.

Tamena Reitter, 90, 143 [F]. Type Psammocryptusrugiceps Reitter, 1897, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Tanchirus Fairmaire, 224 [M]. Type Tanchiruscompactus Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tangiprosodes G.S. Medvedev, 84 [M]. Type Prosodespygmaea Kraatz, 1882, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Note: originally described as a section within a subgenus.

Tanychilus Newman, 487 [M]. Type Tanychilusstriatus Newman, 1838, by subsequent designation (Cazurro Ruiz 1897a: 213). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Tanylypa Pascoe, 152 [F]. Type Tanylypamorio Pascoe, 1869, by monotypy. Status: valid genus in Zolodininae.

Tapenopsis Solier, 5 [F]. Type Tapenopsiscostata Solier, 1843, by original designation. Status: valid genus in Pimeliinae: Leptodini.

Taphrosoma Kirsch, 195 [N]. Type Taphrosomadohrnii Kirsch, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tapinocomus Gebien, 102 [M]. Type Tapinocomussubnudus Gebien, 1928, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Tapinopsis Agassiz, 361 [F]. Type species [automatic]: Tapenopsiscostata Solier, 1843, by original designation. Status: junior synonym of Tapenopsis Solier, 1843 in Pimeliinae: Leptodini. Note: unjustified emendation of Tapenopsis Solier, 1843, not in prevailing usage.

Taraxides C.O. Waterhouse, 289 [M]. Type Helopssinuatus Fabricius, 1801 (= Tenebriolaevigatus Fabricius, 1781), by subsequent designation (Gebien 1941: 340). Status: valid genus in Stenochiinae: Cnodalonini. Note: the original combination of the accepted name of the type species, Tenebriolaevigatus Fabricius, 1781, is a junior primary homonym of Tenebriolaevigatus Linnaeus, 1767.

Tarpela Bates, 272 [F]. Type Tarpelabrownii Bates, 1870, by subsequent designation (Gebien 1943: 407). Status: valid genus in Tenebrioninae: Helopini: Helopina.

Tarphiophasis Wollaston, 227 [F]. Type Tarphiophasistuberculata Wollaston, 1877, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Tarsocnodes Gebien, 82 [M]. Type Psammodesmolossus Haag-Rutenberg, 1871, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Tarsosis Gebien, 33, 34 [F]. Type Stenocaradamarense Péringuey, 1886, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Tauroceras Hope, 130 [N]. Type Tenebriocornutus Fabricius, 1775, by original designation. Status: valid genus in Tenebrioninae: Centronopini.

Tauroceropedus Pic, 4 [M]. Type Tauroceropedusdifformipes Pic, 1913, by subsequent designation (Gebien 1941: 344). Status: junior synonym of Tauroceras Hope, 1841 in Stenochiinae: Cnodalonini. Synonymy: Ferrer et al. (2005: 272).

Taurohelops Keskin & Nabozhenko, 84 [M]. Type Stenomaxincultus Allard, 1877, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Taxes Champion, 226 [M]. Type Taxesdepressus Champion, 1895, by subsequent designation (R. Lucas 1920: 628). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Tearchus Kraatz, 110 [M]. Type Tearchusannulipes Kraatz, 1880, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tedinus Casey, 153 [M]. Type Tedinusangustus Casey, 1891, by monotypy. Status: junior synonym of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina. Synonymy: Bousquet and Campbell (1991: 259).

Telabis Casey, 331 [M]. Type Eurymetoponlongipenne Casey, 1890, by subsequent designation (Casey 1907: 288). Status: valid genus in Pimeliinae: Edrotini. Note: see Alonso-Zarazaga in Bousquet et al.(2018: 116) for comments on the gender of this genus-group name.

Telacis Poey, 322 [M]. Type species [automatic]: Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: replacement name for Cistela Fabricius, 1775, a senior synonym of Cteniopus Solier, 1835.

Telaponium Blaisdell, 209 [N]. Type Telaponiumcastaneum Blaisdell, 1923, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Telchis Champion, 142 [M]. Type Telchisclavicornis Champion, 1886, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Teles Mulsant & Godart, 163 [M]. Type Teleseutymi Mulsant & Godart, 1876, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: the alternative original spelling Tales, used by Mulsant and Godart (1876: 164), was rejected by Rye (1879: 62) who acted as the First Reviser.

Telesicles Champion, 450 [M]. Type Telesiclescordatus Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Telethrus Pascoe, 29 [M]. Type Telethrusebeninus Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Telleus Fairmaire, 465 [M]. Type Telleuscrenatus Fairmaire, 1904, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Temnes Champion, 410 [M]. Type Temnescaeruleus Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Temnoaphelus Ferrer, 377 [M]. Type Temnoaphelushispidus Ferrer, 1988, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Temnophthalmus Gebien, 62, 75 [M]. Type Temnophthalmusscalaris Gebien, 1921, by subsequent designation (Gebien 1941: 341). Status: valid genus in Stenochiinae: Cnodalonini.

Teneatopus Reitter, 23 [M]. Type Tenebrioatronitens Fairmaire, 1891, by monotypy. Status: junior synonym of Ariarathus Fairmaire, 1891 in Tenebrioninae: Tenebrionini. Synonymy: Ardoin (1969b: 126).

Tenebrio Linnaeus, 417 [M]. Type Tenebriomolitor Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Tenebrioninae: Tenebrionini.

Tenebriocamaria Pic, 3 [F]. Type Tenebriocamariaatra Pic, 1919, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tenebriocephalon Pic, 10 [N]. Type Tenebriocephalonpiceum Pic, 1925, by monotypy. Status: valid genus in Pimeliinae: Ceratanisini.

Tenebrioloma Gebien, 386 [N]. Type Tenebriolomasemicostatum Gebien, 1910, by monotypy. Status: junior synonym of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini. Synonymy: Halstead (1967b: 270).

Tenebriomimus Kolbe, 342 [M]. Type Tenebriomimusadansoniarum Kolbe, 1901 (= Palembusocularis Casey, 1891), by monotypy. Status: junior synonym of Ulomoides Blackburn, 1888 in Diaperinae: Diaperini: Diaperina. Synonymy: Gebien (1922b: 268, with Martianus Fairmaire, 1893, a junior synonym of Ulomoides Blackburn, 1888).

Tenebrionellus Crotch, 105 [M]. Type species [automatic]: Tenebriomolitor Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini. Note: unnecessary replacement name for Tenebrio Linnaeus, 1758.

†Tenebrionites Cockerell, 67 [M]. Type Tenebrionitesanglicus Cockerell, 1920, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6); described from Middle Eocene deposits (United Kingdom).

Tenebriopsis Gebien, 170, 186 [F]. Type Tenebriopsissubtilicostis Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tenesis Duvivier, 163 [M]. Type Tenesisfemoratus Duvivier, 1892, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tentyria Latreille, 170 [F]. Type Tentyrialigurica Solier, 1835, by plenary powers (ICZN 2010a, Opinion 2244). Status: valid genus in Pimeliinae: Tentyriini. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2010a, Opinion 2244).

Tentyriina Peyerimhoff, 31 [F]. Type species [automatic]: Pimeliaorbiculata Fabricius, 1775, by subsequent designation (Gebien 1937a: 624; see ICZN 2010a, Opinion 2244). Status: junior synonym of Tentyrina Reitter, 1900 in Pimeliinae: Tentyriini. Note: unjustified emendation of Tentyrina Reitter, 1900, not in prevailing usage.

Tentyrina Reitter, 92, 166 [F]. Type Pimeliaorbiculata Fabricius, 1775, by subsequent designation (Gebien 1937a: 624; see ICZN 2010a, Opinion 2244). Status: valid genus in Pimeliinae: Tentyriini. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2010a, Opinion 2244).

Tentyriomorpha Peyerimhoff, 52 [F]. Type species [automatic]: Tentyromorphatelueti Escalera, 1913, by monotypy. Status: junior synonym of Tentyromorpha Escalera, 1913 in Pimeliinae: Tentyriini. Note: unjustified emendation of Tentyromorpha Escalera, 1913, not in prevailing usage.

Tentyriopsis Gebien, 168, 174 [F]. Type fixed herein (ICZN 1999, Article 70.3) as Tentyriopsispertyi Gebien, 1940, misidentified as Tentyriastriipuncta Perty, 1830 in the original designation by monotypy in Gebien (1928). Status: valid genus in Stenochiinae: Cnodalonini. Note: the type species Tentyriastriipuncta Perty was first established by monotypy; Gebien (1940: 1091) reported that Tentyriastriipuncta Perty of Gebien (1928) was misidentified and proposed the name Tentyriopsispertyi for it; we follow currently accepted concepts (e.g., Blackwelder 1945: 533) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Tentyriastriipuncta Perty, 1830 is also a valid species in the genus Tentyriopsis Gebien, 1928.

Tentyrodera Koch, 572 [F]. Type Microderamarginata Baudi di Selve, 1874, by monotypy. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Tentyromorpha Escalera, 38 [F]. Type Tentyromorphatelueti Escalera, 1913, by monotypy. Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Tentyronota Reitter, 93, 188 [F]. Type Micipsarotundicollis Kraatz, 1865, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Terametus Motschulsky, 193 [M]. Type Terametuscapicola Motschulsky, 1869, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Teremenes Carter, 54 [M]. Type Tenebriolongipennis Hope, 1843, by present designation. Status: junior synonym of Zophophilus Fairmaire, 1881 in Stenochiinae: Cnodalonini. Synonymy: Carter (1930: 547).

Termitonebria Wasmann in Wasmann and Brauns, 105 [F]. Type Termitonebriabraunsi Wasmann, 1925 (= Asyleptusfumosus Péringuey, 1896), by monotypy. Status: junior synonym of Asyleptus Péringuey, 1896 in Tenebrioninae: Amarygmini. Synonymy: Schawaller and Bremer (2013: 81).

Tessaromma Boheman, 91 [N]. Type Tessarommalugubre Boheman, 1858, by subsequent designation (Aalbu and Triplehorn 1991: 170). Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Gemminger in Gemminger and Harold (1870: 1923, with Pedonoeces G.R. Waterhouse, 1845, a junior synonym of Blapstinus Dejean, 1821), Aalbu and Triplehorn (1991: 170). Note: junior homonym of Tessaromma Newman, 1840 [Coleoptera: Cerambycidae].

Tetragonomecus Rye, 87 [M]. Type species [automatic]: Tetragonomenessemiviridis Chevrolat, 1878, by monotypy. Status: junior synonym of Tetragonomenes Chevrolat, 1878 in Stenochiinae: Cnodalonini. Note: unjustified emendation for Tetragonomenes Chevrolat, 1878, not in prevailing usage.

Tetragonomenes Chevrolat, clii [M]. Type Tetragonomenessemiviridis Chevrolat, 1878, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tetranillus Wasmann, 167 [M]. Type Tetranilluscostatus Wasmann, 1899, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Tetranosis G.S. Medvedev, 858 [F]. Type Tetranosisclypeoloba Koch, 1940, by original designation. Status: valid subgenus of Microtelopsis Koch, 1940 in Pimeliinae: Stenosini: Stenosina. Note: as pointed out by Aalbu et al. (2017: 326) this name was first introduced by Koch (1940b: 740); however, Koch did not designate a type species and therefore his name is unavailable; Löbl and Merkl (2003: 251) designated a type species for this genus but did not describe the genus-group taxon as new (ICZN 1999, Article 16.1); G.S. Medvedev (1995a: 858) was the first to provide a description of Tetranosis as well as fixing a type species for the genus, thereby making the name nomenclaturally available for the first time; we hereby act as First Revisers and select Microtelopsis Koch, 1940 as the valid name of the genus based on the Principle of Priority (ICZN 1999, Article 23.3.5).

Tetraphyllus Laporte & Brullé, 332, 404 [M]. Type Tetraphylluslatreillei Laporte & Brullé, 1831, by subsequent designation (Gebien 1935: 64). Status: valid genus in Stenochiinae: Cnodalonini. Note: nomenclatural stability in this genus is threatened by the discovery of an older type species designation (Tetraphyllusformosus Laporte & Brullé, 1831, by subsequent designation by Bates (1879a: 293), which is the type species of Damatris Laporte, 1840); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Gebien (1935: 64).

Tetrethas Koch, 23, 145 [M]. Type Anethasxylophilus Koch, 1962, by original designation. Status: valid subgenus of Anethas Jakobson, 1924 in Pimeliinae: Stenosini: Stenosina.

Tetromma Dejean, 183 [N]. Type Upisunicolor Herbst, 1797, by subsequent designation (Löbl et al. 2008a: 43). Status: junior synonym of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Dejean (1836: 203).

Texaponium Thomas, 658 [N]. Type Cryptadiustriplehorni Berry, 1974, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Thaioblaps Masumoto, 187 [F]. Type Thaioblapspunneeae Masumoto, 1989, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Thalpobia Fairmaire, 385 [F]. Type Thalpobialaevipennis Fairmaire, 1871, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Thalpophila Solier, 253, 370 [F]. Type Akisabbreviata Fabricius, 1801, by subsequent designation (Chevrolat 1849: 544). Status: senior synonym of Thalpophilodes Strand, 1942 in Pimeliinae: Tentyriini. Note: junior homonym of Thalpophila Hübner, 1820 [Lepidoptera].

Thalpophilodes Strand, 391 [M]. Type species [automatic]: Akisabbreviata Fabricius, 1801, by subsequent designation (Chevrolat 1849: 544). Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Thalpophila Solier, 1835.

Tharsus J.L. LeConte, 233 [M]. Type Tharsusseditiosus J.L. LeConte, 1862, by monotypy. Status: junior synonym of Metaclisa Jacquelin du Val, 1861 in Tenebrioninae: Metaclisini. Synonymy: Steiner (2016: 538).

Thaumatoblaps Kaszab & G.S. Medvedev, 79 [F]. Type Thaumatoblapsmarikovskiji Kaszab & G.S. Medvedev, 1984, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Theatetes Champion, 420 [M]. Type Theatetesbasicornis Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Thecacerus Lacordaire, 420 [M]. Type Cnodalonnodosum Gray, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Theresea Pic, 12 [F]. Type Thereseadiversipennis Pic, 1917, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: Masumoto (1999b: 130) transferred this genus from Cnodalonini.

Thesilea Haag-Rutenberg, 103 [F]. Type Thesileaversicolor Haag-Rutenberg, 1878, by subsequent designation (R. Lucas 1920: 638). Status: valid genus in Stenochiinae: Cnodalonini.

Thettea Bates, 290 [F]. Type Thetteatenuitarsis Bates, 1879, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Thinobatis Eschscholtz, 5, 8 [F]. Type Thinobatisferruginea Eschscholtz, 1831, by monotypy. Status: valid genus in Pimeliinae: Thinobatini.

Thoracon Gistel, xi [N]. Type species [automatic]: Silphasabulosa Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Opatrum Fabricius, 1775 in Blaptinae: Opatrini: Opatrina. Note: unnecessary replacement name for Opatrum Fabricius, 1775.

Thoracophorus Hope, 188 [M]. Type Thoracophoruswalckenaerii Hope, 1841, by original designation. Status: senior synonym of Cardiothorax Motschulsky, 1860 in Lagriinae: Adeliini. Synonymy: Motschulsky (1860a: 67). Note: junior homonym of Thoracophorus Motschulsky, 1837 [Coleoptera: Staphylinidae].

Thoracostira Borchmann, 240, 508 [F]. Type Statirasculpta Kirsch, 1873, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Thorictophasis Koch, 80 [F]. Type Caenocrypticusdeserticus Koch, 1950, by original designation. Status: junior synonym of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini. Synonymy: Endrödy-Younga (1996: 13).

Thorictosoma Lea, 257 [N]. Type Thorictosomaectatommae Lea, 1919, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Thorictosomatina.

Thornella Novák, 83 [F]. Type Alleculaholomelaena Fairmaire, 1894, by original designation. Status: valid subgenus of Upinella Mulsant, 1856 in Alleculinae: Alleculini: Alleculina.

Thoseus Pic, 9 [M]. Type Thoseusrufus Pic, 1925, by monotypy. Status: valid genus in Lagriinae: Belopini.

Thraustocolus Kraatz, 414 [M]. Type species [automatic]: Calobamonleptoderus Kraatz, 1865, by monotypy. Status: valid genus & subgenus in Pimeliinae: Tentyriini. Note: replacement name for Calobamon Kraatz, 1865.

Threnus Motschulsky, 404 [M]. Type Threnusniger Motschulsky, 1870, by original designation. Status: senior synonym of Argoporis Horn, 1870 in Tenebrioninae: Cerenopini. Synonymy: Aalbu et al. (1995: 483). Note: see the entry for Argoporis Horn, 1870 regarding the priority of these two names.

Thriptera Solier, 9, 48 [F]. Type Thripteramaillei Solier, 1836 (= Pimeliacrinita Klug, 1830), by subsequent designation (Hope 1841: 118). Status: valid genus in Pimeliinae: Pimeliini.

Thurea Ferrer, 151 [F]. Type Thureapalmi Ferrer, 1998, by original designation. Status: junior synonym of Platycotylus Olliff, 1883 in Tenebrioninae: Palorini. Synonymy: Schawaller (2014: 51).

Thydemorphus Pic, 19 [M]. Type Thydemorphuspilitarsis Pic, 1918, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Thydemus Lewis, 475 [M]. Type Scotaeuspurpurivittatus Marseul, 1876, by original designation. Status: junior synonym of Pseudonautes Fairmaire, 1892 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1914a: 58).

Thylacoderes Solier, 44, 123 [M]. Type Thylacodereseumolpoides Solier, 1843, by original designation. Status: valid genus in Pimeliinae: Praociini.

Tibinella Novák, 91 [F]. Type Upinellapahangica Novák, 2019, by original designation. Status: valid subgenus of Upinella Mulsant, 1856 in Alleculinae: Alleculini: Alleculina.

Tibiocnodes Gearner & Kamiński in Gearner et al., 2021: 8 [M]. Type Psammodeslucidus Fåhraeus, 1870, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Tidiguinia Español, 249 [F]. Type Tidiguiniabolivari Español, 1959, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Timeneca Carter, 78 [F]. Type species [automatic]: Ctimenebreweri Bates, 1873, by monotypy. Status: junior synonym of Mitrothorax Carter, 1914 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 456). Note: replacement name for Ctimene Bates, 1873, a senior synonym of Mitrothorax Carter, 1914.

Timogebienus Ardoin, 308, 331 [M]. Type Hoplonyxcollaris Gebien, 1910, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Timosmithus Ardoin, 457 [M]. Type Timosmithusbasilewskyi Ardoin, 1974, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Tinophyllus Fairmaire, 234 [M]. Type Tinophyllusgracilicornis Fairmaire, 1869, by monotypy. Status: junior synonym of Camariodes Fairmaire, 1869 in Stenochiinae: Cnodalonini. Synonymy: Fairmaire (1886c: 75).

Tisamenes Champion, 64 [M]. Type Tisamenestruquii Champion, 1884, by monotypy. Status: valid subgenus of Philolithus Lacordaire, 1858 in Pimeliinae: Asidini.

Titaena Erichson, 178 [F]. Type Titaenacolumbina Erichson, 1842, by subsequent designation (Chevrolat 1849: 594). Status: valid genus in Tenebrioninae: Titaenini.

Tithassa Pascoe, 125 [F]. Type Tithassacorynomelas Pascoe, 1860, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Tjikoraia Pic, 18 [F]. Type Tjikoraiajavana Pic, 1921, by subsequent designation (Riley 1923: 130). Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Kaszab (1979a: 81).

Tlascalinus Casey, 370 [M]. Type Trimytisflohri Champion, 1892, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Toktokkus Kamiński & Gearner in Kamiński et al., 2020: 10 [M]. Type Toktokkustschinkeli Kamiński & Gearner, 2020, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Tomogria Merkl, 135 [F]. Type Tomogriaperlata Merkl, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Tonibiastes Casey, 617 [M]. Type Notibiuscostipennis Horn, 1894, by original designation. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Tonibius Casey, 617 [M]. Type Notibiussulcatus J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 644). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Tonkinius Fairmaire, 13 [M]. Type Tonkiniussculptilis Fairmaire, 1903, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Toreuma Carter, 84 [N]. Type Toreumacupreum Carter, 1913, by monotypy. Status: senior synonym of Atoreuma Gebien, 1941 in Tenebrioninae: Heleini: Cyphaleina. Note: junior homonym of Toreuma Haeckel, 1880 [Cnidaria].

Toxicum Latreille, 174 [N]. Type Toxicumrichesianum Latreille, 1802, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Toxicini: Toxicina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6).

Toxocnema Fåhraeus, 304 [F]. Type Toxocnemarufitarsis Fåhraeus, 1870, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: transferred from Tenebrioninae: Ulomini by Schawaller (2009b: 363).

Trachasida Reitter, 41, 62 [F]. Type Asidaruficornis Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Gracilasida Escalera, 1905 in Pimeliinae: Asidini. Synonymy: Wilke (1922: 259, with Planasida Escalera, 1907, a junior synonym of Gracilasida Escalera, 1905).

Tracheloeum Hope, 116 [N]. Type Tracheloeumlaticolle Hope, 1841, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6).

Trachelostenus Solier, 255 [M]. Type Trachelostenusinaequalis Solier, 1851, by monotypy. Status: valid genus in Tenebrioninae: Trachelostenini. Note: the family-group name based on this genus was downgraded from the family Trachelostenidae (within the superfamily Tenebrionoidea) to the tribe Trachelostenini (within the tenebrionid subfamily Tenebrioninae) by Matthews and Lawrence (2015: 290).

Trachyderes Koch, 112 [M]. Type Trachynotusbipunctatus Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Trachyderma Latreille, 576 [N]. Type Pimeliahispida Fabricius, 1775 (= Tenebriohispidus Forskål, 1775), by subsequent designation (P.H. Lucas 1839: 50). Status: valid genus and subgenus in Pimeliinae: Pimeliini. Note: gender of genus corrected from feminine to neuter by Löbl and Smetana (2011: 33).

Trachydermum Gistel, xi [N]. Type species [automatic]: Pimeliahispida Fabricius, 1775 (= Tenebriohispidus Forskål, 1775), by subsequent designation (P.H. Lucas 1839: 50). Status: junior synonym of Trachyderma Latreille, 1828 in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Trachyderma Latreille, 1828.

Trachymetus Reitter, 51, 76 [M]. Type Pachypteruselongatus Mulsant & Rey, 1859, by monotypy. Status: junior synonym of Amblysphagus Fairmaire, 1896 in Blaptinae: Opatrini: Neopachypterina. Synonymy: Kaszab (1975a: 27).

Trachynotidus Péringuey, 296 [M]. Type Psammodesthoreyi Haag-Rutenberg, 1871, by subsequent designation (Kamiński et al. 2019b: 18). Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Trachynotus Latreille, 579 [M]. Type Sepidium vittatum Fabricius, 1781, by subsequent designation (Kamiński et al. 2019b: 93). Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Trachyscelis Latreille, 379 [F]. Type Trachyscelisaphodioides Latreille, 1809, by monotypy. Status: valid genus in Diaperinae: Trachyscelini.

Tragardhus Koch, 369 [M]. Type Tragardhusgrandipleurum Koch, 1956, by original designation. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Trestonia Rafinesque, 113 [F]. Type species [automatic]: Toxicumrichesianum Latreille, 1802, by monotypy. Status: junior synonym of Toxicum Latreille, 1802 in Tenebrioninae: Toxicini: Toxicina. Note: unnecessary replacement name for Toxicum Latreille, 1802.

Triangulipenna Louw, 100, 114 [F]. Type Triangulipennalacuna Louw, 1979, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Tribolium W.S. MacLeay, 47 [N]. Type Colydiumcastaneum Herbst, 1797, by monotypy (see ICZN 1988, Opinion 1495). Status: valid genus and subgenus in Tenebrioninae: Triboliini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1988, Opinion 1495).

Trichamarygmus Carter, 46 [M]. Type Trichamarygmuspilosus Carter, 1913, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Trichanemia Ardoin, 361, 406 [F]. Type Anemiaschmitzi Ardoin, 1971, by monotypy. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini.

Tricheleodes Blaisdell, 138 [M]. Type Eleodeshirsuta J.L. LeConte, 1861, by subsequent designation (Johnston 2016: 666). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Trichethmus Gebien, 45 [M]. Type Trichethmuslobicollis Gebien, 1937, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Trichiasida Casey, 77, 172 [F]. Type Pelecyphorushirsutus J.L. LeConte, 1851, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Trichiotes Casey, 432, 443 [M]. Type Trichiotesseriatus Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Trichochianalus Kaszab, 149, 201 [M]. Type Platynoscelismonticola Kaszab, 1940, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini. Note: the spellings Trichochianalis (p. 201 – also used in the directory of new taxa published in volume 30 of the journal Mitteilungen der Münchner Entomologischen Gesellschaft (p. xi)) and Trichochianalus (pp 125, 149, 201) were used in the original publication; Kaszab (1940b: 989) used the spelling Trichochianalus only and therefore acted as the First Reviser (ICZN 1999, Article 24.2.4).

Trichodamatris Chatanay, 506, 524 [M]. Type Porphyrhybaraffrayi Fairmaire, 1884, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Trichoderulus Blaisdell, 281 [M]. Type Trichoderuluslongipilosus Blaisdell, 1923 (= Eleodestribulus Thomas, 2005), by original designation. Status: junior synonym of Pseudeleodes Blaisdell, 1909 in Blaptinae: Amphidorini. Synonymy: Johnston (2016: 672).

Tricholeipopleura Kaszab, 152, 223 [F]. Type Platynoscelislucidicollis Kraatz, 1882, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini. Note: the spellings Tricholeipopleura (pp 131, 223, 225–228, also used in the directory of new taxa published in volume 30 of the journal Mitteilungen der Münchner Entomologischen Gesellschaft (p. xii)) and Tricholeipoleura (pp. 152) were used in the original publication; Kaszab (1940b: 989) used the spelling Tricholeipopleura only and therefore acted as the First Reviser (ICZN 1999, Article 24.2.4).

Trichomyatis Schuster in Reinig, 893 [F]. Type Trichomyatisconradti Schuster, 1931, by original designation. Status: valid genus in Blaptinae: Platyscelidini.

Trichoodescelis Kaszab, 940, 954 [F]. Type Platyscelisacutangula Kraatz, 1884, by original designation. Status: junior synonym of Longuloodescelis Kaszab, 1940 in Blaptinae: Platyscelidini. Synonymy: Egorov (2004: 595).

Trichoplatynoscelis Kaszab, 896 [F]. Type Trichoplatynoscelispamirensis Kaszab, 1940, by original designation. Status: junior synonym of Trichomyatis Schuster, 1931 in Blaptinae: Platyscelidini. Synonymy: Egorov (2004: 603). Note: this name was also used in several instances by Kaszab (1940a: 126, 132, 142, 143, 146); however, it is not available from that publication since there is no type species designated.

Trichoplatyscelis Reinig, 895 [F]. Type Trichoplatyscelispamirensis Reinig, 1931, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Trichopodus Mulsant & Rey, 58, 59 [M]. Type Trichopodusvalidus Mulsant & Rey, 1859, by monotypy. Status: senior synonym of Trichosternum Wollaston, 1861 in Blaptinae: Opatrini: Opatrina. Synonymy: Bates (1872a: 98). Note: junior homonym of Trichopodus Lacepède, 1802 [Pisces].

Trichosaragus Blackburn, 1269 [M]. Type Trichosaraguspilosellus Blackburn, 1890, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Trichosphaena Reitter, 140, 145 [F]. Type Sphenariaolgae Semenov, 1889, by subsequent designation (Gebien 1937a: 577). Status: valid genus in Pimeliinae: Tentyriini.

Trichosternum Wollaston, 205 [N]. Type Trichosternumstriatum Wollaston, 1861, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Trichostethe Koch, 285 [F]. Type Cyphostethepilosa Koch, 1950, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini.

Trichotenebrio Ardoin, 64 [M]. Type Trichotenebrioatronitidus Ardoin, 1962, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Trichotoides Marcuzzi, 23 [M]. Type Scapteshintoni Kaszab, 1949, by monotypy. Status: junior synonym of Ammodonus Mulsant, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Ferrer and Moraguès (2001: 499).

Trichoton Hope, 111 [N]. Type Trichotoncayennense Hope, 1841, by original designation. Status: valid genus in Blaptinae: Opatrini: Blapstinina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6).

Trichotrachys Koch, 201 [M]. Type Trachynotussordidus Gerstaecker, 1854, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Trichotrichus Koch, 108 [M]. Type Trachynotuscrinitus Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Trichotum Agassiz, 376 [N]. Type species [automatic]: Trichotoncayennense Hope, 1841, by original designation. Status: junior synonym of Trichoton Hope, 1841 in Blaptinae: Opatrini: Blapstinina. Note: unjustified emendation of Trichoton Hope, 1841, not in prevailing usage.

Trichulodes Carter, 223 [M]. Type Trichulodespunctatus Carter, 1914, by monotypy. Status: junior synonym of Pseudolyprops Fairmaire, 1882 in Lagriinae: Goniaderini. Synonymy: Doyen et al. (1990: 231).

Trientoma Solier, 253, 256 [F]. Type Trientomavarvasi Solier, 1835, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Trigonocnera Reitter, 202, 213 [F]. Type Trigonoscelispseudopimelia Reitter, 1889, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Trigonopachys Skopin, 300 [M]. Type Trigonopachysmichailovi Skopin, 1968, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Trigonopilus Fairmaire, 22 [M]. Type Trigonopiluslaticeps Fairmaire, 1893, by monotypy. Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Trigonopoda Gebien, 2 [F]. Type Trigonopodacrassipes Gebien, 1914, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Trigonopus Mulsant & Rey, 105 [M]. Type Trigonopuscapicola Mulsant & Rey, 1853, by subsequent designation (Cazurro Ruiz 1897b: 539). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Trigonoscelis Dejean, 179 [F]. Type Pimelianodosa Fischer, 1820, by subsequent designation (Hope 1841: 118). Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Trigonotarsus Hope, 357 [M]. Type Trigonotarsusaustralis Hope, 1843, by original designation. Status: senior synonym of Sobas Pascoe, 1863 in Blaptinae: Opatrini: Opatrina. Note: junior homonym of Trigonotarsus Guérin-Méneville, 1838 [Coleoptera: Curculionidae].

Trilobocara Solier, 129 [N]. Type Trilobocaraciliatum Solier, 1851, by original designation. Status: valid genus in Pimeliinae: Trilobocarini.

Trimytantron Ardoin, 381 [N]. Type Trimytantrondecui Ardoin, 1977, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Trimytis J.L. LeConte, 141 [F]. Type Trimytispruinosa J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Triorophus J.L. LeConte, 141 [M]. Type Triorophuslaevis J.L. LeConte, 1851, by subsequent designation (Casey 1907: 432). Status: valid genus in Pimeliinae: Edrotini.

Triphalopsis Blaisdell, 232 [F]. Type Triphalopsispartida Blaisdell, 1923, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Triphalopsoides Doyen, 222 [M]. Type Triphalopsoideslasiodorsa Doyen, 1990, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Triphalus J.L. LeConte, 104 [M]. Type Triphalusperforatus J.L. LeConte, 1866, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Triplehornia Matthews & Lawrence, 543 [F]. Type Triplehorniametallica Matthews & Lawrence, 2005, by original designation. Status: valid genus in Diaperinae: incertae sedis. Note: Matthews and Lawrence (2019: 651) mentioned that this genus belongs to an unnamed tribe within the subfamily Diaperinae.

Tripolicryptus Strand, 24 [M]. Type species [automatic]: Brachycryptustripolitanus Quedenfeldt, 1891, by monotypy. Status: valid genus in Alleculinae: Cteniopodini. Note: replacement name for Brachycryptus Quedenfeldt, 1891.

Trisilus Haag-Rutenberg, 101 [M]. Type Trisilusfemoralis Haag-Rutenberg, 1878, by monotypy. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 490).

Trogloderus J.L. LeConte, 2 [M]. Type Trogloderuscostatus J.L. LeConte, 1879, by monotypy. Status: valid genus in Blaptinae: Amphidorini.

Troglogeneion Aalbu, 541 [N]. Type Troglogeneionzapoteca Aalbu, 1985, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Tromosternus Harold, 130 [M]. Type Tromosternushaagi Harold, 1876 (= Gnesishelopioides Pascoe, 1866), by monotypy. Status: junior synonym of Gnesis Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Lewis (1894: 476).

Tropidopterus Cazurro Ruiz, 637 [M]. Type Tropidopteruscarinatus Cazurro Ruiz, 1897, by original designation. Status: junior synonym of Adelium W. Kirby, 1819 in Lagriinae: Adeliini. Synonymy: Doyen et al. (1990: 230). Note: Tropidoptère was first used by Blanchard (1845: 35) but it was not formed as a scientific name and is therefore unavailable (ICZN 1999, Article 1.1); the latinized form of this name, Tropidopterus, was used subsequently (e.g., Lacordaire, 438) but was not made available to our knowledge until Cazurro Ruiz (1897b: 637) published a combined description of a new genus and single new species (ICZN 1999, Article 12.2.6); since we believe that the combined description by Cazurro Ruiz (1897b : 637) was based on the same concept as the unavailable name “Tropidoptère Blanchard” we follow the intentions of authors in recent literature (e.g., Doyen et al. 1990: 230; Matthews 1998: 777; Matthews and Lawrence 2019: 615) and treat Tropidopterus Cazurro Ruiz, 1897 as a junior synonym of Adelium W. Kirby, 1819.

Tropitrachys Koch, 229 [M]. Type Trachynotusperegrinator Koch, 1953, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Truncatocamaria Pic, 27 [F]. Type Camariaspinipes Pic, 1917 (as “Camariaspinifer”), by monotypy. Status: junior synonym of Camaria Lepeletier & Audinet-Serville, 1828 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1942a: 320), Masumoto (1993a: 147).

Truncatoodescelis Kaszab, 942, 962 [F]. Type Platyscelislongicollis Kraatz, 1884, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Tuberocnodes Gearner & Kamiński in Gearner et al., 2021: 8 [M]. Type Psammodeshumeralis Haag-Rutenberg, 1871, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Tucumana Gebien, 604 [F]. Type species [automatic]: Eusteniatenuimembris Fairmaire, 1905, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina. Note: replacement name for Eustenia Fairmaire, 1905; placed in Alleculinae by Chatanay (1915a: 526).

Turcmenicola Bogatchev, 44 [M]. Type Turcmenicolajachontovi Bogatchev, 1952, by monotypy. Status: valid subgenus of Colposcelis Dejean, 1834 in Pimeliinae: Tentyriini.

Turkmenohelops G.S. Medvedev, 98, 102 [M]. Type Zophohelopsbalchanicus G.S. Medvedev & Nepesova, 1985, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Turkonalassus Keskin, Nabozhenko & Alpagut-Keskin, 727 [M]. Type Helopsadimonius Allard, 1876, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Tydeolus Champion, 37 [M]. Type Tydeolusatratus Champion, 1884, by subsequent designation (W.F. Kirby 1885b: 80). Status: valid genus in Pimeliinae: Epitragini.

Tylophloeus Bremer, 10, 13 [M]. Type Hypophlaeusflavipennis Motschulsky, 1860, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Tyndarisus Pascoe, 294 [M]. Type Tyndarisuslongitarsus Pascoe, 1869 (= Lepispilusstygianus Pascoe, 1869), by monotypy. Status: junior synonym of Pachycoelia Boisduval, 1835 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1926: 119, with Lepispilus Westwood, 1841, a junior synonym of Pachycoelia Boisduval, 1835).

Tynteria Reitter, 301 [F]. Type Pachychilahumerosa Fairmaire, 1875, by monotypy. Status: junior synonym of Oterophloeus Desbrochers des Loges, 1881 in Pimeliinae: Tentyriini. Synonymy: Reitter (1900c: 89, 143).

Tynthlobia Fairmaire, 261 [F]. Type Tynthlobiaquadricostata Fairmaire, 1888, by monotypy. Status: junior synonym of Ethmus Haag-Rutenberg, 1873 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Fairmaire (1891b: 250).

Typhlophloeus Jeannel & Paulian, 51 [M]. Type Typhlophloeuschappuisi Jeannel & Paulian, 1945, by original designation. Status: valid genus in Diaperinae: Hypophlaeini.

Typhluloma Lea, 475 [N]. Type Typhlulomainops Lea, 1912, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Typhlusechus Linell, 154 [M]. Type Typhlusechussingularis Linell, 1897, by original designation. Status: valid genus in Pimeliinae: Stenosini: Typhlusechina.

Typhobia Pascoe, 279 [F]. Type Typhobiafuliginea Pascoe, 1869, by monotypy. Status: junior synonym of Platydema Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Synonymy: Champion (1886: 181).

Tyrtaeus Champion, 76 [M]. Type Tyrtaeusrufus Champion, 1913, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Ubangia Gebien, 54 [F]. Type Ubangialatifrons Gebien, 1914, by monotypy. Status: junior synonym of Crypsinous Fairmaire, 1891 in Tenebrioninae: Amarygmini. Synonymy: Ardoin (1964a: 839).

Ucalegon Champion, 65 [M]. Type Ucalegonpulchellus Champion, 1884, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Udebra Reitter, 236 [F]. Type Udebrahauseri Reitter, 1896 (= Erodiusfimbriatus Ménétriés, 1849), by monotypy. Status: junior synonym of Adavius Mulsant & Rey, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Kaszab (1942: 28).

Uenomisolampidius Masumoto, 36 [M]. Type Ueonomisolampidiusshunichii Masumoto, 1996, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Uenostrongylium Masumoto, 123 [N]. Type Cryptobateslaosensis Pic, 1928, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Uleda Laporte, 220 [F]. Type Ulomadiaperoides Laporte, 1840, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Uloma Dejean, 67 [N]. Type Tenebrioculinaris Linnaeus, 1758, by plenary powers (ICZN 1975, Opinion 1039). Status: valid genus and subgenus in Tenebrioninae: Ulomini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1975, Opinion 1039). Note: the name Uloma is a Greek noun οὔλωμα (meaning cicatrisation) and the gender is neuter instead of the previously accepted feminine gender (e.g., ICZN 1975, Opinion 1039; see ICZN 2021, Official Correction 134).

Ulomimus Bates, 201 [M]. Type Ulomimusindicus Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Ulomini. Note: unjustified emendation of the original spelling Ulomimimus, introduced by Rye (1875: 288), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Ulomina Baudi di Selve, 112 [F]. Type Ulominacarinata Baudi di Selve, 1876, by monotypy. Status: valid genus in Tenebrioninae: Palorini. Note: redescribed as new by Baudi di Selve (1876b: 235).

Ulomoides Blackburn, 274 [M]. Type Ulomoideshumeralis Blackburn, 1888, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Ulomoides Escalera, 501 [M]. Type species [automatic]: Crypticusviaticus Fairmaire, 1851, by original designation. Status: senior synonym of Platycrypticus Español, 1952 in Diaperinae: Crypticini. Note: junior homonym of Ulomoides Blackburn, 1888 [Coleoptera: Tenebrionidae: Diaperinae: Diaperini: Diaperina].

Ulomotypus Broun, 841 [M]. Type Ulomotypuslaevigatus Broun, 1886, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Ulosonia Laporte, 220 [F]. Type Ulomatricorne Laporte, 1840 (= Phaleriatricornis Dalman, 1823), by subsequent designation (Gebien 1940: 786). Status: junior synonym of Hypogena Dejean, 1834 in Tenebrioninae: Triboliini. Synonymy: Jacquelin du Val (1857: 148).

Ulus Horn, 349, 358 [M]. Type Blapstinuscrassus J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 665). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Umslatus Péringuey, 312 [M]. Type Umslatusagilis Péringuey, 1899, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Uniungulum Koch, 113 [N]. Type Uniungulumhoeschi Koch, 1962, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Upembarus Koch, 220 [M]. Type Upembarussaegeri Koch, 1956, by original designation. Status: valid genus and subgenus in Blaptinae: Platynotini: Platynotina.

Upinella Mulsant, 17 [F]. Type Alleculaaterrima Rosenhauer, 1847, by monotypy. Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Upis Fabricius, 515 [F]. Type Attelabusceramboides Linnaeus, 1758, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Uptona G.S. Medvedev & Lawrence, 582 [F]. Type Uptonapallida G.S. Medvedev & Lawrence, 1986, by original designation. Status: valid genus in Diaperinae: Hyociini: Uptonina.

Uria Gistel, x [F]. Type species [automatic]: Tenebriocadaverinus Fabricius, 1792, by subsequent designation (Westwood 1838: 32). Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Note: unnecessary replacement name for Phaleria Latreille, 1802.

Uriela Reitter, 518 [F]. Type Podhomalafausti Kraatz, 1881, by monotypy. Status: junior synonym of Podhomala Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Sénac (1888: lv).

Urielina Reitter, 331 [F]. Type Podhomalanitida Baudi di Selve, 1876, by monotypy. Status: valid subgenus of Podhomala Solier, 1836 in Pimeliinae: Pimeliini.

Uroblaps Motschulsky, 530 [F]. Type Blaps producta Brullé, 1832 (= Blaps lusitanica Herbst, 1799), by subsequent designation (Nabozhenko 2008: 36). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860)

Uroplatopsis Champion, 53 [F]. Type Uroplatopsisimitator Champion, 1889, by subsequent designation (R. Lucas 1920: 666). Status: valid genus in Lagriinae: Lagriini: Statirina.

Uroprosodes Reitter, 119 [M]. Type Prosodescostifera Kraatz, 1886, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Uyttenboogaartia Koch, 592, 595 [F]. Type Hegetercribricollis Brullé, 1839, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Uzagaria Ancey, 509 [F]. Type Uzagariapubens Ancey, 1881, by monotypy. Status: junior synonym of Emmalus Erichson, 1843 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1910b: 306, as Emmallus).

†Vabole Alekseev & Nabozhenko, 128 [F]. Type Vaboletriplehorni Alekseev & Nabozhenko, 2015, by original designation. Status: valid genus in Tenebrioninae: Palorini. Note: described from Eocene Baltic amber.

Vacronus Casey, 501, 508 [M]. Type Vacronustenuicornis Casey, 1907, by original designation. Status: junior synonym of Alaephus Horn, 1870 in Pimeliinae: Vacronini. Synonymy: Doyen and Lawrence (1979: 350).

Vadalus Mulsant & Rey, 150 [M]. Type Pedinuspunctulatus Mulsant & Rey, 1853, by monotypy. Status: junior synonym of Pedinus Latreille, 1797 in Blaptinae: Pedinini: Pedinina. Synonymy: Gemminger in Gemminger and Harold (1870: 1918), Kamiński and Iwan (2017: 599).

Valdivium Matthews, 709, 721 [N]. Type Adeliumsulcatulum Fairmaire & Germain, 1860, by original designation. Status: valid genus in Lagriinae: Adeliini.

Vaniosus Kulzer, 896 [M]. Type Vaniosusparadoxus Kulzer, 1956, by original designation. Status: valid genus in Pimeliinae: Evaniosomini.

Vansonium Koch, 354 [N]. Type Vansoniumbushmanicum Koch, 1950, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Vansoniina.

Varogeton Bremer, 37, 80 [M]. Type Dietysussubannulipes Pic, 1923, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Vernayella Koch, 129 [F]. Type Vernayellanoctivaga Koch, 1958, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini.

Vieta Laporte, 196 [F]. Type Sepidium vestitum Guérin-Méneville, 1831, by subsequent designation (Hope 1841: 116). Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Vietnalia Novák, 440 [F]. Type Vietnaliacatcatica Novák, 2021, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Vietomorpha Fairmaire, 186 [F]. Type Vietomorphafoveipennis Fairmaire, 1887, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Viettagona G.S. Medvedev & Merkl, 317, 328 [F]. Type Viettagonavietnamensis G.S. Medvedev & Merkl, 2003, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Villiersia Gridelli, 219, 227 [F]. Type Tenebrioclypealis Gebien, 1920, by original designation. Status: senior synonym of Gridellia Kammerer, 2006 in Tenebrioninae: Tenebrionini. Note: junior homonym of Villiersia d’Orbigny, 1837 [Mollusca].

Viriathus Fairmaire, 339 [M]. Type Viriathusstrigipennis Fairmaire, 1902, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Vizcainyx Aalbu & Smith, 198 [M]. Type Vizcainyxandrewsi Aalbu & Smith, 2020, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Vutsimus Péringuey, 308 [M]. Type Vutsimuspraetorius Péringuey, 1899, by subsequent designation (Gebien 1943: 922). Status: valid genus in Tenebrioninae: Amarygmini.

Wahlbergylium Ferrer, 141 [N]. Type Wahlbergyliumviklundi Ferrer, 2011, by original designation. Status: junior synonym of Argutiolana Robiche, 2001 in Stenochiinae: Cnodalonini. Synonymy: Robiche (2019a: 97).

Wallardilagria Pic, 74 [F]. Type Heterogriapallidicolor Pic, 1910, by monotypy. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Warchalowskiellus Iwan, 60 [M]. Type Trigonopuslongulus Mulsant & Rey, 1853, by original designation. Status: junior synonym of Schelodontes Koch, 1956 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan and Kamiński (2014: 171).

Waterhousia Skopin, 109, 110 [F]. Type Trigonoscelislongipes C.O. Waterhouse, 1889, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Wattadelium Emberson, 24 [N]. Type species [automatic]: Edalusopacus Broun, 1893, by subsequent designation (R. Lucas 1920: 255). Status: valid genus in Lagriinae: Adeliini. Note: replacement name for Edalus Broun, 1893.

Wattiana Matthews & Lawrence, 537 [F]. Type Wattianagreensladei Matthews & Lawrence, 2005, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Thorictosomatina.

Wattius Kaszab, 50 [M]. Type Calymmuscucullatus Pascoe, 1871, by original designation. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Weisea Semenov, 370 [F]. Type Weiseasabulicola Semenov, 1891, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Wolladrus Iwan & Kamiński, 483 [M]. Type species [automatic]: Hadrusalpinus Wollaston, 1854, by subsequent designation (R. Lucas 1920: 313). Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: replacement name for Hadrus Wollaston, 1854.

Xanthalia Fairmaire, 395 [F]. Type species [automatic]: Xanthiacurticollis Fairmaire, 1893, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: replacement name for Xanthia Fairmaire, 1893; new placement [OM], previously included in Lagriinae: Lagriini: Lagriina.

Xanthia Fairmaire, 31 [F]. Type Xanthiacurticollis Fairmaire, 1893, by monotypy. Status: senior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Note: junior homonym of Xanthia Hübner, 1813 [Lepidoptera].

Xanthicles Champion, 231 [M]. Type Xanthiclescaraboides Champion, 1886, by subsequent designation (Gebien 1941: 815). Status: valid genus in Lagriinae: Goniaderini.

Xanthobates Gebien, 170, 185 [M]. Type Xanthobatesflavus Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Xanthohelops Nabozhenko, 822 [M]. Type Xanthohelopskarakumicus Nabozhenko & G.S. Medvedev, 2006, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Xanthomus Mulsant, 302 [M]. Type Helopspallidus Curtis, 1830, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Xanthothopeia Mäklin, 223 [F]. Type Xanthothopeiarufipennis Mäklin, 1867, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Xanthothopia Gemminger in Gemminger and Harold, 2001 [F]. Type species [automatic]: Xanthothopeiarufipennis Mäklin, 1867, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Note: unjustified emendation of Xanthothopeia Mäklin, 1867, not in prevailing usage.

Xantusiella Kaszab, 4, 18 [F]. Type Xantusiellacrenulata Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Xenius Champion, 224 [M]. Type Xeniusscabripennis Champion, 1886, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Xenocera Borchmann, 18, 116 [F]. Type Lagriocerafeai (as “feae”) Borchmann, 1911, by original designation. Status: senior synonym of Xenocerogria Merkl, 2007 in Lagriinae: Lagriini: Lagriina. Note: junior homonym of Xenocera Broun, 1881 [Coleoptera: Ptinidae].

Xenocerogria Merkl, 269 [F]. Type species [automatic]: Lagriocerafeai (as “feae”) Borchmann, 1911, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina. Note: replacement name for Xenocera Borchmann, 1936.

Xenogena Borchmann, 22, 211 [F]. Type Adynatacrinita Borchmann, 1915, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Xenogloeus Wollaston, 251 [M]. Type Xenogloeuspolitus Wollaston, 1861, by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Xenolagria Merkl, 124, 126 [F]. Type Lagriatincta Blackburn, 1889, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Xenostethus Bates, 321 [M]. Type Xenostethuslacordairii Bates, 1868, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Xenostira Borchmann, 217, 221 [F]. Type Xenostiragiraffa Borchmann, 1921, by original designation. Status: valid subgenus of Statira Lepeletier & Audinet-Serville, 1828 in Lagriinae: Lagriini: Statirina.

Xenotermes Wasmann, 616 [M]. Type Xenotermesfeai (as “feae”) Wasmann, 1896, by monotypy. Status: valid genus in Tenebrioninae: Rhysopaussini.

Xenus Péringuey, 255 [M]. Type Xenustricorniger Péringuey, 1899, by monotypy. Status: senior synonym of Aphrotus Péringuey, 1904 in Pimeliinae: Epitragini. Note: junior homonym of Xenus Kaup, 1829 [Aves].

Xerolinus Ivie & Hart, 470 [M]. Type Diastolinussallei Mulsant & Rey, 1859, by original designation. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Xyloborus Motschulsky, 64 [M]. Type Xyloboruscrenipennis Motschulsky, 1858, by monotypy. Status: senior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: Merkl and Kompantzeva (1996: 91). Note: although Schwarz and Barber (1914: 175) pointed out that the name Xyloborus was made available for the first time by Motschulsky (1858: 64) there are few occurrences of this name in the literature; reversal of precedence cannot be used to conserve usage of the broadly used name Rhipidandrus J.L. LeConte, 1862 since Xyloborus was used as valid after 1899 (e.g., Swezey 167); an application to the ICZN is necessary to conserve usage Rhipidandrus J.L. LeConte, 1862; the older names Xyloborus Kirby & Spence, 1828 and Xyloborus Dejean, 1834 [Coleoptera] are unavailable because they were published before 1931 without a description, a definition or an indication (ICZN 1999, Article 12.1).

Xylochus Broun, 396 [M]. Type Xylochustibialis Broun, 1880, by subsequent designation (Watt 1992: 24). Status: valid genus in Alleculinae: Alleculini: incertae sedis.

Xylopinus J.L. LeConte, 230 [M]. Type Tenebrioanthracinus Knoch, 1801 (= Tenebriosaperdoides G.-A. Olivier, 1795), by subsequent designation (Gebien 1941: 336). Status: valid genus in Stenochiinae: Cnodalonini.

Xysta Eschscholtz, 9 [F]. Type Eleodesgravida Eschscholtz, 1829, by subsequent designation (Hope 1841: 124). Status: senior synonym of Steneleodes Blaisdell, 1909 in Blaptinae: Amphidorini. Synonymy: Smith and Johnston in Bousquet et al. (2018: 166). Note: junior homonym of Xysta Meigen, 1824 [Diptera].

Xystronia Solier, 238 [F]. Type Xystroniacoerulea Solier, 1835, by monotypy. Status: valid subgenus of Lystronychus Latreille, 1829 in Alleculinae: Alleculini: Xystropodina.

Xystropus Solier, 241 [M]. Type Xystropuspilosus Solier, 1835, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina. Note: see Bousquet et al. (2015: 139) for comments regarding the species originally included in this genus.

Yamatotakeru Ando, 385 [M]. Type Ischnodactylusloripes Lewis, 1894, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

†Yantaroxenos Nabozhenko, Kirejtshuk & Merkl, 564 [M]. Type Yantaroxenoscolydioides Nabozhenko, Kirejtshuk & Merkl, 2016, by original designation. Status: valid genus in Lagriinae: Belopini. Note: described from Eocene Baltic amber.

Yarranum Matthews, 708, 734 [N]. Type Seirotranacrenicollis Pascoe, 1869, by original designation. Status: valid genus in Lagriinae: Adeliini.

Zabroideus Fairmaire, 219 [M]. Type Zabroideuspinguis Fairmaire, 1894, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Zadenos Laporte, 210 [M]. Type Opatrumlongipalpe Wiedemann, 1823, by monotypy. Status: valid subgenus of Selenepistoma Dejean, 1834 in Blaptinae: Dendarini: Melambiina. Note: this name was recently treated as a valid genus (Kamiński 531); however, the older available genus name Selenepistoma Dejean, 1834 has priority and therefore Zadenos is downgraded to a valid subgenus of Dejean’s name; new status.

Zaleucus Champion, 491 [M]. Type species [automatic]: Zamolxisdilatatus Champion, 1884, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini. Note: replacement name for Zamolxis Champion, 1884.

Zambesmia Bouchard & Bousquet, new subgenus [F]. Type Macropodachiyakensis Kuntzen, 1916, by present designation. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: Koch (1944b: 149) introduced the new subgenus name Zambesmia for three nominal species, but unfortunately did not designate a type species; the subgenusZambesmia, which has been treated as valid since 1944, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Macropodachiyakensis Kuntzen, 1916 as type species and referring to Koch (1944b: 149) for the character states that characterise and differentiate Zambesmia.

Zamolxis Champion, 70 [M]. Type Zamolxisdilatatus Champion, 1884, by monotypy. Status: senior synonym of Zaleucus Champion, 1892 in Pimeliinae: Asidini. Note: junior homonym of Zamolxis Stål, 1865 [Hemiptera].

Zarudnionymus Semenov-Tjan-Shansky & Bogatchev, 175 [M]. Type Zarudnionymuspersis Semenov-Tjan-Shansky & Bogatchev, 1947 (= Adelostomagrande Haag-Rutenberg, 1879), by monotypy. Status: valid subgenus of Adelostoma Duponchel, 1827 in Pimeliinae: Adelostomini.

Zeadelium Watt, 32 [N]. Type Adeliumlentum Broun, 1880, by original designation. Status: valid genus in Lagriinae: Adeliini.

Ziaelas Fairmaire, cx [M]. Type Ziaelasinsolitus Fairmaire, 1892, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Zidalus Mulsant & Rey, 71 [M]. Type Opatrinuscorvinus Mulsant & Rey, 1853, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Zizu Novák, 186 [M]. Type Zizukejvali Novák, 2019, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Zodinus Mulsant & Rey, 90 [M]. Type Opatrinusservus Mulsant & Rey, 1853, by subsequent designation (Koch 1956a: 93). Status: junior synonym of Zidalus Mulsant & Rey, 1853 in Blaptinae: Platynotini: Platynotina. Synonymy and First Reviser action (Zodinus Mulsant & Rey, 1853 versus Zidalus Mulsant & Rey, 1853) is Iwan (1995b: 362).

Zolodinus Blanchard, 159 [M]. Type Zolodinuszelandicus Blanchard, 1853, by monotypy. Status: valid genus in Zolodininae.

Zomedes Watt, 25 [M]. Type Zomedesborealis Watt, 1992, by original designation. Status: valid genus in Alleculinae: Alleculini: incertae sedis.

Zophius Dejean, 189 [M]. Type Helopsrufopictus Wiedemann, 1823, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Zophobas Dejean, 204 [M]. Type Helopsmorio Fabricius, 1777 (= Tenebrioatratus Fabricius, 1775), by subsequent designation (Motschulsky 1872: 26). Status: valid genus and subgenus in Tenebrioninae: Tenebrionini. Note: according to Matthews and Lawrence (2019: 628) more research is needed to establish the correct placement of this genus within the subfamily Tenebrioninae.

Zophodes Fåhraeus, 298 [M]. Type Zophodestristis Fåhraeus, 1870, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Zophohelops Reitter, 221 [M]. Type species [automatic]: Euryhelopstiro Reitter, 1902, by subsequent designation (Gebien 1943: 424). Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina. Note: replacement name for Euryhelops Reitter, 1902.

Zophondrus Nabozhenko, 240 [M]. Type Zophohelopsiranensis Nabozhenko, 2014, by original designation. Status: valid subgenus of Zophohelops Reitter, 1902 in Tenebrioninae: Helopini: Cylindrinotina.

Zophophilus Fairmaire, 359 [M]. Type Zophophiluscurticornis Fairmaire, 1881, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Zophoserodius Reitter, 58, 60 [M]. Type Erodiuszophosoides Allard, 1865, by subsequent designation (Löbl et al. 2008a: 43). Status: valid subgenus of Erodius Fabricius, 1775 in Pimeliinae: Erodiini.

Zophosis Latreille, 167 [F]. Type Erodiustestudinarius Fabricius, 1781, by monotypy. Status: valid genus and subgenus in Pimeliinae: Zophosini.

Zophosodactylus Koch, 146 [M]. Type Protodactylussanctaemariae Koch, 1962, by original designation. Status: junior synonym of Protodactylus Koch, 1952 in Pimeliinae: Zophosini. Synonymy: Penrith (1981c: 129).

Zoutpansbergia Koch, 388 [F]. Type Zoutpansbergiaserricostata Koch, 1956, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina. Note: combined description of new genus-group taxon and new species (ICZN 1999, Article 13.4).

Zuercheria Reitter, 134 [F]. Type Zuercheriamatthiesseni Reitter, 1908, by subsequent designation (Löbl et al. 2008a: 43). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1911b: 590).

Zygas Pascoe, 487 [M]. Type Eurychoracimicoides Quensel, 1806, by original designation. Status: junior synonym of Lycanthropa J. Thomson, 1860 in Pimeliinae: Adelostomini. Synonymy: Haag-Rutenberg (1875b: 61–64).

Zypoetes Champion, 532 [M]. Type Zypoetesepieroides Champion, 1893, by monotypy. Status: valid genus in Phrenapatinae: Penetini.